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<document id="D1C99D9ABB56866C04906D6174E641F7" ID-DOI="10.11646/zootaxa.3652.3.2" ID-GBIF-Dataset="9cb4b047-ae44-419e-898a-eb3bc27c3d15" ID-ISSN="1175-5326" ID-Zenodo-Dep="224020" ID-ZooBank="B334624E-A7F5-4077-BF64-D2AD6DB215D5" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460609979718" checkinUser="plazi" docAuthor="Mattos, Ingrid &amp; Mermudes, José Ricardo M." docDate="2013" docId="03CE87A7FFBD694DCFEFFF15FB7EF95D" docLanguage="en" docName="zt03652p342.pdf" docOrigin="Zootaxa 3652 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paxillus chapadensis Mattos &amp; Mermudes, new species" docType="treatment" docVersion="8" lastPageNumber="337" masterDocId="FFF7FFDFFFB76947CF78FFA2FFF8FFF8" masterDocTitle="Synopsis of Paxillus MacLeay, 1819 (Coleoptera: Passalidae): distributional records and descriptions of four new species from Brazil" masterLastPageNumber="342" masterPageNumber="327" pageNumber="337" updateTime="1698328243075" updateUser="plazi">
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<mods:title id="2B53BC2D1207C9A2A2279008A321901F">Synopsis of Paxillus MacLeay, 1819 (Coleoptera: Passalidae): distributional records and descriptions of four new species from Brazil</mods:title>
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<mods:name id="7D6C17ABDABD743DBF033A85264A9C67" type="personal">
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<mods:namePart id="64842DAFC0C1A4ADB28B14441CBC21F7">Mattos, Ingrid</mods:namePart>
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<mods:namePart id="2A7006E7E3DA67336F38D399FBA007B9">Mermudes, José Ricardo M.</mods:namePart>
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<mods:title id="7D414A9CEAC0F098F624B2F15A97ECB9">Zootaxa</mods:title>
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<mods:date id="56AE682C60EE7C6142892028EE56EA44">2013</mods:date>
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<mods:number id="DD3BF1734F290FA5D4E54050D0D649E6">3652</mods:number>
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<mods:identifier id="6387946355D986B675663DB0CBDF91DA" type="DOI">10.11646/zootaxa.3652.3.2</mods:identifier>
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<treatment id="03CE87A7FFBD694DCFEFFF15FB7EF95D" ID-DOI="http://doi.org/10.5281/zenodo.6154351" ID-GBIF-Taxon="119568115" ID-Zenodo-Dep="6154351" LSID="urn:lsid:plazi:treatment:03CE87A7FFBD694DCFEFFF15FB7EF95D" httpUri="http://treatment.plazi.org/id/03CE87A7FFBD694DCFEFFF15FB7EF95D" lastPageNumber="337" pageId="10" pageNumber="337">
<subSubSection id="C37D653AFFBD694DCFEFFF15FED6FF0D" pageId="10" pageNumber="337" type="nomenclature">
<paragraph id="8BD836B1FFBD694DCFEFFF15FCB9FF29" blockId="10.[151,833,183,245]" box="[151,833,183,209]" pageId="10" pageNumber="337">
<heading id="D09081DDFFBD694DCFEFFF15FCB9FF29" bold="true" box="[151,833,183,209]" fontSize="11" level="1" pageId="10" pageNumber="337" reason="1">
<emphasis id="B913EAA3FFBD694DCFEFFF15FCB9FF29" bold="true" box="[151,833,183,209]" pageId="10" pageNumber="337">
<taxonomicName id="4C674D32FFBD694DCFEFFF15FD5DFF29" authority="Mattos &amp; Mermudes" authorityName="Mattos &amp; Mermudes" box="[151,677,183,209]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="337" phylum="Arthropoda" rank="species" species="chapadensis" status="sp. nov.">
<emphasis id="B913EAA3FFBD694DCFEFFF15FE6DFF29" bold="true" box="[151,405,183,209]" italics="true" pageId="10" pageNumber="337">Paxillus chapadensis</emphasis>
Mattos &amp; Mermudes
</taxonomicName>
,
<taxonomicNameLabel id="A22057D8FFBD694DCDCAFF15FCB9FF29" box="[690,833,183,209]" pageId="10" pageNumber="337" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD836B1FFBD694DCFEFFF7FFED6FF0D" blockId="10.[151,833,183,245]" box="[151,302,221,245]" pageId="10" pageNumber="337">
(
<figureCitation id="135C2A34FFBD694DCFE7FF7FFEDEFF0D" box="[159,294,221,245]" captionStart="FIGURES 22 27" captionStartId="11.[151,269,1926,1950]" captionTargetBox="[158,1429,195,1894]" captionTargetId="figure@11.[151,1436,151,1901]" captionTargetPageId="11" captionText="FIGURES 22 27. Paxillus chapadensis, male: 22, head and anterior margin of pronotum, dorsal. 23 24, antennae: 23, dorsal; 24, ventral. 25 27, Aedeagus: 25, dorsal; 26, lateral; 27, ventral. Scale bars = 0.5 mm (Figs. 25 27 to the same scale); DS = dorsal spine, VI = ventral impression." httpUri="https://zenodo.org/record/224026/files/figure.png" pageId="10" pageNumber="337">Figs. 2227</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBD694DCFEFFE85FB38FA67" pageId="10" pageNumber="337" type="description">
<paragraph id="8BD836B1FFBD694DCFEFFE85FD64FAF2" blockId="10.[151,1437,295,1702]" pageId="10" pageNumber="337">
<emphasis id="B913EAA3FFBD694DCFEFFE85FED9FEB8" bold="true" box="[151,289,295,320]" pageId="10" pageNumber="337">Description</emphasis>
. Length
<quantity id="4C9F9B54FFBD694DCEFDFE85FDF9FEC7" box="[389,513,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="10" pageNumber="337" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">1619 mm</quantity>
. Head (
<figureCitation id="135C2A34FFBD694DCD2DFE85FD52FEC7" box="[597,682,295,319]" captionStart="FIGURES 22 27" captionStartId="11.[151,269,1926,1950]" captionTargetBox="[158,1429,195,1894]" captionTargetId="figure@11.[151,1436,151,1901]" captionTargetPageId="11" captionText="FIGURES 22 27. Paxillus chapadensis, male: 22, head and anterior margin of pronotum, dorsal. 23 24, antennae: 23, dorsal; 24, ventral. 25 27, Aedeagus: 25, dorsal; 26, lateral; 27, ventral. Scale bars = 0.5 mm (Figs. 25 27 to the same scale); DS = dorsal spine, VI = ventral impression." httpUri="https://zenodo.org/record/224026/files/figure.png" pageId="10" pageNumber="337">Fig. 22</figureCitation>
): labrum with anterior margin slightly concave, with fine setiferous punctures; anterior frontal area moderately to densely punctate, shallow, wide, and opaque, interstices unevenly narrowed; anterior margin of frontal area straight; median frontal area with weak elevation anterior to central tubercle; anterior and posterior frontal ridges elevated, evident, extending from central to medial frontal tubercles. Medial frontal tubercles elevated, robust, surpassing anterior margin of frontal area. Inner tubercles distinct, elevated, smaller than medial frontal tubercles; central tubercle short, not free, dilated at base, continuous with posterior lateral tubercles; these evident. Lateral post-frontal area usually punctate; bridge strong and smooth. Supraocular margin with tubercles and very fine punctures; canthus rounded at apex, ocular keel with fine and sparse punctures. Postocular sulcus strong and densely punctuate; punctures confluent, deep, and opaque. Antennae (
<figureCitation id="135C2A34FFBD694DCE6BFDD5FE66FD77" box="[275,414,631,655]" captionStart="FIGURES 22 27" captionStartId="11.[151,269,1926,1950]" captionTargetBox="[158,1429,195,1894]" captionTargetId="figure@11.[151,1436,151,1901]" captionTargetPageId="11" captionText="FIGURES 22 27. Paxillus chapadensis, male: 22, head and anterior margin of pronotum, dorsal. 23 24, antennae: 23, dorsal; 24, ventral. 25 27, Aedeagus: 25, dorsal; 26, lateral; 27, ventral. Scale bars = 0.5 mm (Figs. 25 27 to the same scale); DS = dorsal spine, VI = ventral impression." httpUri="https://zenodo.org/record/224026/files/figure.png" pageId="10" pageNumber="337">Figs. 2324</figureCitation>
): first antennomere of club short, second antennomere longer; scape with dorsal subapical spine and ventrally impressed with sparse setae. Lateral lobes of mentum somewhat pubescent and with few coarse punctures; scars irregular and open to outside, internally opaque (fossae of each lobe irregularly rounded, deeper and with the margin elevated and clearly marked on the inner portion). Pronotum: anterior margin sinuate; anterior marginal groove narrow and punctate, reaching middle of anterior margin, usually touching margin. Lateral marginal groove narrow and punctate, with fine, sparse pubescence. Sides of pronotum with irregular punctures, which are often fused; with punctate scar; posterior margin with or without short pubescence, or with yellowishorange tuft of setae. Prosternal process strongly impressed and opaque in middle; with shallow, irregular, confluent punctures; laterally smooth and shiny. Scutellum with median circular depression. Elytra with tuft of moderately long and dense setae, dorsal striae with small punctures; lateral striae with large punctures. Mesosternum smooth, with scar usually elongate (rarely oval), narrow and shallow deep; lateral anterior area punctate. Metasternum: disc evident by edges of margin; anterior lateral area punctate and pubescent; posterior lateral area moderately finely punctate, punctures contiguous with margin of disc. Lateral fossae of metasternum narrowing anteriorly with lower margin elevated and in posterior region slightly wider, lacking elevated margin, surface opaque. Sternite 7 rounded or slightly sinuate at apex. Mesotibia dorsally pubescent; with 1 lateral median spine. Aedeagus (
<figureCitation id="135C2A34FFBD694DCB94FB20FA8CFB62" box="[1260,1396,1154,1178]" captionStart="FIGURES 22 27" captionStartId="11.[151,269,1926,1950]" captionTargetBox="[158,1429,195,1894]" captionTargetId="figure@11.[151,1436,151,1901]" captionTargetPageId="11" captionText="FIGURES 22 27. Paxillus chapadensis, male: 22, head and anterior margin of pronotum, dorsal. 23 24, antennae: 23, dorsal; 24, ventral. 25 27, Aedeagus: 25, dorsal; 26, lateral; 27, ventral. Scale bars = 0.5 mm (Figs. 25 27 to the same scale); DS = dorsal spine, VI = ventral impression." httpUri="https://zenodo.org/record/224026/files/figure.png" pageId="10" pageNumber="337">Figs. 2527</figureCitation>
) as long as wide. Phallus subglobular, longer than parameres and phallobase together (lateral view). Phallobase and parameres fused, lacking suture line. Parameres weakly elongate, conspicuously curved and acuminate at apex, with a distinct downward curve (lateral view).
</paragraph>
<paragraph id="8BD836B1FFBD694DCFBFFAB5FB6EFAAD" blockId="10.[151,1437,295,1702]" pageId="10" pageNumber="337">
<emphasis id="B913EAA3FFBD694DCFBFFAB5FEAAFAC8" bold="true" box="[199,338,1303,1328]" pageId="10" pageNumber="337">Dimensions</emphasis>
(mm). Total length (from anterior margin of labrum to apex of elytra) 1619; elytral length (median line) 1011; pronotal length (median line) 45; pronotal width 6; humeral width 5.
</paragraph>
<paragraph id="8BD836B1FFBD694DCFBFFAC0FB38FA67" blockId="10.[151,1437,295,1702]" pageId="10" pageNumber="337">
<emphasis id="B913EAA3FFBD694DCFBFFAC0FEBDFA83" bold="true" box="[199,325,1378,1403]" pageId="10" pageNumber="337">Variability</emphasis>
. Frontal area with shallow punctures, these varying from contiguous to aligned in two or three irregular rows. In one of the
<typeStatus id="54DC8813FFBD694DCEA0FA2AFDBEFA58" box="[472,582,1416,1440]" pageId="10" pageNumber="337" type="paratype">paratypes</typeStatus>
, the scape of antennae has a spine on only the right side.
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBD694DCFBFFA0FFA61F9F7" pageId="10" pageNumber="337" type="discussion">
<paragraph id="8BD836B1FFBD694DCFBFFA0FFA61F9F7" blockId="10.[151,1437,295,1702]" pageId="10" pageNumber="337">
<emphasis id="B913EAA3FFBD694DCFBFFA0FFEC3FA3D" bold="true" box="[199,315,1453,1477]" pageId="10" pageNumber="337">Remarks.</emphasis>
<taxonomicName id="4C674D32FFBD694DCE3AFA0CFE1FFA3D" box="[322,487,1453,1477]" class="Insecta" family="Passallidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="337" phylum="Arthropoda" rank="species" species="chapadensis">
<emphasis id="B913EAA3FFBD694DCE3AFA0CFE1FFA3D" box="[322,487,1453,1477]" italics="true" pageId="10" pageNumber="337">P. chapadensis</emphasis>
</taxonomicName>
is distinguished from other species of the genus by the scape with a dorsal subapical spine and a ventral impression (
<figureCitation id="135C2A34FFBD694DCE86FA70FD70FA12" box="[510,648,1490,1514]" captionStart="FIGURES 22 27" captionStartId="11.[151,269,1926,1950]" captionTargetBox="[158,1429,195,1894]" captionTargetId="figure@11.[151,1436,151,1901]" captionTargetPageId="11" captionText="FIGURES 22 27. Paxillus chapadensis, male: 22, head and anterior margin of pronotum, dorsal. 23 24, antennae: 23, dorsal; 24, ventral. 25 27, Aedeagus: 25, dorsal; 26, lateral; 27, ventral. Scale bars = 0.5 mm (Figs. 25 27 to the same scale); DS = dorsal spine, VI = ventral impression." httpUri="https://zenodo.org/record/224026/files/figure.png" pageId="10" pageNumber="337">Figs. 2324</figureCitation>
), with sparse setae; and the prosternal process strongly impressed and opaque in the middle. The parameres (
<figureCitation id="135C2A34FFBD694DCD3DFA55FD6FF9F7" box="[581,663,1527,1551]" captionStart="FIGURES 22 27" captionStartId="11.[151,269,1926,1950]" captionTargetBox="[158,1429,195,1894]" captionTargetId="figure@11.[151,1436,151,1901]" captionTargetPageId="11" captionText="FIGURES 22 27. Paxillus chapadensis, male: 22, head and anterior margin of pronotum, dorsal. 23 24, antennae: 23, dorsal; 24, ventral. 25 27, Aedeagus: 25, dorsal; 26, lateral; 27, ventral. Scale bars = 0.5 mm (Figs. 25 27 to the same scale); DS = dorsal spine, VI = ventral impression." httpUri="https://zenodo.org/record/224026/files/figure.png" pageId="10" pageNumber="337">Fig. 26</figureCitation>
) are conspicuously curved and acuminate at the apex in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBD694DCFBFF9BFFD38F987" pageId="10" pageNumber="337" type="materials_examined">
<paragraph id="8BD836B1FFBD694DCFBFF9BFFD38F987" blockId="10.[151,1437,295,1702]" pageId="10" pageNumber="337">
<emphasis id="B913EAA3FFBD694DCFBFF9BFFE80F9CE" bold="true" box="[199,376,1565,1590]" pageId="10" pageNumber="337">
<typeStatus id="54DC8813FFBD694DCFBFF9BFFEFAF9CD" box="[199,258,1565,1589]" pageId="10" pageNumber="337">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54DC8813FFBD694DCEFAF9BFFE12F9CD" box="[386,490,1565,1589]" pageId="10" pageNumber="337" type="holotype">Holotype</typeStatus>
male from
<collectingCountry id="F3707621FFBD694DCD0DF9BFFD46F9CD" box="[629,702,1565,1589]" name="Brazil" pageId="10" pageNumber="337">Brazil</collectingCountry>
: Mato Grosso, Chapada dos Guimarães,
<date id="FFD91071FFBD694DCBE0F9BFFAE1F9CD" box="[1176,1305,1565,1589]" pageId="10" pageNumber="337" value="1983-11-27">27.XI.1983</date>
, Exp. Dep. Zool. UFPR leg., Mattos &amp; Mermudes det. 2012 (DZUP).
<typeStatus id="54DC8813FFBD694DCC53F9E1FC64F9A3" box="[811,924,1603,1627]" pageId="10" pageNumber="337" type="paratype">Paratypes</typeStatus>
: 3 (
<specimenCount id="9D61FD38FFBD694DCCB0F9E0FBECF9A2" box="[968,1044,1602,1626]" pageId="10" pageNumber="337" type="male">1 male</specimenCount>
,
<specimenCount id="9D61FD38FFBD694DCB59F9E0FB6AF9A2" box="[1057,1170,1602,1626]" pageId="10" pageNumber="337" type="female">2 females</specimenCount>
),
<date id="FFD91071FFBD694DCBDDF9E0FAD0F9A2" box="[1189,1320,1602,1626]" pageId="10" pageNumber="337" value="1983-11-27">27.XI.1983</date>
,
<specimenCount id="9D61FD38FFBD694DCA4BF9E0FA6FF9A2" box="[1331,1431,1602,1626]" pageId="10" pageNumber="337" type="female">1 female</specimenCount>
,
<date id="FFD91071FFBD694DCFEFF9C5FEE1F987" box="[151,281,1639,1663]" pageId="10" pageNumber="337" value="1983-11-18">18.XI.1983</date>
, Exp. Dep. Zool. UFPR leg. (DZUP).
</paragraph>
</subSubSection>
<subSubSection id="C37D653AFFBD694DCFBFF92FFB7EF95D" box="[199,1158,1677,1702]" pageId="10" pageNumber="337" type="etymology">
<paragraph id="8BD836B1FFBD694DCFBFF92FFB7EF95D" blockId="10.[151,1437,295,1702]" box="[199,1158,1677,1702]" pageId="10" pageNumber="337">
<emphasis id="B913EAA3FFBD694DCFBFF92FFEBEF95D" bold="true" box="[199,326,1677,1701]" pageId="10" pageNumber="337">Etymology</emphasis>
. The species is named after the
<typeStatus id="54DC8813FFBD694DCDD5F92CFD25F95E" box="[685,733,1678,1702]" pageId="10" pageNumber="337">type</typeStatus>
locality, the Chapada dos Guimarães.
</paragraph>
</subSubSection>
</treatment>
</document>