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<document ID-DOI="http://doi.org/10.5281/zenodo.3792242" ID-GBIF-Dataset="6bed073a-175c-42e6-bb99-645c99d55b4f" ID-GBIF-Taxon="163750593" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792242" ID-ZooBank="C631A347-306E-4773-84A4-E4712329186B" LSID="urn:lsid:zoobank.org:act:BD950E49-6462-434C-8BDD-587EAE361BB3" checkinTime="1587984202003" checkinUser="plazi" docAuthor="Miller, Jeremy, Griswold, Charles &amp; Yin, Chang" docDate="2009" docId="03ACB915FFB17424FF4E40E0FE36FEF6" docLanguage="en" docName="ZK_article_1993.pdf" docOrigin="ZooKeys 11 (11)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Maymena paquini Miller, Griswold &amp; Yin 2009, sp. n." docType="treatment" docUuid="BD950E49-6462-434C-8BDD-587EAE361BB3" docUuidSource="ZooBank" docVersion="8" lastPageId="48" lastPageNumber="56" masterDocId="FF95C16DFF9F7414FFC2455DFFF1FFB8" masterDocTitle="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers" masterLastPageNumber="195" masterPageNumber="9" pageId="46" pageNumber="55" updateTime="1635491590422" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Jeremy</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Griswold, Charles</mods:namePart>
<mods:affiliation>California Academy of Sciences ,, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Chang</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University ,, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-05-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>11</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>11</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>9</mods:start>
<mods:end>195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.11.160</mods:identifier>
<mods:identifier type="GBIF-Dataset">6bed073a-175c-42e6-bb99-645c99d55b4f</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576459</mods:identifier>
<mods:identifier type="ZooBank">C631A347-306E-4773-84A4-E4712329186B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792242" ID-GBIF-Taxon="163750593" ID-Zenodo-Dep="3792242" LSID="urn:lsid:zoobank.org:act:BD950E49-6462-434C-8BDD-587EAE361BB3" httpUri="http://treatment.plazi.org/id/03ACB915FFB17424FF4E40E0FE36FEF6" lastPageId="48" lastPageNumber="56" pageId="46" pageNumber="55">
<subSubSection box="[140,722,1469,1495]" pageId="46" pageNumber="55" type="nomenclature">
<paragraph blockId="46.[140,981,1469,1566]" box="[140,722,1469,1495]" pageId="46" pageNumber="55">
<heading box="[140,722,1469,1495]" fontSize="11" level="2" pageId="46" pageNumber="55" reason="3">
<emphasis bold="true" box="[140,722,1469,1495]" pageId="46" pageNumber="55">
<taxonomicName authority="Miller, Griswold &amp; Yin" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[140,645,1469,1495]" class="Arachnida" family="Mysmenidae" genus="Maymena" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="55" phylum="Arthropoda" rank="species" species="paquini" status="sp. n.">
<emphasis bold="true" box="[140,360,1469,1495]" italics="true" pageId="46" pageNumber="55">
<emphasis box="[140,258,1469,1495]" inLineHeading="true" pageId="46" pageNumber="55" reason="1">Maymena</emphasis>
paquini
</emphasis>
Miller, Griswold &amp; Yin
</taxonomicName>
,
<taxonomicNameLabel box="[657,722,1469,1495]" pageId="46" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="55" type="description">
<paragraph blockId="46.[140,981,1469,1566]" box="[140,981,1504,1531]" pageId="46" pageNumber="55">
<uri box="[140,981,1504,1531]" pageId="46" pageNumber="55">
urn:lsid:zoobank.org:act:
<uuid box="[419,981,1504,1531]" pageId="46" pageNumber="55">BD950E49-6462-434C-8BDD-587EAE361BB3</uuid>
</uri>
</paragraph>
<paragraph blockId="46.[140,981,1469,1566]" box="[140,392,1539,1566]" pageId="46" pageNumber="55">
<figureCitation box="[140,221,1539,1566]" captionStart="Figure 50" captionStartId="122.[140,212,958,980]" captionTargetBox="[140,1108,165,938]" captionTargetId="figure@122.[140,1109,164,939]" captionTargetPageId="122" captionText="Figure 50. A-C, Chanea suukyii sp. n. from Maku ridge; D-F, Maymena paquini sp. n. from Wayala Ku cave; G, H; Maymena kehen sp. n. from Lishadi, unlabeled arrow indicates diagnostic notch in scape; I, Gaiziapis zhizhuba sp. n. from Nankang Yakou.A, D, G, epigynum, ventral veiw; B, F, H, I, cleared vulva, dorsal view; C, male palp, cleared and expanded. BC: booklung cover; BH: basal hematodocha; CD: copulatory duct; FD: fertilization duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768845" httpUri="https://zenodo.org/record/3768845/files/figure.png" pageId="46" pageNumber="55">Figs 50</figureCitation>
D-F,
<figureCitation box="[280,350,1539,1566]" captionStart-0="Figure 53" captionStart-1="Figure 54" captionStart-2="Figure 55" captionStart-3="Figure 56" captionStart-4="Figure 57" captionStartId-0="125.[140,212,1162,1184]" captionStartId-1="126.[140,212,1340,1362]" captionStartId-2="127.[140,212,1203,1225]" captionStartId-3="128.[140,212,1204,1226]" captionStartId-4="129.[140,212,1206,1228]" captionTargetBox-0="[159,1089,183,1131]" captionTargetBox-1="[150,1096,173,1309]" captionTargetBox-2="[140,1108,165,1184]" captionTargetBox-3="[140,1108,165,1184]" captionTargetBox-4="[140,1108,165,1186]" captionTargetId-0="figure@125.[140,1108,164,1143]" captionTargetId-1="figure@126.[140,1109,164,1320]" captionTargetId-2="figure@127.[140,1109,164,1184]" captionTargetId-3="figure@128.[140,1108,164,1184]" captionTargetId-4="figure@129.[140,1109,164,1186]" captionTargetPageId-0="125" captionTargetPageId-1="126" captionTargetPageId-2="127" captionTargetPageId-3="128" captionTargetPageId-4="129" captionText-0="Figure 53. Maymena paquini sp. n. from Wayala Ku cave. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, female, posterior." captionText-1="Figure 54. Maymena paquini sp. n. from Wayala Ku cave, male palp. A, prolateral; B, retrolateral. CB: cymbium; E: embolus; T: tegulum." captionText-2="Figure 55. Maymena paquini sp. n. from Wayala Ku cave, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral detail, arrow indicates hook-like cymbial apophysis; D, palpal tibia, dorsal, arrows indicate trichobothria; E, metatarsus I; F, tarsus, arrow indicates tarsal organ. CB: cymbium; E: embolus; T: tegulum." captionText-3="Figure 56. Maymena paquini sp. n. from Wayala Ku cave, SEM. A, epigynum, ventral; B, epigynum, lateral; C, epigynum, posterior; D, epiandrous gland spigots; E, female labrum, anterior, dorsal to the right; F, labrum, lateral, dorsal to the right." captionText-4="Figure 57. Maymena paquini sp. n. from Wayala Ku cave, SEM of spinnerets. A-D, female; E, F, male. A, spinnerets; B, PLS; C, E, PMS; D, F, PLS, arrow indicates modified PLS seta. AC: aciniform gland spigot; AG: aggregate gland spigot; AGn: aggregate gland spigot nubbin; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL: flagelliform gland spigot; FLn: flagelliform gland spigot nubbin; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi-0="http://doi.org/10.5281/zenodo.3768849" figureDoi-1="http://doi.org/10.5281/zenodo.3768851" figureDoi-2="http://doi.org/10.5281/zenodo.3768853" figureDoi-3="http://doi.org/10.5281/zenodo.3768855" figureDoi-4="http://doi.org/10.5281/zenodo.3768857" httpUri-0="https://zenodo.org/record/3768849/files/figure.png" httpUri-1="https://zenodo.org/record/3768851/files/figure.png" httpUri-2="https://zenodo.org/record/3768853/files/figure.png" httpUri-3="https://zenodo.org/record/3768855/files/figure.png" httpUri-4="https://zenodo.org/record/3768857/files/figure.png" pageId="46" pageNumber="55">53-57</figureCitation>
,
<figureCitation box="[363,392,1540,1566]" captionStart="Figure 99" captionStartId="170.[140,212,1410,1432]" captionTargetBox="[140,1108,165,1390]" captionTargetId="figure@170.[140,1109,164,1390]" captionTargetPageId="170" captionText="Figure 99. Distribution map, inset map shows study area. White: Maymena paquini sp. n.; purple: Maymena kehen sp. n.; red: Gaiziapis zhizhuba sp. n.; yellow: Patu qiqi sp. n.; blue: Patu xiaoxiao sp. n." figureDoi="http://doi.org/10.5281/zenodo.3768931" httpUri="https://zenodo.org/record/3768931/files/figure.png" pageId="46" pageNumber="55">99</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="47" lastPageNumber="56" pageId="46" pageNumber="55" type="materials_examined">
<paragraph blockId="46.[140,1108,1609,1672]" pageId="46" pageNumber="55">
<emphasis bold="true" box="[140,377,1609,1635]" pageId="46" pageNumber="55">Material Examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608939562" collectingDate="2005-08-03" country="China" county="Lushui Co." elevation="910" latitude="26.13198" location="Wayala Ku" longLatPrecision="1" longitude="98.86149" municipality="Daxingdi" pageId="46" pageNumber="55" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus box="[385,496,1610,1636]" pageId="46" pageNumber="55">Holotype</typeStatus>
:
<collectingCountry box="[507,606,1610,1636]" name="China" pageId="46" pageNumber="55">CHINA</collectingCountry>
:
<collectingRegion box="[617,709,1611,1637]" country="China" name="Yunnan" pageId="46" pageNumber="55">Yunnan</collectingRegion>
:
<collectingCounty box="[720,850,1610,1636]" pageId="46" pageNumber="55">Lushui Co.</collectingCounty>
,
<collectingMunicipality box="[863,972,1610,1636]" pageId="46" pageNumber="55">Daxingdi</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03ACB915FFB17424FF4E40E0FE36FEF6:8EDA5ED8FFB1743AFC1A4317FBA2F9DD" box="[984,1107,1610,1637]" country="China" county="Lushui Co." latitude="26.13198" longLatPrecision="1" longitude="98.86149" municipality="Daxingdi" name="Wayala Ku" pageId="46" pageNumber="55" stateProvince="Yunnan">Wayala Ku</location>
(cave), km 128 rd S-228,
<geoCoordinate box="[429,575,1645,1671]" degrees="26.13198" direction="north" orientation="latitude" pageId="46" pageNumber="55" precision="1" value="26.13198">26.13198°N</geoCoordinate>
,
<geoCoordinate box="[586,725,1645,1671]" degrees="98.86149" direction="east" orientation="longitude" pageId="46" pageNumber="55" precision="1" value="98.86149">98.86149°E</geoCoordinate>
,
<quantity box="[736,813,1645,1672]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="55" unit="m" value="910.0">
<elevation box="[736,813,1645,1672]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="46" pageNumber="55" unit="m" value="910.0">910 m</elevation>
</quantity>
,
<date box="[825,995,1645,1672]" pageId="46" pageNumber="55" value="2005-08-03">
<collectingDate box="[825,995,1645,1672]" pageId="46" pageNumber="55" value="2005-08-03">3 August 2005</collectingDate>
</date>
, cave, hu-
</materialsCitation>
</paragraph>
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
man impacted and flooded, P. Paquin, D. Kavanaugh, PP-1705 (CASENT 9022598,
<collectionCode box="[140,212,203,229]" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="47" pageNumber="56">HNU</collectionCode>
), 1 ♁.
</paragraph>
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
<materialsCitation ID-GBIF-Occurrence="2608939544" box="[185,1018,237,265]" collectingDate="2005-08-03" collectionCode="CAS" country="China" county="Lushui Co." elevation="910" latitude="26.13198" location="Wayala Ku" longLatPrecision="1" longitude="98.86149" municipality="Daxingdi" pageId="47" pageNumber="56" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus box="[185,295,238,264]" pageId="47" pageNumber="56">Paratypes</typeStatus>
: [same data as holotype] (CASENT 9022595,
<collectionCode box="[838,895,237,263]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="56" type="Museum">CAS</collectionCode>
), 1 ♁,
<specimenCount box="[980,1018,237,264]" pageId="47" pageNumber="56" type="female">3 ♀</specimenCount>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939574" collectingDate="2005-08-03" collectionCode="HNU" country="China" county="Lushui Co." elevation="910" latitude="26.13198" location="Wayala Ku" longLatPrecision="1" longitude="98.86149" municipality="Daxingdi" pageId="47" pageNumber="56" specimenCount="9" specimenCount-female="9" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9022599,
<collectionCode box="[595,667,273,299]" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="47" pageNumber="56">HNU</collectionCode>
),
<specimenCount box="[690,728,273,299]" pageId="47" pageNumber="56" type="female">9 ♀</specimenCount>
, 6 juvs
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939559" collectingDate="2005-08-03" collectionCode="CAS" country="China" county="Lushui Co." elevation="910" latitude="26.13198" location="Wayala Ku" longLatPrecision="1" longitude="98.86149" municipality="Daxingdi" pageId="47" pageNumber="56" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9022596,
<collectionCode box="[394,451,308,334]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="56" type="Museum">CAS</collectionCode>
), 1 ♁,
<specimenCount box="[542,583,308,334]" pageId="47" pageNumber="56" type="female">2 ♀</specimenCount>
, 4 juvs
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939587" collectingDate="2005-08-03" collectionCode="CAS" country="China" county="Lushui Co." elevation="910" latitude="26.13198" location="Wayala Ku" longLatPrecision="1" longitude="98.86149" municipality="Daxingdi" pageId="47" pageNumber="56" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9022594,
<collectionCode box="[257,314,343,369]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="56" type="Museum">CAS</collectionCode>
),
<specimenCount box="[334,371,343,369]" pageId="47" pageNumber="56" type="female">2 ♀</specimenCount>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939543" box="[393,1026,342,370]" collectingDate="2005-08-03" collectionCode="CAS" country="China" county="Lushui Co." elevation="910" latitude="26.13198" location="Wayala Ku" longLatPrecision="1" longitude="98.86149" municipality="Daxingdi" pageId="47" pageNumber="56" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9022597,
<collectionCode box="[912,969,343,369]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="56" type="Museum">CAS</collectionCode>
),
<specimenCount box="[989,1026,343,369]" pageId="47" pageNumber="56" type="female">1 ♀</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="56" type="etymology">
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
<emphasis bold="true" box="[185,320,378,404]" pageId="47" pageNumber="56">
<emphasis box="[185,315,378,404]" inLineHeading="true" pageId="47" pageNumber="56" reason="1">Etymology</emphasis>
.
</emphasis>
Patronymic in honor of Pierre Paquin in recognition of his contributions to biospeleology.
</paragraph>
</subSubSection>
<subSubSection pageId="47" pageNumber="56" type="diagnosis">
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
<emphasis bold="true" box="[185,312,448,474]" pageId="47" pageNumber="56">
<emphasis box="[185,307,448,474]" inLineHeading="true" pageId="47" pageNumber="56" reason="1">Diagnosis</emphasis>
.
</emphasis>
Male distinguished from other
<taxonomicName authorityName="Gertsch" authorityYear="1960" box="[679,787,449,474]" class="Arachnida" family="Mysmenidae" genus="Maymena" kingdom="Animalia" order="Araneae" pageId="47" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis box="[679,787,449,474]" italics="true" pageId="47" pageNumber="56">Maymena</emphasis>
</taxonomicName>
for which males are known (i.e., American species) by the elongate palpal tibia (
<figureCitation box="[751,856,483,510]" captionStart="Figure 54" captionStartId="126.[140,212,1340,1362]" captionTargetBox="[150,1096,173,1309]" captionTargetId="figure@126.[140,1109,164,1320]" captionTargetPageId="126" captionText="Figure 54. Maymena paquini sp. n. from Wayala Ku cave, male palp. A, prolateral; B, retrolateral. CB: cymbium; E: embolus; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768851" httpUri="https://zenodo.org/record/3768851/files/figure.png" pageId="47" pageNumber="56">Fig. 54A</figureCitation>
; compare to
<bibRefCitation author="Gertsch WJ" pageId="47" pageNumber="56" pagination="1 - 40" refId="ref45779" refString="Gertsch WJ (1960) Descriptions of American spiders of the family Symphytognathidae. American Museum Novitates 1981: 1 - 40." type="journal article" year="1960">Gertsch, 1960</bibRefCitation>
: fig. 51).
</paragraph>
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
Female distinguished from American species by the shape of the epigynal scape, which is relatively long and narrow (
<figureCitation box="[547,629,589,616]" captionStart="Figure 50" captionStartId="122.[140,212,958,980]" captionTargetBox="[140,1108,165,938]" captionTargetId="figure@122.[140,1109,164,939]" captionTargetPageId="122" captionText="Figure 50. A-C, Chanea suukyii sp. n. from Maku ridge; D-F, Maymena paquini sp. n. from Wayala Ku cave; G, H; Maymena kehen sp. n. from Lishadi, unlabeled arrow indicates diagnostic notch in scape; I, Gaiziapis zhizhuba sp. n. from Nankang Yakou.A, D, G, epigynum, ventral veiw; B, F, H, I, cleared vulva, dorsal view; C, male palp, cleared and expanded. BC: booklung cover; BH: basal hematodocha; CD: copulatory duct; FD: fertilization duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768845" httpUri="https://zenodo.org/record/3768845/files/figure.png" pageId="47" pageNumber="56">Figs 50</figureCitation>
D-F, 56A-C); distinguished from
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[1006,1108,589,615]" class="Arachnida" family="Mysmenidae" genus="Maymena" kingdom="Animalia" order="Araneae" pageId="47" pageNumber="56" phylum="Arthropoda" rank="species" species="kehen">
<emphasis box="[1006,1108,589,615]" italics="true" pageId="47" pageNumber="56">M. kehen</emphasis>
</taxonomicName>
by the wider scape, wider separation of the spermathecae, and by the lateral margin of the scape, which is uniquely notched in
<taxonomicName authorityName="Miller &amp; Griswold &amp; Yin" authorityYear="2009" box="[538,638,660,686]" class="Arachnida" family="Mysmenidae" genus="Maymena" kingdom="Animalia" order="Araneae" pageId="47" pageNumber="56" phylum="Arthropoda" rank="species" species="kehen">
<emphasis box="[538,638,660,686]" italics="true" pageId="47" pageNumber="56">M. kehen</emphasis>
</taxonomicName>
(
<figureCitation box="[652,733,659,686]" captionStart="Figure 50" captionStartId="122.[140,212,958,980]" captionTargetBox="[140,1108,165,938]" captionTargetId="figure@122.[140,1109,164,939]" captionTargetPageId="122" captionText="Figure 50. A-C, Chanea suukyii sp. n. from Maku ridge; D-F, Maymena paquini sp. n. from Wayala Ku cave; G, H; Maymena kehen sp. n. from Lishadi, unlabeled arrow indicates diagnostic notch in scape; I, Gaiziapis zhizhuba sp. n. from Nankang Yakou.A, D, G, epigynum, ventral veiw; B, F, H, I, cleared vulva, dorsal view; C, male palp, cleared and expanded. BC: booklung cover; BH: basal hematodocha; CD: copulatory duct; FD: fertilization duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768845" httpUri="https://zenodo.org/record/3768845/files/figure.png" pageId="47" pageNumber="56">Fig. 50</figureCitation>
G-H). While distinctions between the two known Chinese
<taxonomicName authorityName="Gertsch" authorityYear="1960" box="[410,516,696,721]" class="Arachnida" family="Mysmenidae" genus="Maymena" kingdom="Animalia" order="Araneae" pageId="47" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis box="[410,516,696,721]" italics="true" pageId="47" pageNumber="56">Maymena</emphasis>
</taxonomicName>
species are subtle, they do seem to be consistent.
</paragraph>
</subSubSection>
<subSubSection lastPageId="48" lastPageNumber="57" pageId="47" pageNumber="56" type="description">
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
<emphasis bold="true" box="[185,335,729,755]" pageId="47" pageNumber="56">
<emphasis box="[185,330,729,755]" inLineHeading="true" pageId="47" pageNumber="56" reason="1">Description</emphasis>
.
</emphasis>
Carapace pale orange. Labrum not swollen, weakly sclerotized, short labral tongue with flat ventral margin (
<figureCitation box="[603,689,765,792]" captionStart="Figure 56" captionStartId="128.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@128.[140,1108,164,1184]" captionTargetPageId="128" captionText="Figure 56. Maymena paquini sp. n. from Wayala Ku cave, SEM. A, epigynum, ventral; B, epigynum, lateral; C, epigynum, posterior; D, epiandrous gland spigots; E, female labrum, anterior, dorsal to the right; F, labrum, lateral, dorsal to the right." figureDoi="http://doi.org/10.5281/zenodo.3768855" httpUri="https://zenodo.org/record/3768855/files/figure.png" pageId="47" pageNumber="56">Fig. 56</figureCitation>
E-F). Sternum orange. Legs orange. Femoral spots on legs I and II in female, leg I only in male. Tarsal organs set distal to proximal margin of tarsus (ratio of distance from proximal margin of tarsus to tarsal organ over total tarsus length
<emphasis box="[491,522,872,897]" italics="true" pageId="47" pageNumber="56">ca.</emphasis>
0.15,
<figureCitation box="[601,703,871,898]" captionStart="Figure 55" captionStartId="127.[140,212,1203,1225]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@127.[140,1109,164,1184]" captionTargetPageId="127" captionText="Figure 55. Maymena paquini sp. n. from Wayala Ku cave, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral detail, arrow indicates hook-like cymbial apophysis; D, palpal tibia, dorsal, arrows indicate trichobothria; E, metatarsus I; F, tarsus, arrow indicates tarsal organ. CB: cymbium; E: embolus; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768853" httpUri="https://zenodo.org/record/3768853/files/figure.png" pageId="47" pageNumber="56">Fig. 55F</figureCitation>
). Abdomen light gray with pair of dorsolateral dark gray patches (
<figureCitation box="[490,574,906,933]" captionStart="Figure 53" captionStartId="125.[140,212,1162,1184]" captionTargetBox="[159,1089,183,1131]" captionTargetId="figure@125.[140,1108,164,1143]" captionTargetPageId="125" captionText="Figure 53. Maymena paquini sp. n. from Wayala Ku cave. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, female, posterior." figureDoi="http://doi.org/10.5281/zenodo.3768849" httpUri="https://zenodo.org/record/3768849/files/figure.png" pageId="47" pageNumber="56">Fig. 53</figureCitation>
).
</paragraph>
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
Male palp: Palpal patella and tibia elongate (
<figureCitation box="[698,802,941,968]" captionStart="Figure 54" captionStartId="126.[140,212,1340,1362]" captionTargetBox="[150,1096,173,1309]" captionTargetId="figure@126.[140,1109,164,1320]" captionTargetPageId="126" captionText="Figure 54. Maymena paquini sp. n. from Wayala Ku cave, male palp. A, prolateral; B, retrolateral. CB: cymbium; E: embolus; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768851" httpUri="https://zenodo.org/record/3768851/files/figure.png" pageId="47" pageNumber="56">Fig. 54A</figureCitation>
); palpal tibia with two trichobothria (
<figureCitation box="[281,391,976,1003]" captionStart="Figure 55" captionStartId="127.[140,212,1203,1225]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@127.[140,1109,164,1184]" captionTargetPageId="127" captionText="Figure 55. Maymena paquini sp. n. from Wayala Ku cave, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral detail, arrow indicates hook-like cymbial apophysis; D, palpal tibia, dorsal, arrows indicate trichobothria; E, metatarsus I; F, tarsus, arrow indicates tarsal organ. CB: cymbium; E: embolus; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768853" httpUri="https://zenodo.org/record/3768853/files/figure.png" pageId="47" pageNumber="56">Fig. 55D</figureCitation>
). Cymbium covers ventral part of palpal bulb, dorsal part exposed (
<figureCitation box="[222,305,1011,1038]" captionStart="Figure 55" captionStartId="127.[140,212,1203,1225]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@127.[140,1109,164,1184]" captionTargetPageId="127" captionText="Figure 55. Maymena paquini sp. n. from Wayala Ku cave, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral detail, arrow indicates hook-like cymbial apophysis; D, palpal tibia, dorsal, arrows indicate trichobothria; E, metatarsus I; F, tarsus, arrow indicates tarsal organ. CB: cymbium; E: embolus; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768853" httpUri="https://zenodo.org/record/3768853/files/figure.png" pageId="47" pageNumber="56">Fig. 55</figureCitation>
A-B). Hook-like apophysis on prolateral face of cymbium (
<figureCitation box="[987,1094,1011,1038]" captionStart="Figure 55" captionStartId="127.[140,212,1203,1225]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@127.[140,1109,164,1184]" captionTargetPageId="127" captionText="Figure 55. Maymena paquini sp. n. from Wayala Ku cave, SEM of male. A, palp, prolateral; B, palp, retrolateral; C, palp, prolateral detail, arrow indicates hook-like cymbial apophysis; D, palpal tibia, dorsal, arrows indicate trichobothria; E, metatarsus I; F, tarsus, arrow indicates tarsal organ. CB: cymbium; E: embolus; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768853" httpUri="https://zenodo.org/record/3768853/files/figure.png" pageId="47" pageNumber="56">Fig. 55C</figureCitation>
). Embolus long and filiform with proximal origin (
<figureCitation box="[697,800,1047,1074]" captionStart="Figure 54" captionStartId="126.[140,212,1340,1362]" captionTargetBox="[150,1096,173,1309]" captionTargetId="figure@126.[140,1109,164,1320]" captionTargetPageId="126" captionText="Figure 54. Maymena paquini sp. n. from Wayala Ku cave, male palp. A, prolateral; B, retrolateral. CB: cymbium; E: embolus; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768851" httpUri="https://zenodo.org/record/3768851/files/figure.png" pageId="47" pageNumber="56">Fig. 54B</figureCitation>
).
</paragraph>
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
<taxonomicName authority=": Epigynum" authorityName="Epigynum" box="[185,381,1082,1108]" class="Polychaeta" family="Polychaeta (awaiting allocation)" genus="Vulva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polychaeta (awaiting allocation)" pageId="47" pageNumber="56" phylum="Annelida" rank="genus">Vulva: Epigynum</taxonomicName>
a scape extending nearly to tracheal spiracle (
<figureCitation box="[890,1028,1082,1109]" captionStart="Figure 53" captionStartId="125.[140,212,1162,1184]" captionTargetBox="[159,1089,183,1131]" captionTargetId="figure@125.[140,1108,164,1143]" captionTargetPageId="125" captionText="Figure 53. Maymena paquini sp. n. from Wayala Ku cave. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, female, posterior." figureDoi="http://doi.org/10.5281/zenodo.3768849" httpUri="https://zenodo.org/record/3768849/files/figure.png" pageId="47" pageNumber="56">Fig. 53B, D</figureCitation>
). Spermathecae round, ducts arise from anteroectal part, run posteriorly; fertilization ducts diverge from copulatory ducts, run anteromesally; copulatory ducts loop near base of scape (
<figureCitation box="[215,315,1187,1214]" captionStart="Figure 50" captionStartId="122.[140,212,958,980]" captionTargetBox="[140,1108,165,938]" captionTargetId="figure@122.[140,1109,164,939]" captionTargetPageId="122" captionText="Figure 50. A-C, Chanea suukyii sp. n. from Maku ridge; D-F, Maymena paquini sp. n. from Wayala Ku cave; G, H; Maymena kehen sp. n. from Lishadi, unlabeled arrow indicates diagnostic notch in scape; I, Gaiziapis zhizhuba sp. n. from Nankang Yakou.A, D, G, epigynum, ventral veiw; B, F, H, I, cleared vulva, dorsal view; C, male palp, cleared and expanded. BC: booklung cover; BH: basal hematodocha; CD: copulatory duct; FD: fertilization duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768845" httpUri="https://zenodo.org/record/3768845/files/figure.png" pageId="47" pageNumber="56">Fig. 50F</figureCitation>
), terminate in paired atria near middle of scape (
<figureCitation box="[864,946,1187,1214]" captionStart="Figure 56" captionStartId="128.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@128.[140,1108,164,1184]" captionTargetPageId="128" captionText="Figure 56. Maymena paquini sp. n. from Wayala Ku cave, SEM. A, epigynum, ventral; B, epigynum, lateral; C, epigynum, posterior; D, epiandrous gland spigots; E, female labrum, anterior, dorsal to the right; F, labrum, lateral, dorsal to the right." figureDoi="http://doi.org/10.5281/zenodo.3768855" httpUri="https://zenodo.org/record/3768855/files/figure.png" pageId="47" pageNumber="56">Fig. 56</figureCitation>
A-C).
</paragraph>
<paragraph blockId="47.[140,1108,167,1672]" pageId="47" pageNumber="56">
Male (CASENT 9022595): Total length 2.20, carapace 1.06 long, 0.95 wide, clypeus 0.25, sternum 0.56 long, 0.60 wide, coxa IV separated by 0.44 times their width. Leg I: femur d1, p2, r2, patella d2, tibia d1, p3, r1, metatarsus p1; Leg II: femur d1, p1, r2, patella d2, tibia d1, p2-3, r1, metatarsus with all prolateral setae strong; Leg III: femur d1, p1, r1, patella d2, tibia d1, p1, r1, metatarsus p1; Leg IV: femur d1, p1, r1, patella d2, tibia d1, p2, r1. Metatarsal macrosetae kinked proximal to center. Metatarsal trichobothria: TmI: 0.24; TmII: 0.24; TmIII: 0.33. Leg measurements: see Appendix A. Epiandrous gland spigots in four clusters of two-three spigots each (
<figureCitation captionStart="Figure 56" captionStartId="128.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@128.[140,1108,164,1184]" captionTargetPageId="128" captionText="Figure 56. Maymena paquini sp. n. from Wayala Ku cave, SEM. A, epigynum, ventral; B, epigynum, lateral; C, epigynum, posterior; D, epiandrous gland spigots; E, female labrum, anterior, dorsal to the right; F, labrum, lateral, dorsal to the right." figureDoi="http://doi.org/10.5281/zenodo.3768855" httpUri="https://zenodo.org/record/3768855/files/figure.png" pageId="47" pageNumber="56">Fig. 56D</figureCitation>
). Posterior lateral spinnerets with flagelliform and aggregate gland spigots reduced to nubbins (
<figureCitation box="[278,379,1539,1566]" captionStart="Figure 57" captionStartId="129.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@129.[140,1109,164,1186]" captionTargetPageId="129" captionText="Figure 57. Maymena paquini sp. n. from Wayala Ku cave, SEM of spinnerets. A-D, female; E, F, male. A, spinnerets; B, PLS; C, E, PMS; D, F, PLS, arrow indicates modified PLS seta. AC: aciniform gland spigot; AG: aggregate gland spigot; AGn: aggregate gland spigot nubbin; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL: flagelliform gland spigot; FLn: flagelliform gland spigot nubbin; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi="http://doi.org/10.5281/zenodo.3768857" httpUri="https://zenodo.org/record/3768857/files/figure.png" pageId="47" pageNumber="56">Fig. 57F</figureCitation>
).
</paragraph>
<paragraph blockId="47.[140,1108,167,1672]" lastBlockId="48.[140,1108,167,334]" lastPageId="48" lastPageNumber="57" pageId="47" pageNumber="56">
Female (CASENT 9022595): Total length 2.50 carapace 1.18 long, 0.99 wide, clypeus 0.23, sternum 0.66 long, 0.64 wide, coxa IV separated by 0.59 times their width. Leg I: femur d1, p1-3, r2, patella d2, tibia d1, p3, r1, metatarsus with all prolateral setae strong; Leg II: femur d1, p1, r1-2, patella d2, tibia d1, p2-3, r1, metatarsus with all prolateral setae strong; Leg III: femur d1, p1, r1, patella d2, tibia d1, p2, r1, metatarsus p1; Leg IV: femur d1, r0-1, patella d2, tibia d1, p2, r1. Metatarsal trichobothria: TmI: 0.25; TmII: 0.27; TmIII: 0.34. Leg measurements: see Appendix A. Spinnerets (
<figureCitation box="[306,387,307,334]" captionStart="Figure 57" captionStartId="129.[140,212,1206,1228]" captionTargetBox="[140,1108,165,1186]" captionTargetId="figure@129.[140,1109,164,1186]" captionTargetPageId="129" captionText="Figure 57. Maymena paquini sp. n. from Wayala Ku cave, SEM of spinnerets. A-D, female; E, F, male. A, spinnerets; B, PLS; C, E, PMS; D, F, PLS, arrow indicates modified PLS seta. AC: aciniform gland spigot; AG: aggregate gland spigot; AGn: aggregate gland spigot nubbin; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL: flagelliform gland spigot; FLn: flagelliform gland spigot nubbin; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi="http://doi.org/10.5281/zenodo.3768857" httpUri="https://zenodo.org/record/3768857/files/figure.png" pageId="48" pageNumber="57">Fig. 57</figureCitation>
A-D).
</paragraph>
</subSubSection>
</treatment>
</document>
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