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<mods:title id="EB454D6721AF88CD12C238228D87035E">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<taxonomicName id="4C674D4CFFC3FF96CC8B0556FD21FAC9" authority="Abrams and Harvey" authorityName="Abrams and Harvey" authorityYear="2020" box="[151,705,1322,1349]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="warramboo" status="sp. nov.">
<emphasis id="B913EADDFFC3FF96CC8B0556FE5CFAC9" bold="true" box="[151,444,1322,1349]" italics="true" pageId="69" pageNumber="70">Draculoides warramboo</emphasis>
Abrams and Harvey
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,
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<paragraph id="8BD836CFFFC3FF96CC8B0533FE86FAE5" blockId="69.[151,1314,1322,1421]" box="[151,358,1359,1385]" pageId="69" pageNumber="70">
(
<figureCitation id="135C2A4AFFC3FF96CC830533FEEBFAE5" box="[159,267,1359,1385]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support &lt;80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="69" pageNumber="70">Figs. 18</figureCitation>
,
<figureCitation id="135C2A4AFFC3FF96CD0B0533FEBFFAE5" box="[279,351,1359,1385]" captionStart-0="FIGURE 41" captionStart-1="FIGURE 42" captionStart-2="FIGURE 43" captionStartId-0="70.[151,250,1650,1674]" captionStartId-1="71.[151,250,1415,1439]" captionStartId-2="72.[151,250,1390,1414]" captionTargetBox-0="[153,1436,489,1626]" captionTargetBox-1="[152,1435,813,1391]" captionTargetBox-2="[151,1430,181,1366]" captionTargetId-0="figure-128@70.[152,1436,489,1626]" captionTargetId-1="figure-318@71.[151,1436,813,1392]" captionTargetId-2="figure-20@72.[151,1436,181,1366]" captionTargetPageId-0="70" captionTargetPageId-1="71" captionTargetPageId-2="72" captionText-0="FIGURE 41. Draculoides warramboo sp. nov., holotype male (WAM T139912): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." captionText-1="FIGURE 42. Draculoides warramboo sp. nov., paratype female (WAM T138501): A. Body, dorsal; B. Body, ventral; C. Body, lateral. The scale bar shown in image A is 2 mm and also applies to B and C." captionText-2="FIGURE 43. Draculoides warramboo sp. nov.: AC, holotype male (WAM T139912): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. D, paratype female (WAM T138501): D. Spermathecae, ventral. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4416951" figureDoi-1="http://doi.org/10.5281/zenodo.4416953" figureDoi-2="http://doi.org/10.5281/zenodo.4416957" httpUri-0="https://zenodo.org/record/4416951/files/figure.png" httpUri-1="https://zenodo.org/record/4416953/files/figure.png" httpUri-2="https://zenodo.org/record/4416957/files/figure.png" pageId="69" pageNumber="70">4143</figureCitation>
)
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Zoobank Code: http://zoobank.org/NomenclaturalActs/
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<taxonomicName id="4C674D4CFFC3FF96CC8B05CBFE9CFA43" authority="SCH" authorityName="SCH" box="[151,380,1463,1487]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFC3FF96CC8B05CBFEA1FA43" box="[151,321,1463,1487]" italics="true" pageId="69" pageNumber="70">Paradraculoides</emphasis>
SCH
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057:
<bibRefCitation id="EFF64B3EFFC3FF96CDB705CBFD8EFA43" author="Abrams, K. M. &amp; Huey, J. A. &amp; Hillyer, M. J. &amp; Humphreys, W. F. &amp; Didham, R. K. &amp; Harvey, M. S." box="[427,622,1463,1487]" pageId="69" pageNumber="70" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. &amp; Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
Abrams
<emphasis id="B913EADDFFC3FF96CE1E05C4FDCEFA43" box="[514,558,1463,1487]" italics="true" pageId="69" pageNumber="70">et al</emphasis>
. 2019
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MPE 106532: 8, fig. 2.
</paragraph>
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<emphasis id="B913EADDFFC3FF96CC8B0581FE62F99B" bold="true" box="[151,386,1533,1559]" pageId="69" pageNumber="70">Material examined.</emphasis>
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<typeStatus id="54DC886DFFC3FF96CD960581FE11F99B" box="[394,497,1533,1559]" pageId="69" pageNumber="70" type="holotype">Holotype</typeStatus>
male.
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<collectingCountry id="F370765FFFC3FF96CE220581FD01F99B" box="[574,737,1533,1559]" name="Australia" pageId="69" pageNumber="70">AUSTRALIA</collectingCountry>
:
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<collectingRegion id="49A3F82DFFC3FF96CEF10582FC56F99B" box="[749,950,1533,1559]" country="Australia" name="Western Australia" pageId="69" pageNumber="70">Western Australia</collectingRegion>
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:
<collectingMunicipality id="6BBCACB5FFC3FF96CFDA0581FB3DF99B" box="[966,1245,1533,1559]" pageId="69" pageNumber="70">Warramboo Robe Valley</collectingMunicipality>
ca.
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W. of
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<emphasis id="B913EADDFFC3FF96CCDB0761FE9CF8BB" box="[199,380,1821,1847]" italics="true" pageId="69" pageNumber="70">Other material.</emphasis>
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Harvey
<emphasis id="B913EADDFFC3FF96CD6E071AFE46F8F3" box="[370,422,1893,1919]" italics="true" pageId="69" pageNumber="70">et al</emphasis>
., 2008
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)
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W Pannawonica
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(
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sc) (
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;
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W Pannawonica
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(
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) (
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;
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W Pannawonica
</locationDeviation>
,
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,
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,
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,
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(
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,
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-
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) (
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) (DNA:
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, 28S, 18S,
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;
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<specimenCount id="9D61FD46FFC0FF95CECE00A3FCA8FF75" box="[722,840,223,249]" pageId="70" pageNumber="71" type="juvenile">1 juvenile</specimenCount>
,
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,
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W Pannawonica
</locationDeviation>
,
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,
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,
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(
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,
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-
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) (
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) (DNA:
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, 28S, 18S,
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;
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<specimenCount id="9D61FD46FFC0FF95C80B015BFB6CFECD" box="[1047,1164,295,321]" pageId="70" pageNumber="71" type="juvenile">1 juvenile</specimenCount>
,
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,
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W Pannawonica
</locationDeviation>
,
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,
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, troglofauna scrape,
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(
<collectorName id="26925319FFC0FF95C8D80137FEC1FE05" pageId="70" pageNumber="71">Biota Environmental Sciences</collectorName>
,
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-
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scB) (
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)
</materialsCitation>
;
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<specimenCount id="9D61FD46FFC0FF95CF640113FC0CFE05" box="[888,1004,367,393]" pageId="70" pageNumber="71" type="juvenile">1 juvenile</specimenCount>
,
<location id="8EB86014FFC0FF95CFE40113FB9FFE05" LSID="urn:lsid:plazi:treatment:03CE87D9FFC3FF9BCC8B0556FF2AF82E:8EB86014FFC0FF95CFE40113FB9FFE05" box="[1016,1151,367,393]" country="Australia" latitude="-21.648577" longLatPrecision="1" longitude="115.83836" name="Warramboo" pageId="70" pageNumber="71" stateProvince="Western Australia">Warramboo</location>
,
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W Pannawonica
</locationDeviation>
,
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,
<date id="FFD9100FFFC0FF95CE1301EFFD4CFE21" box="[527,684,402,429]" pageId="70" pageNumber="71" value="2015-06-06">
<collectingDate id="EF9DE9E7FFC0FF95CE1301EFFD4CFE21" box="[527,684,402,429]" pageId="70" pageNumber="71" value="2015-06-06">06 June 2015</collectingDate>
</date>
, troglofauna scrape,
<collectorName id="26925319FFC0FF95CF8001EFFBCBFE21" box="[924,1067,403,429]" pageId="70" pageNumber="71">J. Alexander</collectorName>
(
<collectorName id="26925319FFC0FF95C82001EFFA78FE21" box="[1084,1432,403,429]" pageId="70" pageNumber="71">Biota Environmental Sciences</collectorName>
,
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-
<specimenCode id="DBC19EB4FFC0FF95CDA401CAFE31FE5D" box="[440,465,438,465]" collectionCode="MEARC" pageId="70" pageNumber="71">01</specimenCode>
scC) (
<collectionCode id="ED76AE0AFFC0FF95CE0D01CBFDB8FE5D" box="[529,600,439,465]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="70" pageNumber="71" type="Museum">WAM</collectionCode>
<specimenCode id="DBC19EB4FFC0FF95CE4201CBFD27FE5D" box="[606,711,439,465]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="70" pageNumber="71">T139911</specimenCode>
)
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.
</paragraph>
</subSubSection>
<caption id="DF186647FFC0FF95CC8B060EFBD2F95D" ID-DOI="http://doi.org/10.5281/zenodo.4416951" ID-Zenodo-Dep="4416951" httpUri="https://zenodo.org/record/4416951/files/figure.png" pageId="70" pageNumber="71" startId="70.[151,250,1650,1674]" targetBox="[153,1436,489,1626]" targetPageId="70">
<paragraph id="8BD836CFFFC0FF95CC8B060EFBD2F95D" blockId="70.[151,1436,1650,1746]" pageId="70" pageNumber="71">
<emphasis id="B913EADDFFC0FF95CC8B060EFEC1F906" bold="true" box="[151,289,1650,1674]" pageId="70" pageNumber="71">FIGURE 41.</emphasis>
<taxonomicName id="4C674D4CFFC0FF95CD34060EFDC2F906" authority="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey, 2020" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[296,546,1650,1674]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="warramboo" status="sp. nov.">
<emphasis id="B913EADDFFC0FF95CD34060EFDC2F906" box="[296,546,1650,1674]" italics="true" pageId="70" pageNumber="71">Draculoides warramboo</emphasis>
</taxonomicName>
<emphasis id="B913EADDFFC0FF95CE35060EFD9BF906" bold="true" box="[553,635,1650,1674]" pageId="70" pageNumber="71">
<taxonomicNameLabel id="A22057A6FFC0FF95CE35060EFD9BF906" box="[553,635,1650,1674]" pageId="70" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (WAM T139912): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F.
</paragraph>
</caption>
<subSubSection id="C37D6544FFC0FF94CCDB077CFE7CFEE9" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="8BD836CFFFC0FF94CCDB077CFE7CFEE9" blockId="70.[151,1437,1792,2035]" lastBlockId="71.[151,1437,151,789]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
<emphasis id="B913EADDFFC0FF95CCDB077CFEA2F896" bold="true" box="[199,322,1792,1818]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
The shape of the male flagellum of
<taxonomicName id="4C674D4CFFC0FF95CEC3077CFC14F896" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[735,1012,1792,1818]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFC0FF95CEC3077CFC14F896" box="[735,1012,1792,1818]" italics="true" pageId="70" pageNumber="71">Draculoides warramboo</emphasis>
</taxonomicName>
most closely resembles
<emphasis id="B913EADDFFC0FF95C915077DFEC0F8B2" italics="true" pageId="70" pageNumber="71">
<taxonomicName id="4C674D4CFFC0FF95C915077DFA95F896" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[1289,1397,1792,1818]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="affinis">D. affinis</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95C99D077DFEC0F8B2" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="anachoretus">D. anachoretus</taxonomicName>
</emphasis>
,
<taxonomicName id="4C674D4CFFC0FF95CD330759FE42F8B2" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[303,418,1828,1854]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFC0FF95CD330759FE42F8B2" box="[303,418,1828,1854]" italics="true" pageId="70" pageNumber="71">D. bythius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CDAC0759FDA7F8B2" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[432,583,1828,1854]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="cochranus">
<emphasis id="B913EADDFFC0FF95CDAC0759FDA7F8B2" box="[432,583,1828,1854]" italics="true" pageId="70" pageNumber="71">D. cochranus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CE480759FD15F8B2" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[596,757,1828,1854]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFC0FF95CE480759FD15F8B2" box="[596,757,1828,1854]" italics="true" pageId="70" pageNumber="71">D. gnophicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CF1E0759FC9DF8B2" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[770,893,1829,1854]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFC0FF95CF1E0759FC9DF8B2" box="[770,893,1829,1854]" italics="true" pageId="70" pageNumber="71">D. eremius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CF960759FC1EF8B2" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[906,1022,1828,1854]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFC0FF95CF960759FC1EF8B2" box="[906,1022,1828,1854]" italics="true" pageId="70" pageNumber="71">D. kryptus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFC0FF95C8290759FAE8F8B2" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[1077,1288,1828,1854]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFC0FF95C8290759FAE8F8B2" box="[1077,1288,1828,1854]" italics="true" pageId="70" pageNumber="71">D. mckechnieorum</emphasis>
</taxonomicName>
especially in the presence of a broad base. It differs from
<emphasis id="B913EADDFFC0FF95CE900735FBCDF8EE" box="[652,1069,1864,1890]" italics="true" pageId="70" pageNumber="71">
<taxonomicName id="4C674D4CFFC0FF95CE900735FD18F8EE" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[652,760,1864,1890]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="affinis">D. affinis</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CF180735FC60F8EE" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[772,896,1865,1890]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="eremius">D. eremius</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CF910735FBCDF8EE" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[909,1069,1864,1890]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="gnophicola">D. gnophicola</taxonomicName>
</emphasis>
and
<taxonomicName id="4C674D4CFFC0FF95C8790735FB3AF8EE" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1125,1242,1864,1890]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFC0FF95C8790735FB3AF8EE" box="[1125,1242,1864,1890]" italics="true" pageId="70" pageNumber="71">D. kryptus</emphasis>
</taxonomicName>
by the sub-distal placement of dm4 (close to distal margin in
<emphasis id="B913EADDFFC0FF95CE880711FBDCF80A" box="[660,1084,1900,1926]" italics="true" pageId="70" pageNumber="71">
<taxonomicName id="4C674D4CFFC0FF95CE880711FCE1F80A" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[660,769,1900,1926]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="affinis">D. affinis</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CF120711FC6CF80A" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[782,908,1901,1926]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="eremius">D. eremius</taxonomicName>
,
<taxonomicName id="4C674D4CFFC0FF95CF860711FBDCF80A" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[922,1084,1900,1926]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="gnophicola">D. gnophicola</taxonomicName>
</emphasis>
and
<taxonomicName id="4C674D4CFFC0FF95C86A0711FB0CF80A" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1142,1260,1900,1926]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFC0FF95C86A0711FB0CF80A" box="[1142,1260,1900,1926]" italics="true" pageId="70" pageNumber="71">D. kryptus</emphasis>
</taxonomicName>
), from
<taxonomicName id="4C674D4CFFC0FF95C9590711FF16F826" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="B913EADDFFC0FF95C9590711FF16F826" italics="true" pageId="70" pageNumber="71">D. anachoretus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C674D4CFFC0FF95CD3107EDFE40F826" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[301,416,1936,1962]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFC0FF95CD3107EDFE40F826" box="[301,416,1936,1962]" italics="true" pageId="70" pageNumber="71">D. bythius</emphasis>
</taxonomicName>
by the close proximity of dl1 and vl1 (far apart in
<emphasis id="B913EADDFFC0FF95CFCA07EDFB77F826" box="[982,1175,1936,1962]" italics="true" pageId="70" pageNumber="71">
<taxonomicName id="4C674D4CFFC0FF95CFCA07EDFB61F826" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[982,1153,1936,1962]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="anachoretus">D. anachoretus</taxonomicName>
a
</emphasis>
nd
<taxonomicName id="4C674D4CFFC0FF95C8A507EDFACDF826" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1209,1325,1936,1962]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFC0FF95C8A507EDFACDF826" box="[1209,1325,1936,1962]" italics="true" pageId="70" pageNumber="71">D. bythius</emphasis>
</taxonomicName>
), from
<taxonomicName id="4C674D4CFFC0FF95C99D07EDFEECF842" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="cochranus">
<emphasis id="B913EADDFFC0FF95C99D07EDFEECF842" italics="true" pageId="70" pageNumber="71">D. cochranus</emphasis>
</taxonomicName>
by the anterior position of vm3, close to vm1 and vm2 (vm3 is situated in the middle of the flagellum of
<emphasis id="B913EADDFFC0FF95CCA407A5FEB8F87E" box="[184,344,2008,2034]" italics="true" pageId="70" pageNumber="71">
<taxonomicName id="4C674D4CFFC0FF95CCA407A5FEB3F87E" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[184,339,2008,2034]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="cochranus">D. cochranus</taxonomicName>
,
</emphasis>
midway between vm5 and vm1) and from
<taxonomicName id="4C674D4CFFC0FF95CF5207A5FBC4F87E" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[846,1060,2008,2034]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFC0FF95CF5207A5FBC4F87E" box="[846,1060,2008,2034]" italics="true" pageId="70" pageNumber="71">D. mckechnieorum</emphasis>
</taxonomicName>
by the more anterior position of dm4 which is not level with dl3 as in
<emphasis id="B913EADDFFC1FF94CE5800EBFCC1FF3D" box="[580,801,151,177]" italics="true" pageId="71" pageNumber="72">
<taxonomicName id="4C674D4CFFC1FF94CE5800EBFCFDFF3D" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[580,797,151,177]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="mckechnieorum">D. mckechnieorum</taxonomicName>
.
</emphasis>
Female flagellum is unknown.
<taxonomicName id="4C674D4CFFC1FF94C89B00EBFA7DFF3D" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[1159,1437,151,177]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFC1FF94C89B00EBFA7DFF3D" box="[1159,1437,151,177]" italics="true" pageId="71" pageNumber="72">Draculoides warramboo</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFC1FF94CE1E00C7FD6CFF59" box="[514,652,187,213]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFC1FF94CE1E00C7FD6CFF59" box="[514,652,187,213]" italics="true" pageId="71" pageNumber="72">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at COI by the 50bp mini-barcode shown in
<figureCitation id="135C2A4AFFC1FF94CCA900A3FEF6FF75" box="[181,278,223,249]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="71" pageNumber="72">Figure 3</figureCitation>
.
<taxonomicName id="4C674D4CFFC1FF94CD3F00A3FDD9FF75" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[291,569,223,249]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFC1FF94CD3F00A3FDD9FF75" box="[291,569,223,249]" italics="true" pageId="71" pageNumber="72">Draculoides warramboo</emphasis>
</taxonomicName>
can be diagnosed from all other
<taxonomicName id="4C674D4CFFC1FF94CFB300A3FBD9FF75" box="[943,1081,223,249]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B913EADDFFC1FF94CFB300A3FBD9FF75" box="[943,1081,223,249]" italics="true" pageId="71" pageNumber="72">Draculoides</emphasis>
</taxonomicName>
species that were sequenced at ITS2 (
<figureCitation id="135C2A4AFFC1FF94CCC2017FFEC1FE91" box="[222,289,259,285]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="71" pageNumber="72">Fig. 7</figureCitation>
) except for
<taxonomicName id="4C674D4CFFC1FF94CDB4017FFDACFE91" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[424,588,259,285]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="anachoretus">
<emphasis id="B913EADDFFC1FF94CDB4017FFDACFE91" box="[424,588,259,285]" italics="true" pageId="71" pageNumber="72">D.anachoretus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94CE46017FFD2DFE91" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[602,717,259,285]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="bythius">
<emphasis id="B913EADDFFC1FF94CE46017FFD2DFE91" box="[602,717,259,285]" italics="true" pageId="71" pageNumber="72">D. bythius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94CEC7017FFCB5FE90" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[731,853,259,284]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="eremius">
<emphasis id="B913EADDFFC1FF94CEC7017FFCB5FE90" box="[731,853,259,284]" italics="true" pageId="71" pageNumber="72">D. eremius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94CF7E017FFBE2FE91" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[866,1026,259,285]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="gnophicola">
<emphasis id="B913EADDFFC1FF94CF7E017FFBE2FE91" box="[866,1026,259,285]" italics="true" pageId="71" pageNumber="72">D. gnophicola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94C80C017FFB64FE91" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1040,1156,259,285]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="kryptus">
<emphasis id="B913EADDFFC1FF94C80C017FFB64FE91" box="[1040,1156,259,285]" italics="true" pageId="71" pageNumber="72">D. kryptus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94C88E017FFA85FE91" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[1170,1381,259,285]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="mckechnieorum">
<emphasis id="B913EADDFFC1FF94C88E017FFA85FE91" box="[1170,1381,259,285]" italics="true" pageId="71" pageNumber="72">D. mckechnieorum</emphasis>
</taxonomicName>
,,
<taxonomicName id="4C674D4CFFC1FF94C99D017FFEF0FECD" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="immortalis">
<emphasis id="B913EADDFFC1FF94C99D017FFEF0FECD" italics="true" pageId="71" pageNumber="72">D. immortalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94CD3C015BFE5FFECD" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[288,447,295,321]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="belalugosii">
<emphasis id="B913EADDFFC1FF94CD3C015BFE5FFECD" box="[288,447,295,321]" italics="true" pageId="71" pageNumber="72">D. belalugosii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94CDD2015BFD7CFECD" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[462,668,295,321]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="christopherleei">
<emphasis id="B913EADDFFC1FF94CDD2015BFD7CFECD" box="[462,668,295,321]" italics="true" pageId="71" pageNumber="72">D. christopherleei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C674D4CFFC1FF94CEB6015BFCA5FECD" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[682,837,295,321]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="piscivultus">
<emphasis id="B913EADDFFC1FF94CEB6015BFCA5FECD" box="[682,837,295,321]" italics="true" pageId="71" pageNumber="72">D. piscivultus</emphasis>
</taxonomicName>
and
<emphasis id="B913EADDFFC1FF94CF62015BFBE3FECD" box="[894,1027,295,321]" italics="true" pageId="71" pageNumber="72">
<taxonomicName id="4C674D4CFFC1FF94CF62015BFBE0FECD" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[894,1024,295,321]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="akashae">D. akashae</taxonomicName>
,
</emphasis>
which are not distinguishable using the ITS2 mini-barcode.
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFC1FF9BCCDB0112FD13F9D2" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="description">
<paragraph id="8BD836CFFFC1FF94CCDB0112FB2EFE05" blockId="71.[151,1437,151,789]" box="[199,1230,366,393]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB0112FE5BFE04" bold="true" box="[199,443,366,392]" pageId="71" pageNumber="72">Description (adults).</emphasis>
<emphasis id="B913EADDFFC1FF94CDDE0113FDF1FE05" box="[450,529,367,393]" italics="true" pageId="71" pageNumber="72">Colour</emphasis>
. Yellow-brown; propeltidium and pedipalps somewhat darker.
</paragraph>
<paragraph id="8BD836CFFFC1FF94CCDB01EFFB93FE79" blockId="71.[151,1437,151,789]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB01EFFE8FFE21" box="[199,367,403,429]" italics="true" pageId="71" pageNumber="72">Cephalothorax</emphasis>
. Propeltidium with 2 +1 apical setae in a triangular formation on anterior process and 2 + 2 + 2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 12 (
<specimenCount id="9D61FD46FFC1FF94C91201CBFAC4FE5D" box="[1294,1316,439,465]" pageId="71" pageNumber="72" type="male"></specimenCount>
) (
<specimenCount id="9D61FD46FFC1FF94C92301CBFAB1FE5D" box="[1343,1361,439,465]" pageId="71" pageNumber="72" type="female"></specimenCount>
) setae (including 2 sternapophysial setae); posterior sternum triangular with 6 (
<specimenCount id="9D61FD46FFC1FF94CFD801A7FC3BFE79" box="[964,987,475,501]" pageId="71" pageNumber="72" type="male"></specimenCount>
), 7 (
<specimenCount id="9D61FD46FFC1FF94C80C01A7FBC2FE79" box="[1040,1058,475,501]" pageId="71" pageNumber="72" type="female"></specimenCount>
) setae.
</paragraph>
<paragraph id="8BD836CFFFC1FF94CCDB0183FAADFD25" blockId="71.[151,1437,151,789]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB0183FED5FD95" box="[199,309,511,537]" italics="true" pageId="71" pageNumber="72">Chelicera</emphasis>
. Fixed finger with 2 large teeth plus 4 (
<specimenCount id="9D61FD46FFC1FF94CEE60183FCF1FD95" box="[762,785,511,537]" pageId="71" pageNumber="72" type="male"></specimenCount>
), 4 (
<specimenCount id="9D61FD46FFC1FF94CF5B0183FCB9FD95" box="[839,857,511,537]" pageId="71" pageNumber="72" type="female"></specimenCount>
) smaller teeth between these and 1 lateral tooth on proximal large tooth; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 composed of 6 (
<specimenCount id="9D61FD46FFC1FF94CE34023BFDDFFDED" box="[552,575,583,609]" pageId="71" pageNumber="72" type="male"></specimenCount>
), 8 (
<specimenCount id="9D61FD46FFC1FF94CE6F023BFD65FDED" box="[627,645,583,609]" pageId="71" pageNumber="72" type="female"></specimenCount>
) setae; G3 composed of 5 (
<specimenCount id="9D61FD46FFC1FF94CFAB023BFC2DFDED" box="[951,973,583,609]" pageId="71" pageNumber="72" type="male"></specimenCount>
) (
<specimenCount id="9D61FD46FFC1FF94CFFB023BFC19FDED" box="[999,1017,583,609]" pageId="71" pageNumber="72" type="female"></specimenCount>
) setae; internal face of chelicera with 3 (
<specimenCount id="9D61FD46FFC1FF94CCA90217FF2CFD09" box="[181,204,619,645]" pageId="71" pageNumber="72" type="male"></specimenCount>
), 5 (
<specimenCount id="9D61FD46FFC1FF94CCE30217FEF1FD09" box="[255,273,619,645]" pageId="71" pageNumber="72" type="female"></specimenCount>
) short whip-like setae (G4); brush at base of fixed finger composed of 8 setae (G5A) each densely pilose in distal half and G5B composed of 10 (
<specimenCount id="9D61FD46FFC1FF94CE4502F3FD90FD25" box="[601,624,655,681]" pageId="71" pageNumber="72" type="male"></specimenCount>
), 9 (
<specimenCount id="9D61FD46FFC1FF94CEB902F3FD57FD25" box="[677,695,655,681]" pageId="71" pageNumber="72" type="female"></specimenCount>
) setae; G6 with one seta; G7 composed of 4 (
<specimenCount id="9D61FD46FFC1FF94C8A502F3FB2FFD25" box="[1209,1231,655,681]" pageId="71" pageNumber="72" type="male"></specimenCount>
) (
<specimenCount id="9D61FD46FFC1FF94C8F602F3FB1CFD25" box="[1258,1276,655,681]" pageId="71" pageNumber="72" type="female"></specimenCount>
) setae.
</paragraph>
<paragraph id="8BD836CFFFC1FF94CCDB02CFFEABFC99" blockId="71.[151,1437,151,789]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB02CFFECCFD41" box="[199,300,691,717]" italics="true" pageId="71" pageNumber="72">Pedipalp</emphasis>
. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 8 (
<specimenCount id="9D61FD46FFC1FF94CCC102ABFF14FD7D" box="[221,244,727,753]" pageId="71" pageNumber="72" type="male"></specimenCount>
), 7 (
<specimenCount id="9D61FD46FFC1FF94CD3602ABFEDCFD7D" box="[298,316,727,753]" pageId="71" pageNumber="72" type="female"></specimenCount>
) stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.7 (
<specimenCount id="9D61FD46FFC1FF94C97A02ABFA9CFD7D" box="[1382,1404,727,753]" pageId="71" pageNumber="72" type="male"></specimenCount>
) × length of tarsus.
</paragraph>
<caption id="DF186647FFC1FF94CC8B05FBFC41FA4F" ID-DOI="http://doi.org/10.5281/zenodo.4416953" ID-Zenodo-Dep="4416953" httpUri="https://zenodo.org/record/4416953/files/figure.png" pageId="71" pageNumber="72" startId="71.[151,250,1415,1439]" targetBox="[152,1435,813,1391]" targetPageId="71">
<paragraph id="8BD836CFFFC1FF94CC8B05FBFC41FA4F" blockId="71.[151,1436,1415,1475]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CC8B05FBFEC0FA13" bold="true" box="[151,288,1415,1439]" pageId="71" pageNumber="72">FIGURE 42.</emphasis>
<taxonomicName id="4C674D4CFFC1FF94CD3905FBFDFFFA13" authority="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey, 2020" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[293,543,1415,1439]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="warramboo" status="sp. nov.">
<emphasis id="B913EADDFFC1FF94CD3905FBFDFFFA13" box="[293,543,1415,1439]" italics="true" pageId="71" pageNumber="72">Draculoides warramboo</emphasis>
</taxonomicName>
<emphasis id="B913EADDFFC1FF94CE3905FBFD95FA13" bold="true" box="[549,629,1415,1439]" pageId="71" pageNumber="72">
<taxonomicNameLabel id="A22057A6FFC1FF94CE3905FBFD95FA13" box="[549,629,1415,1439]" pageId="71" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female (WAM T138501): A. Body, dorsal; B. Body, ventral; C. Body, lateral. The scale bar shown in image A is 2 mm and also applies to B and C.
</paragraph>
</caption>
<paragraph id="8BD836CFFFC1FF94CCDB058EFB0EF987" blockId="71.[151,1437,1521,2016]" box="[199,1262,1521,1548]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB058EFEE4F987" box="[199,260,1522,1547]" italics="true" pageId="71" pageNumber="72">Legs.</emphasis>
Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
</paragraph>
<paragraph id="8BD836CFFFC1FF94CCDB0669FB09F9D8" blockId="71.[151,1437,1521,2016]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB0669FED1F9A3" box="[199,305,1557,1583]" italics="true" pageId="71" pageNumber="72">Abdomen</emphasis>
. Chaetotaxy of tergites IIX: 2 macrosetae + 4 microsetae: 3 macrosetae + 6 microsetae (microsetae in column): 2: 2: 2: 2: 2: 2: 2 (
<specimenCount id="9D61FD46FFC1FF94CDF70645FDE2F9DF" box="[491,514,1593,1619]" pageId="71" pageNumber="72" type="male"></specimenCount>
), 2: 2: 2: 2: 2: 4: 4 (
<specimenCount id="9D61FD46FFC1FF94CEF80645FD17F9DF" box="[740,759,1593,1619]" pageId="71" pageNumber="72" type="female"></specimenCount>
); segment XII without small dorsal process.
</paragraph>
<paragraph id="8BD836CFFFC1FF94CCDB0621FDEBF933" blockId="71.[151,1437,1521,2016]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB0621FE6AF9FB" box="[199,394,1629,1655]" italics="true" pageId="71" pageNumber="72">Female genitalia</emphasis>
. Two pairs of spermathecae with outer lobe sub-equal to inner lobe, each pair connected basally before connection with bursa (
<figureCitation id="135C2A4AFFC1FF94CE4C06FDFD5BF917" box="[592,699,1665,1691]" captionStart="FIGURE 43" captionStartId="72.[151,250,1390,1414]" captionTargetBox="[151,1430,181,1366]" captionTargetId="figure-20@72.[151,1436,181,1366]" captionTargetPageId="72" captionText="FIGURE 43. Draculoides warramboo sp. nov.: AC, holotype male (WAM T139912): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. D, paratype female (WAM T138501): D. Spermathecae, ventral. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4416957" httpUri="https://zenodo.org/record/4416957/files/figure.png" pageId="71" pageNumber="72">Fig. 43D</figureCitation>
), distally round and smooth; sparsely covered with small pores; gonopod short, distally bifurcate.
</paragraph>
<paragraph id="8BD836CFFFC1FF94CCDB06B5FCCCF8FF" blockId="71.[151,1437,1521,2016]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB06B5FEDAF96F" box="[199,314,1737,1763]" italics="true" pageId="71" pageNumber="72">Flagellum</emphasis>
. Male: Dorsoventrally compressed 2 × longer than broad (
<figureCitation id="135C2A4AFFC1FF94CFCF06B5FBCEF96F" box="[979,1070,1737,1763]" captionStart="FIGURE 41" captionStartId="70.[151,250,1650,1674]" captionTargetBox="[153,1436,489,1626]" captionTargetId="figure-128@70.[152,1436,489,1626]" captionTargetPageId="70" captionText="FIGURE 41. Draculoides warramboo sp. nov., holotype male (WAM T139912): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 1 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." figureDoi="http://doi.org/10.5281/zenodo.4416951" httpUri="https://zenodo.org/record/4416951/files/figure.png" pageId="71" pageNumber="72">Figs. 41</figureCitation>
DF, 43AC); seta dm1 situated dorso-medially, slightly closer to anterior margin; seta dm4 situated close to posterior margin; dl1 between dl3 and vl1 but much closer to vl1; dl3 on posterior margin; vm2 situated slightly above vm1; vm5 situated on approximately same level as dl1, closer to vl2 than to vm5; at least three pairs of microsetae between vl1 and dl3. Female flagellum unknown, specimen collected without flagellum.
</paragraph>
<paragraph id="8BD836CFFFC1FF94CCDB0702FE61F86C" blockId="71.[151,1437,1521,2016]" pageId="71" pageNumber="72">
<emphasis id="B913EADDFFC1FF94CCDB0702FE6DF81B" box="[199,397,1917,1943]" italics="true" pageId="71" pageNumber="72">Dimensions (mm)</emphasis>
.
<typeStatus id="54DC886DFFC1FF94CD860701FDE2F81B" box="[410,514,1917,1943]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
male (WAM T139912): Body length 3.51. Propeltidium 1.10/0.42. Chelicera 0.69. Flagellum 0.38/0.19. Pedipalp: trochanter 0.54, femur 0.48, patella 0.48, tibia 0.42, tarsus 0.19, claw 0.13, total excluding claw 2.25.
</paragraph>
<caption id="DF186647FFCEFF9BCC8B0512FAF9FA26" ID-DOI="http://doi.org/10.5281/zenodo.4416957" ID-Zenodo-Dep="4416957" httpUri="https://zenodo.org/record/4416957/files/figure.png" pageId="72" pageNumber="73" startId="72.[151,250,1390,1414]" targetBox="[151,1430,181,1366]" targetPageId="72">
<paragraph id="8BD836CFFFCEFF9BCC8B0512FAF9FA26" blockId="72.[151,1437,1390,1451]" pageId="72" pageNumber="73">
<emphasis id="B913EADDFFCEFF9BCC8B0512FEC2FA0A" bold="true" box="[151,290,1390,1414]" pageId="72" pageNumber="73">FIGURE 43.</emphasis>
<taxonomicName id="4C674D4CFFCEFF9BCD370512FDC6FA0A" authority="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey, 2020" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[299,550,1390,1414]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="warramboo" status="sp. nov.">
<emphasis id="B913EADDFFCEFF9BCD370512FDC6FA0A" box="[299,550,1390,1414]" italics="true" pageId="72" pageNumber="73">Draculoides warramboo</emphasis>
</taxonomicName>
<emphasis id="B913EADDFFCEFF9BCE330512FD62FA0A" bold="true" box="[559,642,1390,1414]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="A22057A6FFCEFF9BCE330512FD62FA0A" box="[559,642,1390,1414]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: AC, holotype male (WAM T139912): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. D, paratype female (WAM T138501): D. Spermathecae, ventral. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BD836CFFFCEFF9BCCDB05A5FF30F9B6" blockId="72.[151,1437,1496,1954]" pageId="72" pageNumber="73">
<typeStatus id="54DC886DFFCEFF9BCCDB05A5FEC9FA7F" box="[199,297,1497,1523]" pageId="72" pageNumber="73" type="paratype">Paratype</typeStatus>
female (WAM T138501): Body length 4.76. Propeltidium 1.56/0.83. Chelicera 0.96. Flagellum missing. Pedipalp: trochanter 0.75, femur 0.52, patella 0.67, tibia 0.65, tarsus 0.35, claw 0.10, total excluding claw 2.94.
</paragraph>
<paragraph id="8BD836CFFFCEFF9BCCDB0639FD13F9D2" blockId="72.[151,1437,1496,1954]" box="[199,755,1604,1631]" pageId="72" pageNumber="73">
<emphasis id="B913EADDFFCEFF9BCCDB0639FECFF9D2" box="[199,303,1605,1630]" italics="true" pageId="72" pageNumber="73">Variation</emphasis>
. Body length (males) 3.463.51 (n = 2).
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFCEFF9BCCDB0614FCFAF8D6" pageId="72" pageNumber="73" type="discussion">
<paragraph id="8BD836CFFFCEFF9BCCDB0614FEDCF8BA" blockId="72.[151,1437,1496,1954]" pageId="72" pageNumber="73">
<emphasis id="B913EADDFFCEFF9BCCDB0614FEDBF90E" bold="true" box="[199,315,1640,1666]" pageId="72" pageNumber="73">Remarks.</emphasis>
<taxonomicName id="4C674D4CFFCEFF9BCD5D0614FDB4F90E" authorityName="Abrams &amp; Huey &amp; Hillyer &amp; Didham &amp; Harvey" authorityYear="2020" box="[321,596,1640,1666]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="warramboo">
<emphasis id="B913EADDFFCEFF9BCD5D0614FDB4F90E" box="[321,596,1640,1666]" italics="true" pageId="72" pageNumber="73">Draculoides warramboo</emphasis>
</taxonomicName>
is known from several locations within two areas known as Warramboo and Tod Bore (
<figureCitation id="135C2A4AFFCEFF9BCD0C06F0FE89F92A" box="[272,361,1676,1702]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="72" pageNumber="73">Fig. 1A</figureCitation>
). Warramboo is not a discrete mesa but appears to be the western extension of pisolitic geology from Mesa A (
<bibRefCitation id="EFF64B3EFFCEFF9BCD2606CCFDFAF946" author="Harvey, M. S. &amp; Berry, O. &amp; Edward, K. L. &amp; Humphreys, G." box="[314,538,1712,1739]" pageId="72" pageNumber="73" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. &amp; Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">
Harvey
<emphasis id="B913EADDFFCEFF9BCD8A06CDFE2FF946" box="[406,463,1712,1738]" italics="true" pageId="72" pageNumber="73">et al.</emphasis>
, 2008
</bibRefCitation>
), despite this, molecular data suggest no gene flow occurring between Warramboo and Mesa A (
<figureCitation id="135C2A4AFFCEFF9BCD7D06A8FE48F962" box="[353,424,1748,1774]" captionStart="FIGURE 2" captionStartId="17.[151,250,1910,1934]" captionTargetBox="[332,1254,181,1881]" captionTargetId="figure-17@17.[332,1254,181,1886]" captionTargetPageId="17" captionText="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support &lt;80 are not shown, except for the common ancestor of the Draculoides radiation." figureDoi="http://doi.org/10.5281/zenodo.4416855" httpUri="https://zenodo.org/record/4416855/files/figure.png" pageId="72" pageNumber="73">Fig. 2</figureCitation>
). Juveniles were sequenced by
<bibRefCitation id="EFF64B3EFFCEFF9BCF1306A9FC16F962" author="Harvey, M. S. &amp; Berry, O. &amp; Edward, K. L. &amp; Humphreys, G." box="[783,1014,1748,1775]" pageId="72" pageNumber="73" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. &amp; Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">
Harvey
<emphasis id="B913EADDFFCEFF9BCF7506A9FC7CF962" box="[873,924,1748,1774]" italics="true" pageId="72" pageNumber="73">et al</emphasis>
. (2008)
</bibRefCitation>
but not named due to a lack of adult specimens. The specimens listed above are associated with this species by locality and, in two cases, by sequence data (
<figureCitation id="135C2A4AFFCEFF9BCCC90760FECEF8BA" box="[213,302,1820,1846]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="72" pageNumber="73">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="8BD836CFFFCEFF9BCCDB073CFCFAF8D6" blockId="72.[151,1437,1496,1954]" box="[199,794,1856,1883]" pageId="72" pageNumber="73">
<emphasis id="B913EADDFFCEFF9BCCDB073CFE81F8D6" bold="true" box="[199,353,1856,1882]" pageId="72" pageNumber="73">Other names</emphasis>
. WAM SCH057 (
<bibRefCitation id="EFF64B3EFFCEFF9BCE35073CFCEEF8D6" author="Abrams, K. M. &amp; Huey, J. A. &amp; Hillyer, M. J. &amp; Humphreys, W. F. &amp; Didham, R. K. &amp; Harvey, M. S." box="[553,782,1856,1883]" pageId="72" pageNumber="73" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. &amp; Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
Abrams
<emphasis id="B913EADDFFCEFF9BCE97073DFD5CF8D6" box="[651,700,1856,1882]" italics="true" pageId="72" pageNumber="73">et al</emphasis>
., 2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D6544FFCEFF9BCCDB0718FF2AF82E" pageId="72" pageNumber="73" type="etymology">
<paragraph id="8BD836CFFFCEFF9BCCDB0718FF2AF82E" blockId="72.[151,1437,1496,1954]" pageId="72" pageNumber="73">
<emphasis id="B913EADDFFCEFF9BCCDB0718FEAEF8F2" bold="true" box="[199,334,1892,1918]" pageId="72" pageNumber="73">Etymology.</emphasis>
The specific epithet refers to the
<typeStatus id="54DC886DFFCEFF9BCED50719FD19F8F3" box="[713,761,1893,1919]" pageId="72" pageNumber="73">type</typeStatus>
locality, Warramboo. It is to be treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>