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<document ID-DOI="http://doi.org/10.5281/zenodo.3792218" ID-GBIF-Dataset="6bed073a-175c-42e6-bb99-645c99d55b4f" ID-GBIF-Taxon="163750555" ID-ISSN="13132970" ID-Zenodo-Dep="3792218" ID-ZooBank="C631A347-306E-4773-84A4-E4712329186B" LSID="urn:lsid:zoobank.org:act:3745AD70-626C-4FFB-9B20-B60D83C8995C" checkinTime="1587984202003" checkinUser="plazi" docAuthor="Miller, Jeremy, Griswold, Charles &amp; Yin, Chang" docDate="2009" docId="03ACB915FFA07454FF4E463BFC14FC1D" docLanguage="en" docName="ZK_article_1993.pdf" docOrigin="ZooKeys 11 (11)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Crassignatha quanqu Miller, Griswold &amp; Yin 2009, sp. n." docType="treatment" docUuid="3745AD70-626C-4FFB-9B20-B60D83C8995C" docUuidSource="ZooBank" docVersion="9" lastPageId="64" lastPageNumber="73" masterDocId="FF95C16DFF9F7414FFC2455DFFF1FFB8" masterDocTitle="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers" masterLastPageNumber="195" masterPageNumber="9" pageId="63" pageNumber="72" updateTime="1635491590422" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): Systematics and diversity of micro-orbweavers</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Jeremy</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Griswold, Charles</mods:namePart>
<mods:affiliation>California Academy of Sciences ,, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Chang</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University ,, China</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-05-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>11</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>11</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>9</mods:start>
<mods:end>195</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.11.160</mods:identifier>
<mods:identifier type="GBIF-Dataset">6bed073a-175c-42e6-bb99-645c99d55b4f</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576459</mods:identifier>
<mods:identifier type="ZooBank">C631A347-306E-4773-84A4-E4712329186B</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792218" ID-GBIF-Taxon="163750555" ID-Zenodo-Dep="3792218" LSID="urn:lsid:zoobank.org:act:3745AD70-626C-4FFB-9B20-B60D83C8995C" httpUri="http://treatment.plazi.org/id/03ACB915FFA07454FF4E463BFC14FC1D" lastPageId="64" lastPageNumber="73" pageId="63" pageNumber="72">
<subSubSection box="[140,756,870,897]" pageId="63" pageNumber="72" type="nomenclature">
<paragraph blockId="63.[140,977,870,968]" box="[140,756,870,897]" pageId="63" pageNumber="72">
<heading box="[140,756,870,897]" fontSize="11" level="2" pageId="63" pageNumber="72" reason="3">
<emphasis bold="true" box="[140,756,870,897]" pageId="63" pageNumber="72">
<taxonomicName authority="Miller, Griswold &amp; Yin" authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[140,679,870,897]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="species" species="quanqu" status="sp. n.">
<emphasis bold="true" box="[140,394,870,897]" italics="true" pageId="63" pageNumber="72">
<emphasis box="[140,295,870,897]" inLineHeading="true" pageId="63" pageNumber="72" reason="1">Crassignatha</emphasis>
quanqu
</emphasis>
Miller, Griswold &amp; Yin
</taxonomicName>
,
<taxonomicNameLabel box="[691,756,871,897]" pageId="63" pageNumber="72" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="72" type="description">
<paragraph blockId="63.[140,977,870,968]" box="[140,977,906,933]" pageId="63" pageNumber="72">
<uri box="[140,977,906,933]" pageId="63" pageNumber="72">
urn:lsid:zoobank.org:act:
<uuid box="[421,977,906,933]" pageId="63" pageNumber="72">3745AD70-626C-4FFB-9B20-B60D83C8995C</uuid>
</uri>
</paragraph>
<paragraph blockId="63.[140,977,870,968]" box="[140,478,941,968]" pageId="63" pageNumber="72">
<figureCitation box="[140,222,941,967]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="63" pageNumber="72">Figs 76</figureCitation>
E-G,
<figureCitation box="[284,312,941,967]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="63" pageNumber="72">79</figureCitation>
E-F,
<figureCitation box="[367,437,941,968]" captionStart-0="Figure 83" captionStart-1="Figure 84" captionStart-2="Figure 85" captionStartId-0="155.[140,212,1138,1160]" captionStartId-1="156.[140,212,1261,1283]" captionStartId-2="157.[140,212,1202,1224]" captionTargetBox-0="[152,1100,178,1110]" captionTargetBox-1="[150,1103,167,1233]" captionTargetBox-2="[140,1108,165,1182]" captionTargetId-0="figure@155.[140,1108,164,1118]" captionTargetId-1="figure@156.[140,1109,164,1241]" captionTargetId-2="figure@157.[140,1108,164,1182]" captionTargetPageId-0="155" captionTargetPageId-1="156" captionTargetPageId-2="157" captionText-0="Figure 83. Crassignatha quanqu sp. n. from Bang Bie village. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." captionText-1="Figure 84. Crassignatha quanqu sp. n. from Bang Bie village, male palp. A, prolateral; B, retrolateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum." captionText-2="Figure 85. Crassignatha quanqu sp. n. from Bang Bie village, SEM of spinnerets. A-E, female; F, male. A, spinnerets; B, ALS; C, PMS; D, F, PLS; E, base of ALS showing lack of colulus. AC: aciniform gland spigot; AG: aggregate gland spigot; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL:flagelliform gland spigot; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi-0="http://doi.org/10.5281/zenodo.3768902" figureDoi-1="http://doi.org/10.5281/zenodo.3768904" figureDoi-2="http://doi.org/10.5281/zenodo.3768906" httpUri-0="https://zenodo.org/record/3768902/files/figure.png" httpUri-1="https://zenodo.org/record/3768904/files/figure.png" httpUri-2="https://zenodo.org/record/3768906/files/figure.png" pageId="63" pageNumber="72">83-85</figureCitation>
,
<figureCitation box="[449,478,941,967]" captionStart="Figure 96" captionStartId="167.[140,212,1410,1432]" captionTargetBox="[140,1108,165,1390]" captionTargetId="figure@167.[140,1109,164,1390]" captionTargetPageId="167" captionText="Figure 96. Distribution map, inset map shows study area. Black: Wendilgarda muji sp. n.; yellow: Mysmena changouzi sp. n.; blue: Mysmena bizi sp. n.; red: Mosu huogou sp. n.; purple: Crassignatha quanqu sp. n.; white: Crassignatha gudu sp. n." figureDoi="http://doi.org/10.5281/zenodo.3768924" httpUri="https://zenodo.org/record/3768924/files/figure.png" pageId="63" pageNumber="72">96</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="72" type="materials_examined">
<paragraph blockId="63.[140,1108,1011,1672]" pageId="63" pageNumber="72">
<emphasis bold="true" box="[140,377,1011,1037]" pageId="63" pageNumber="72">
<emphasis box="[140,241,1011,1037]" inLineHeading="true" pageId="63" pageNumber="72" reason="1">Material</emphasis>
Examined.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608939436" collectingDate="2005-05-24" collectionCode="HNU" collectorName="Route S &amp; C. Griswold" country="China" county="Longling Co." elevation="1553" latitude="24.81333" location="Bang Bie village" longLatPrecision="1" longitude="98.8328" municipality="Zhen'an Township" pageId="63" pageNumber="72" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus box="[385,496,1012,1038]" pageId="63" pageNumber="72">Holotype</typeStatus>
:
<collectingCountry box="[508,607,1012,1038]" name="China" pageId="63" pageNumber="72">CHINA</collectingCountry>
:
<collectingRegion box="[618,710,1012,1038]" country="China" name="Yunnan" pageId="63" pageNumber="72">Yunnan</collectingRegion>
:
<collectingCounty box="[721,876,1012,1038]" pageId="63" pageNumber="72">Longling Co.</collectingCounty>
,
<collectingMunicipality box="[889,1103,1012,1038]" pageId="63" pageNumber="72">Zhenan Township</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03ACB915FFA07454FF4E463BFC14FC1D:8EDA5ED8FFA0742BFF4E414AFECEFB89" box="[140,319,1047,1073]" country="China" county="Longling Co." latitude="24.81333" longLatPrecision="1" longitude="98.8328" municipality="Zhen'an Township" name="Bang Bie village" pageId="63" pageNumber="72" stateProvince="Yunnan">Bang Bie village</location>
at stream at km 6.8 on
<collectorName box="[586,676,1047,1073]" pageId="63" pageNumber="72">Route S</collectorName>
317,
<geoCoordinate box="[733,878,1047,1073]" degrees="24.81333" direction="north" orientation="latitude" pageId="63" pageNumber="72" precision="1" value="24.81333">24.81333°N</geoCoordinate>
,
<geoCoordinate box="[889,1029,1047,1073]" degrees="98.83280" direction="east" orientation="longitude" pageId="63" pageNumber="72" precision="1" value="98.8328">98.83280°E</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5525" metricValueMax="1.56" metricValueMin="1.545" pageId="63" pageNumber="72" unit="m" value="1552.5" valueMax="1560.0" valueMin="1545.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.5525" metricValueMax="1.56" metricValueMin="1.545" pageId="63" pageNumber="72" unit="m" value="1552.5" valueMax="1560.0" valueMin="1545.0">1545- 1560 m</elevation>
</quantity>
,
<date box="[248,409,1082,1109]" pageId="63" pageNumber="72" value="2005-05-24">
<collectingDate box="[248,409,1082,1109]" pageId="63" pageNumber="72" value="2005-05-24">24 May 2005</collectingDate>
</date>
, shaded embankments along stream, dusting webs in understory,
<collectorName box="[208,350,1117,1143]" pageId="63" pageNumber="72">C. Griswold</collectorName>
, CGY119 (CASENT 9029323,
<collectionCode box="[718,790,1118,1144]" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="63" pageNumber="72">HNU</collectionCode>
), 1 ♁.
</materialsCitation>
</paragraph>
<paragraph blockId="63.[140,1108,1011,1672]" pageId="63" pageNumber="72">
<materialsCitation ID-GBIF-Occurrence="2608939444" box="[185,1018,1152,1180]" collectingDate="2005-05-24" collectionCode="CAS" collectorName="Route S &amp; C. Griswold" country="China" county="Longling Co." elevation="1553" latitude="24.81333" location="Bang Bie village" longLatPrecision="1" longitude="98.8328" municipality="Zhen'an Township" pageId="63" pageNumber="72" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus box="[185,295,1153,1179]" pageId="63" pageNumber="72">Paratypes</typeStatus>
: [same data as holotype] (CASENT 9022387,
<collectionCode box="[838,895,1153,1179]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="63" pageNumber="72" type="Museum">CAS</collectionCode>
), 1 ♁,
<specimenCount box="[980,1018,1153,1179]" pageId="63" pageNumber="72" type="female">1 ♀</specimenCount>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608939439" collectingDate="2005-05-24" collectionCode="HNU" collectorName="Route S &amp; C. Griswold" country="China" county="Longling Co." elevation="1553" latitude="24.81333" location="Bang Bie village" longLatPrecision="1" longitude="98.8328" municipality="Zhen'an Township" pageId="63" pageNumber="72" specimenCount="2" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
[same data as holotype] (CASENT 9022388,
<collectionCode box="[586,658,1188,1214]" country="China" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="63" pageNumber="72">HNU</collectionCode>
), 1 ♁,
<specimenCount box="[737,774,1188,1214]" pageId="63" pageNumber="72" type="female">1 ♀</specimenCount>
,
<specimenCount box="[796,859,1188,1214]" pageId="63" pageNumber="72" type="generic">1 juv.</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="72" type="etymology">
<paragraph blockId="63.[140,1108,1011,1672]" pageId="63" pageNumber="72">
<emphasis bold="true" box="[185,320,1223,1249]" pageId="63" pageNumber="72">
<emphasis box="[185,315,1223,1249]" inLineHeading="true" pageId="63" pageNumber="72" reason="1">Etymology</emphasis>
.
</emphasis>
Formed from the Chinese word for twisted (quán qū), referring to the shape of the embolus in the male palp.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="72" type="diagnosis">
<paragraph blockId="63.[140,1108,1011,1672]" pageId="63" pageNumber="72">
<emphasis bold="true" box="[185,312,1293,1319]" pageId="63" pageNumber="72">
<emphasis box="[185,307,1293,1319]" inLineHeading="true" pageId="63" pageNumber="72" reason="1">Diagnosis</emphasis>
.
</emphasis>
Male distinguished by the unique embolus shape, which is a rigid spiral ribbon making a single turn, not simply tapered but with a narrow waste near the midpoint (
<figureCitation box="[262,366,1364,1390]" captionStart="Figure 84" captionStartId="156.[140,212,1261,1283]" captionTargetBox="[150,1103,167,1233]" captionTargetId="figure@156.[140,1109,164,1241]" captionTargetPageId="156" captionText="Figure 84. Crassignatha quanqu sp. n. from Bang Bie village, male palp. A, prolateral; B, retrolateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768904" httpUri="https://zenodo.org/record/3768904/files/figure.png" pageId="63" pageNumber="72">Fig. 84A</figureCitation>
).
</paragraph>
<paragraph blockId="63.[140,1108,1011,1672]" pageId="63" pageNumber="72">
Female distinguished from other
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[558,696,1399,1425]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis box="[558,696,1399,1425]" italics="true" pageId="63" pageNumber="72">Crassignatha</emphasis>
</taxonomicName>
species except
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[864,965,1399,1425]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="species" species="yinzhi">
<emphasis box="[864,965,1399,1425]" italics="true" pageId="63" pageNumber="72">C. yinzhi</emphasis>
</taxonomicName>
by the distal lobe of the scape, which is more narrow than and projects well beyond the basal lobe (
<figureCitation box="[147,255,1469,1495]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="63" pageNumber="72">Figs 76G</figureCitation>
,
<figureCitation box="[267,297,1469,1495]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="63" pageNumber="72">79</figureCitation>
E-F); other
<taxonomicName authorityName="Wunderlich" authorityYear="1995" box="[430,568,1469,1495]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis box="[430,568,1469,1495]" italics="true" pageId="63" pageNumber="72">Crassignatha</emphasis>
</taxonomicName>
species either have the lobes not well differentiated or they are the same width (
<figureCitation box="[500,602,1504,1531]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="63" pageNumber="72">Fig. 79A</figureCitation>
); distinguished from
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[838,938,1504,1531]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="species" species="yinzhi">
<emphasis box="[838,938,1504,1531]" italics="true" pageId="63" pageNumber="72">C. yinzhi</emphasis>
</taxonomicName>
by the orientation of the scape, projecting ventrally in
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[591,708,1540,1566]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="species" species="quanqu">
<emphasis box="[591,708,1540,1566]" italics="true" pageId="63" pageNumber="72">C. quanqu</emphasis>
</taxonomicName>
(
<figureCitation box="[722,819,1539,1566]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="63" pageNumber="72">Figs 76F</figureCitation>
,
<figureCitation box="[830,879,1540,1566]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="63" pageNumber="72">79F</figureCitation>
), posteroventrally in
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[140,243,1575,1601]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="species" species="yinzhi">
<emphasis box="[140,243,1575,1601]" italics="true" pageId="63" pageNumber="72">C. yinzhi</emphasis>
</taxonomicName>
(
<figureCitation box="[258,368,1575,1601]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="63" pageNumber="72">Figs 79D</figureCitation>
,
<figureCitation box="[381,431,1575,1601]" captionStart="Figure 80" captionStartId="152.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@152.[140,1109,164,1270]" captionTargetPageId="152" captionText="Figure 80. Crassignatha yinzhi from Xiao Hei Shan Nature Reserve. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." figureDoi="http://doi.org/10.5281/zenodo.3813021" httpUri="https://zenodo.org/record/3813021/files/figure.png" pageId="63" pageNumber="72">80B</figureCitation>
), and elements of somatic morphology including abdomen color and shape, and the more distinct division between the cephalic and thoracic regions in
<taxonomicName authorityName="Miller, Griswold &amp; Yin" authorityYear="2009" box="[237,339,1645,1672]" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="72" phylum="Arthropoda" rank="species" species="yinzhi">
<emphasis box="[237,339,1645,1672]" italics="true" pageId="63" pageNumber="72">C. yinzhi</emphasis>
</taxonomicName>
(contrast
<figureCitation box="[453,550,1645,1671]" captionStart="Figure 80" captionStartId="152.[140,212,1290,1312]" captionTargetBox="[140,1108,165,1270]" captionTargetId="figure@152.[140,1109,164,1270]" captionTargetPageId="152" captionText="Figure 80. Crassignatha yinzhi from Xiao Hei Shan Nature Reserve. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." figureDoi="http://doi.org/10.5281/zenodo.3813021" httpUri="https://zenodo.org/record/3813021/files/figure.png" pageId="63" pageNumber="72">Fig. 80B</figureCitation>
with
<figureCitation box="[616,717,1645,1671]" captionStart="Figure 83" captionStartId="155.[140,212,1138,1160]" captionTargetBox="[152,1100,178,1110]" captionTargetId="figure@155.[140,1108,164,1118]" captionTargetPageId="155" captionText="Figure 83. Crassignatha quanqu sp. n. from Bang Bie village. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." figureDoi="http://doi.org/10.5281/zenodo.3768902" httpUri="https://zenodo.org/record/3768902/files/figure.png" pageId="63" pageNumber="72">Fig. 83B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="64" pageNumber="73" type="description">
<paragraph blockId="64.[140,1108,166,933]" pageId="64" pageNumber="73">
<emphasis bold="true" box="[185,335,166,192]" pageId="64" pageNumber="73">
<emphasis box="[185,330,166,192]" inLineHeading="true" pageId="64" pageNumber="73" reason="1">Description</emphasis>
.
</emphasis>
Carapace orange-brown with small tubercles and sulci. Sternum dark brown. Legs orange. Abdomen subspherical, pale orange, with numerous small sclerotized patches, some baring long setae, male with single orange scutum laterally and posteriorly, female without sclerite around spinnerets (
<figureCitation box="[803,886,272,299]" captionStart="Figure 83" captionStartId="155.[140,212,1138,1160]" captionTargetBox="[152,1100,178,1110]" captionTargetId="figure@155.[140,1108,164,1118]" captionTargetPageId="155" captionText="Figure 83. Crassignatha quanqu sp. n. from Bang Bie village. A, male, lateral; B, female, lateral; C, female, dorsal; D, female, ventral; E, male leg II, prolateral." figureDoi="http://doi.org/10.5281/zenodo.3768902" httpUri="https://zenodo.org/record/3768902/files/figure.png" pageId="64" pageNumber="73">Fig. 83</figureCitation>
).
</paragraph>
<paragraph blockId="64.[140,1108,166,933]" pageId="64" pageNumber="73">
Male palp: Median apophysis with two tapered distal processes. Embolus a moderately long, rigid spiral ribbon making a single turn, shape complex, not simply tapered (
<figureCitation box="[148,252,378,404]" captionStart="Figure 84" captionStartId="156.[140,212,1261,1283]" captionTargetBox="[150,1103,167,1233]" captionTargetId="figure@156.[140,1109,164,1241]" captionTargetPageId="156" captionText="Figure 84. Crassignatha quanqu sp. n. from Bang Bie village, male palp. A, prolateral; B, retrolateral. CB: cymbium; CT: cymbial tooth; E: embolus; EM: embolic membrane; MA: median apophysis; T: tegulum." figureDoi="http://doi.org/10.5281/zenodo.3768904" httpUri="https://zenodo.org/record/3768904/files/figure.png" pageId="64" pageNumber="73">Fig. 84A</figureCitation>
).
</paragraph>
<paragraph blockId="64.[140,1108,166,933]" pageId="64" pageNumber="73">
<taxonomicName box="[185,253,413,439]" class="Polychaeta" family="Polychaeta (awaiting allocation)" genus="Vulva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polychaeta (awaiting allocation)" pageId="64" pageNumber="16" phylum="Annelida" rank="genus">Vulva</taxonomicName>
: Scape projects ventrally (
<figureCitation box="[549,649,413,439]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="64" pageNumber="73">Figs 76F</figureCitation>
,
<figureCitation box="[661,711,413,439]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="64" pageNumber="73">79F</figureCitation>
), with narrow distal lobe on wider basal lobe (
<figureCitation box="[264,371,448,474]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="64" pageNumber="73">Figs 76G</figureCitation>
,
<figureCitation box="[383,434,449,475]" captionStart="Figure 79" captionStartId="151.[140,212,1204,1226]" captionTargetBox="[140,1108,165,1184]" captionTargetId="figure@151.[140,1108,164,1184]" captionTargetPageId="151" captionText="Figure 79. SEM, epigynum. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E, F, Crassignatha quanqu sp. n. from Bang Bie village. A, C, E, ventral; B, D, F, lateral, anterior to the right. BL: basal lobe of scape; DL: distal lobe of scape." figureDoi="http://doi.org/10.5281/zenodo.3768898" httpUri="https://zenodo.org/record/3768898/files/figure.png" pageId="64" pageNumber="73">79E</figureCitation>
). Spermathecae separated by half their diameter. Ducts arise from posteromesal part of spermathecae, follow complex path before opening near scape apex (
<figureCitation box="[273,380,519,546]" captionStart="Figure 76" captionStartId="148.[140,212,962,984]" captionTargetBox="[140,1108,165,937]" captionTargetId="figure@148.[140,1108,164,937]" captionTargetPageId="148" captionText="Figure 76. A, B, Crassignatha pianma sp. n. from Chang Yan He; C, D, Crassignatha yinzhi from Xiao Hei Shan Nature Reserve; E-G, Crassignatha quanqu sp. n. from Bang Bie village; H, I, Crassignatha yamu sp. n. from Yamu He. A, F, epigynum, lateral view; B, D, G, I, cleared vulva, dorsal view; C, E, H, epigynum, ventral view. CD: copulatory duct; S: spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3768892" httpUri="https://zenodo.org/record/3768892/files/figure.png" pageId="64" pageNumber="73">Fig. 76G</figureCitation>
).
</paragraph>
<paragraph blockId="64.[140,1108,166,933]" pageId="64" pageNumber="73">
Male (CASENT 9022387): Total length 0.73, carapace 0.35 long, 0.34 wide, clypeus 0.11, sternum 0.25 long, 0.24 wide, coxa IV separated by 2.00 times their width. Leg I: patella d1, tibia d2, p1, metatarsus unmodified; Leg II: patella d1, tibia d2, v2, metatarsus excavated proximoventrally; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.42; TmII: 0.42; TmIII: absent. Leg measurements: see Appendix A. Spinnerets (
<figureCitation box="[687,784,730,757]" captionStart="Figure 85" captionStartId="157.[140,212,1202,1224]" captionTargetBox="[140,1108,165,1182]" captionTargetId="figure@157.[140,1108,164,1182]" captionTargetPageId="157" captionText="Figure 85. Crassignatha quanqu sp. n. from Bang Bie village, SEM of spinnerets. A-E, female; F, male. A, spinnerets; B, ALS; C, PMS; D, F, PLS; E, base of ALS showing lack of colulus. AC: aciniform gland spigot; AG: aggregate gland spigot; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL:flagelliform gland spigot; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi="http://doi.org/10.5281/zenodo.3768906" httpUri="https://zenodo.org/record/3768906/files/figure.png" pageId="64" pageNumber="73">Fig. 85F</figureCitation>
)
</paragraph>
<paragraph blockId="64.[140,1108,166,933]" pageId="64" pageNumber="73">
Female (CASENT 9022387): Total length 1.00, carapace 0.41 long, 0.38 wide, clypeus 0.09, sternum 0.25 long, 0.24 wide, coxa IV separated by 1.63 times their width. Leg I: patella d1, tibia d2, p1; Leg II: patella d1, tibia d2; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.41; TmII: 0.41; TmIII: absent. Leg measurements: see Appendix A. Spinnerets (
<figureCitation box="[857,936,906,933]" captionStart="Figure 85" captionStartId="157.[140,212,1202,1224]" captionTargetBox="[140,1108,165,1182]" captionTargetId="figure@157.[140,1108,164,1182]" captionTargetPageId="157" captionText="Figure 85. Crassignatha quanqu sp. n. from Bang Bie village, SEM of spinnerets. A-E, female; F, male. A, spinnerets; B, ALS; C, PMS; D, F, PLS; E, base of ALS showing lack of colulus. AC: aciniform gland spigot; AG: aggregate gland spigot; ALS: anterior lateral spinneret; CY: cylindrical gland spigot; FL:flagelliform gland spigot; MAP: major ampullate gland spigot; mAP: minor ampullate gland spigot; n: nubbin; PI: piriform gland spigot; PLS: posterior lateral spinneret; PMS: posterior median spinneret; t: tartipore." figureDoi="http://doi.org/10.5281/zenodo.3768906" httpUri="https://zenodo.org/record/3768906/files/figure.png" pageId="64" pageNumber="73">Fig. 85</figureCitation>
A-D)
</paragraph>
</subSubSection>
</treatment>
</document>