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<document id="A2513D19444F97AFBBD7A4B19E43AFE0" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-Zenodo-Dep="13742850" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFD0892F9D34FC85FB77FB1B" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis punctatostriatus Janssens 1937" docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="39" updateTime="1726239157283" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="642F3D1F185A98906A2ADB3D8259E5C3">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
</mods:titleInfo>
<mods:name id="E32670EF3C109DC0ECFA4CECD910A7CD" type="personal">
<mods:role id="31BEC00E6C38C6633ED5D50CFCAAED1C">
<mods:roleTerm id="BDCC8FB63B925DF2AEF850D121A80718">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CC20DBCB422E666104CB1FE70144C23">Karimbumkara, Seena Narayanan</mods:namePart>
<mods:nameIdentifier id="0DF2B315E502F4497807B1690B57CFFF" type="LSID">A6B50103-8D6D-4977-9111-EF8BA4AA0477</mods:nameIdentifier>
<mods:affiliation id="0BA9B25CFDA813CCBCCA70295DA63AE7">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="84DADA43BB446DFFA4A457A74F695172" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
</mods:name>
<mods:name id="FFA45CB413D5BD7852A1211D700C9057" type="personal">
<mods:role id="E5FD13FFF2D3B1AAD739D3EB3EAB2EAB">
<mods:roleTerm id="DA672FC6506B48CFECFDA590F8BD723C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="18A1E166DCFC190F43C07D8E5E67C04B">Priyadarsanan, Dharma Rajan</mods:namePart>
<mods:nameIdentifier id="88561B8A7AC33333ED6664E00AE2A69E" type="LSID">03907E93-F783-4720-9F7B-CB662E7FC745</mods:nameIdentifier>
<mods:affiliation id="D716DF390F20DAC0FE034954650C561F">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="CE00244A9D9B1FABDDFCF9E2CF770647" type="email">priyan@atree.org</mods:nameIdentifier>
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<mods:typeOfResource id="8F770D62384FD14CE8AF64E4C9E5565E">text</mods:typeOfResource>
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<mods:title id="C639F0EB34D25B933DE6E2BAFA3A065E">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
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<mods:date id="F409B3CA0E8CD84239092F0241D58A5A">2024</mods:date>
<mods:detail id="9AB74F39E400D84AD3FD86E0EFFB2EE5" type="pubDate">
<mods:number id="78FCE92B09EBCA5D4999F897D1731915">2024-09-10</mods:number>
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<mods:detail id="5F3A3B5064B4AC911066A7EB875F89C6" type="volume">
<mods:number id="46CDBB9C05DABCD9B8BC060BD24E86BA">956</mods:number>
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<mods:classification id="DD8EF18D2D239AC929BCC70152250123">journal article</mods:classification>
<mods:identifier id="E43235FC1A5540197B400C4774D8D32C" type="DOI">10.5852/ejt.2024.956.2657</mods:identifier>
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<treatment id="557187D2FFD0892F9D34FC85FB77FB1B" ID-DOI="http://doi.org/10.5281/zenodo.13742917" ID-Zenodo-Dep="13742917" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B" httpUri="http://treatment.plazi.org/id/557187D2FFD0892F9D34FC85FB77FB1B" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39">
<subSubSection id="95C2654FFFD089209D34FC85FC06FCC4" box="[565,1023,820,846]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="DD6736C4FFD089209D34FC85FC06FCC4" blockId="38.[560,1027,820,882]" box="[565,1023,820,846]" pageId="38" pageNumber="39">
<heading id="862F81A8FFD089209D34FC85FC06FCC4" box="[565,1023,820,846]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="39" reason="2">
<taxonomicName id="1AD84D47FFD089209D34FC85FC06FCC4" ID-CoL="49KKQ" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[565,1023,820,846]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD089209D34FC85FCB0FCC4" bold="true" box="[565,841,820,846]" italics="true" pageId="38" pageNumber="39">Onitis punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFD089209C51FC85FC06FCC4" author="Janssens A." box="[848,1023,820,846]" pageId="38" pageNumber="39" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD089209D31FCE6FBFAFCFB" box="[560,1027,855,882]" pageId="38" pageNumber="39" type="description">
<paragraph id="DD6736C4FFD089209D31FCE6FBFAFCFB" blockId="38.[560,1027,820,882]" box="[560,1027,855,882]" pageId="38" pageNumber="39">
<figureCitation id="45E32A41FFD089209D31FCE6FD48FCFB" box="[560,689,855,881]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="38" pageNumber="39">Figs 6M–P</figureCitation>
,
<figureCitation id="45E32A41FFD089209DBCFCE6FD18FCFB" box="[701,737,855,881]" captionStart="Fig" captionStartId="31.[189,232,1858,1884]" captionTargetBox="[303,1358,297,1749]" captionTargetId="figure-1@31.[189,1398,262,1791]" captionTargetPageId="31" captionText="Fig. 9. Head of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/NBAIR/COL/ONIT/H- 271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742875" httpUri="https://zenodo.org/record/13742875/files/figure.png" pageId="38" pageNumber="39">9B</figureCitation>
,
<figureCitation id="45E32A41FFD089209DEFFCE6FCE5FCFB" box="[750,796,855,881]" captionStart="Fig" captionStartId="32.[190,233,1814,1840]" captionTargetBox="[325,1212,290,1775]" captionTargetId="figure-1@32.[189,1398,262,1779]" captionTargetPageId="32" captionText="Fig. 10. A. Onitisfeae Felsche, 1907, cotype, (image: Olaf Jäger, SNSD). B. O. humerosus (Pallas, 1771), BMNH (E) 1237143, (image: BMNH). C. O. lama Lansberge, 1875, syntype, ♂, MNHN EC4810, (image: Antoine Mantilleri, MNHN). D. O. naviauxi Cambefort, 1988, holotype, ♂, MNHN EC4807, (image: Antoine Mantilleri, MNHN). E. O. philemon Fabricius, 1801, lectotype, ZMUK Fabricius 003706, (image: Alexey Solodovnikov, ZMUC). F. O. punctatostriatus Janssens, 1937, holotype, ♂, (image: Jonathan Brecko, IRSNB)." figureDoi="http://doi.org/10.5281/zenodo.13742879" httpUri="https://zenodo.org/record/13742879/files/figure.png" pageId="38" pageNumber="39">10F</figureCitation>
,
<figureCitation id="45E32A41FFD089209C29FCE6FCA2FCFB" box="[808,859,855,881]" captionStart="Fig" captionStartId="37.[189,232,1857,1883]" captionTargetBox="[297,1300,330,1792]" captionTargetId="figure-1@37.[189,1398,285,1802]" captionTargetPageId="37" captionText="Fig. 14. Pronotum of male specimens. A. Onitis bhomorensis sp. nov., holotype, NIM/NBAIR/ COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/ Sc1000317. E. O.visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742891" httpUri="https://zenodo.org/record/13742891/files/figure.png" pageId="38" pageNumber="39">14B</figureCitation>
,
<figureCitation id="45E32A41FFD089209C66FCE6FC63FCF8" box="[871,922,855,882]" captionStart="Fig" captionStartId="39.[189,233,1858,1884]" captionTargetBox="[302,1276,327,1774]" captionTargetId="figure-1@39.[189,1398,264,1787]" captionTargetPageId="39" captionText="Fig. 15. Pronotal basal foveae of male specimens. A. Onitisbhomorensis sp. nov., holotype, NIM/ NBAIR/COL/ONIT/H-271023A. B. O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. O. kethai sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023B. D. O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. O. visthara sp. nov., holotype, NIM/NBAIR/COL/ONIT/H-271023C. F. O. singhalensis Lansberge, 1875, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="38" pageNumber="39">15B</figureCitation>
,
<figureCitation id="45E32A41FFD089209CA6FCE6FC23FCFB" box="[935,986,855,881]" captionStart="Fig 16" captionStartId="40.[189,229,1858,1884]" captionTargetBox="[303,1269,309,1806]" captionTargetId="figure-1@40.[189,1398,262,1810]" captionTargetPageId="40" captionText="Fig 16. A. Protibia of Onitis bhomorensis sp. nov. showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A. B. Anterior tip of protibia of O. punctatostriatus Janssens, 1937, AIMB/Co/Sc1000492. C. Mid-leg of O. kethai sp. nov, NIM/NBAIR/COL/ONIT/H-271023B. D. Mid-leg of O. philemon Fabricius, 1801, AIMB/Co/Sc1000317. E. Mid-leg of O. visthara sp. nov, NIM/NBAIR/COL/ONIT/H- 271023C. F. Mid-leg of O. singhalensis Lansberge, AIMB/Co/Sc1000684. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="38" pageNumber="39">16B</figureCitation>
,
<figureCitation id="45E32A41FFD089209CE7FCE6FBFAFCFB" box="[998,1027,855,881]" captionStart="Fig" captionStartId="49.[189,232,1927,1953]" captionTargetBox="[227,1362,257,1876]" captionTargetId="figure-1@49.[212,1375,244,1888]" captionTargetPageId="49" captionText="Fig. 19. Map showing the distribution of Onitislama Lansberge, 1875; O. naviauxi Cambefort, 1988; O. philemon Fabricius, 1801; O. punctatostriatus Janssens, 1937 and O. singhalensis Lansberge, 1875 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="38" pageNumber="39">19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD089209FBCFC28FC6FFBB0" pageId="38" pageNumber="39" type="reference_group">
<paragraph id="DD6736C4FFD089209FBCFC28FC57FC38" blockId="38.[189,942,920,947]" box="[189,942,920,947]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FBCFC28FE32FC39" box="[189,459,921,947]" italics="true" pageId="38" pageNumber="39">Onitispunctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFD089209ED3FC28FD5FFC38" author="Janssens A." box="[466,678,920,947]" pageId="38" pageNumber="39" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 19</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFD089209FBCFC6BFC6FFBB0" blockId="38.[189,1399,986,1083]" pageId="38" pageNumber="39">
<treatmentCitationGroup id="FDC811EAFFD089209FBCFC6BFC6FFBB0" pageId="38" pageNumber="39">
<taxonomicName id="1AD84D47FFD089209FBCFC6BFE32FC7E" ID-CoL="49KKQ" authorityName="Janssens" authorityYear="1937" box="[189,459,986,1012]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD089209FBCFC6BFE32FC7E" box="[189,459,986,1012]" italics="true" pageId="38" pageNumber="39">Onitis punctatostriatus</emphasis>
</taxonomicName>
–
<treatmentCitation id="5C7910D5FFD089209EE6FC6BFD3DFC7F" author="Balthasar V." box="[487,708,986,1013]" page="35" pageId="38" pageNumber="39" year="1963">
<bibRefCitation id="B9494B35FFD089209EE6FC6BFD3DFC7F" author="Balthasar V." box="[487,708,986,1013]" pageId="38" pageNumber="39" refId="ref27200" refString="Balthasar V. 1963. Monographie der Scarabaeidae undAphodiidae der palaeoarktischen und orientalischen Region (Coleoptera: Lamellicornia). Band 2: Coprinae (Onitini, Oniticellini, Onthophagini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prague." type="book" year="1963">Balthasar 1963: 35</bibRefCitation>
</treatmentCitation>
(monograph). —
<treatmentCitation id="5C7910D5FFD089209C96FC6BFB34FC7F" author="Bezdek A. &amp; Krell F. - T." box="[919,1229,986,1013]" page="159" pageId="38" pageNumber="39" year="2006">
<bibRefCitation id="B9494B35FFD089209C96FC6BFB34FC7F" author="Bezdek A. &amp; Krell F. - T." box="[919,1229,986,1013]" pageId="38" pageNumber="39" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). — Siddiqui
<emphasis id="EFACEAD6FFD089209E7AFC4FFE47FB9D" box="[379,446,1021,1047]" italics="true" pageId="38" pageNumber="39">et. al.</emphasis>
2014: 300 (distribution). —
<treatmentCitation id="5C7910D5FFD089209C15FC4CFBD5FB9D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[788,1068,1021,1048]" page="1037" pageId="38" pageNumber="39" year="2015">
<bibRefCitation id="B9494B35FFD089209C15FC4CFBD5FB9D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[788,1068,1021,1048]" pageId="38" pageNumber="39" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFD089209C62FC4FFC66FB9D" box="[867,927,1021,1047]" italics="true" pageId="38" pageNumber="39">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFD089209BA1FC4CFA8FFB9D" author="Bezdek A." box="[1184,1398,1021,1047]" page="179" pageId="38" pageNumber="39" year="2016">
<bibRefCitation id="B9494B35FFD089209BA1FC4CFA8FFB9D" author="Bezdek A." box="[1184,1398,1021,1047]" pageId="38" pageNumber="39" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFD089209EB8FB91FD4AFBB0" author="Schoolmeesters P." box="[441,691,1056,1082]" pageId="38" pageNumber="39" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD089209FBCFBD0FC88FA9E" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="DD6736C4FFD089209FBCFBD0FE3FFBF1" blockId="38.[189,454,1121,1147]" box="[189,454,1121,1147]" pageId="38" pageNumber="39">
<heading id="862F81A8FFD089209FBCFBD0FE3FFBF1" bold="true" box="[189,454,1121,1147]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="EFACEAD6FFD089209FBCFBD0FE3FFBF1" bold="true" box="[189,454,1121,1147]" pageId="38" pageNumber="39">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD089209FBCFB20FC88FA9E" blockId="38.[189,1399,1168,1300]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FBCFB20FE35FB21" box="[189,460,1169,1195]" italics="true" pageId="38" pageNumber="39">Onitispunctatostriatus</emphasis>
is similar to
<taxonomicName id="1AD84D47FFD089209D6AFB20FCF5FB20" authorityName="Arrow" authorityYear="1931" box="[619,780,1168,1195]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFD089209D6AFB20FCF5FB20" box="[619,780,1168,1195]" italics="true" pageId="38" pageNumber="39">O. subopacus</emphasis>
</taxonomicName>
in having frontal carina broadly interrupted and the metasternum smooth and shining behind with scattered punctures. It differs from the latter in having deep and broad elytral striae and the pronotum being strongly ocellate punctured. This species is reported for the first time from the state of
<collectingRegion id="1F1CF826FFD089209D1EFB4BFD96FA9E" box="[543,623,1274,1300]" country="India" name="Assam" pageId="38" pageNumber="39">Assam</collectingRegion>
in northeastern
<collectingCountry id="A5CF7654FFD089209C2CFB4BFC94FA9E" box="[813,877,1274,1300]" name="India" pageId="38" pageNumber="39">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD0892F9FBCFA8AFDF7FECD" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="DD6736C4FFD089209FBCFA8AFE15FADF" blockId="38.[189,492,1339,1365]" box="[189,492,1339,1365]" pageId="38" pageNumber="39">
<heading id="862F81A8FFD089209FBCFA8AFE15FADF" bold="true" box="[189,492,1339,1365]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="EFACEAD6FFD089209FBCFA8AFE15FADF" bold="true" box="[189,492,1339,1365]" pageId="38" pageNumber="39">Type material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFD089209FEBFADBFEE2FA67" collectionCode="IRSNB" collectorName="R. P. Verschraeghen &amp; A. Janssens" country="India" determinerName="J. J. Gillet" location="INDIA" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="DD6736C4FFD089209FEBFADBFEA5FA0E" blockId="38.[189,1399,1386,1517]" box="[234,348,1386,1412]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FEBFADBFEA5FA0E" bold="true" box="[234,348,1386,1412]" pageId="38" pageNumber="39">
<typeStatus id="02638866FFD089209FEBFADBFEA5FA0E" box="[234,348,1386,1412]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD089209FBCFA3CFEE2FA67" blockId="38.[189,1399,1386,1517]" pageId="38" pageNumber="39">
<collectingCountry id="A5CF7654FFD089209FBCFA3CFEF7FA2D" box="[189,270,1421,1447]" name="India" pageId="38" pageNumber="39">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFD089209E2AFA3CFEBDFA2D" box="[299,324,1421,1447]" count="1" pageId="38" pageNumber="39" type="male">♂</specimenCount>
; “Darjeeling 1904, R.P. Verschraeghen, cf. Mem. Mus. Hist. Nat. Belg., 2: Serie, fasc. 11, 1937, p. 48, Ex. Typis, A. Janssens rev., 1936:
<taxonomicName id="1AD84D47FFD089209DF3FA00FB25FA40" authority="Janss., J. J. Gillet" authorityName="Janss., J. J. Gillet" box="[754,1244,1456,1483]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD089209DF3FA00FBFBFA40" box="[754,1026,1457,1483]" italics="true" pageId="38" pageNumber="39">Onitis punctatostriatus</emphasis>
Janss.,
<determinerName id="28BBDA00FFD089209B60FA00FB25FA40" box="[1121,1244,1456,1483]" pageId="38" pageNumber="39">J. J. Gillet</determinerName>
</taxonomicName>
det., vend.”;
<collectionCode id="BBC9AE01FFD089209FBCFA62FEEEFA67" box="[189,279,1491,1517]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="39">IRSNB</collectionCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="6DB03C99FFD089209FEBF9A4FD3BF9FF" collectionCode="IRSNB" collectorName="Onitisspinipes Drury R. M. H. N. Belg." country="India" determinerName="J. Gillet" location="INDIA" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<paragraph id="DD6736C4FFD089209FEBF9A4FEA5F9A5" blockId="38.[189,1398,1557,1654]" box="[234,348,1557,1583]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FEBF9A4FEA5F9A5" bold="true" box="[234,348,1557,1583]" pageId="38" pageNumber="39">
<typeStatus id="02638866FFD089209FEBF9A4FEA5F9A5" box="[234,348,1557,1583]" pageId="38" pageNumber="39" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD089209FBCF989FD3FF9FF" blockId="38.[189,1398,1557,1654]" pageId="38" pageNumber="39">
<collectingCountry id="A5CF7654FFD089209FBCF989FEF7F9D8" box="[189,270,1592,1618]" name="India" pageId="38" pageNumber="39">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFD089209E2DF988FEBAF9D9" box="[300,323,1593,1619]" count="1" pageId="38" pageNumber="39" type="female">♀</specimenCount>
; “
<emphasis id="EFACEAD6FFD089209E60F989FDEAF9D8" box="[353,531,1592,1618]" italics="true" pageId="38" pageNumber="39">Onitisspinipes</emphasis>
Drury R.M.H.N. Belg. 10. 640,
<emphasis id="EFACEAD6FFD089209CA1F988FBF9F9D8" box="[928,1024,1593,1618]" italics="true" pageId="38" pageNumber="39">spinipes</emphasis>
Drury;
<determinerName id="28BBDA00FFD089209B63F988FB3BF9D8" box="[1122,1218,1592,1619]" pageId="38" pageNumber="39">J. Gillet</determinerName>
det. 1923, Ex-Typis”; label data same as for male;
<collectionCode id="BBC9AE01FFD089209D69F9EAFD3BF9FF" box="[616,706,1627,1653]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="38" pageNumber="39">IRSNB</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="DD6736C4FFD089209FEBF92DFD07F93D" blockId="38.[189,1398,1692,2001]" box="[234,766,1692,1719]" pageId="38" pageNumber="39">
<emphasis id="EFACEAD6FFD089209FEBF92DFDBDF93C" bold="true" box="[234,580,1692,1718]" pageId="38" pageNumber="39">Additional material studied</emphasis>
(
<specimenCount id="CBDEFD4DFFD089209D55F92CFD0FF93D" box="[596,758,1693,1719]" count="10" pageId="38" pageNumber="39" type="generic">10 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFD0892F9FBCF971FDF7FECD" blockId="38.[189,1398,1692,2001]" lastBlockId="41.[189,1398,263,327]" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39">
<materialsCitation id="6DB03C99FFD089209FBCF971FD93F974" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Rajkamal Goswami" country="India" location="Majuli Island" pageId="38" pageNumber="39" specimenCode="Co/Sc1000492 to 1000495" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Assam">
<collectingCountry id="A5CF7654FFD089209FBCF971FEE8F950" box="[189,273,1728,1754]" name="India" pageId="38" pageNumber="39">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFD089209E33F971FE85F950" box="[306,380,1728,1754]" count="2" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD089209E8CF971FE2CF950" box="[397,469,1728,1754]" count="2" pageId="38" pageNumber="39" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFD089209EE6F971FDC4F950" box="[487,573,1728,1754]" country="India" name="Assam" pageId="38" pageNumber="39">Assam</collectingRegion>
,
<location id="D807601FFFD089209D4DF971FD01F950" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209D4DF971FD01F950" box="[588,760,1728,1754]" country="India" name="Majuli Island" pageId="38" pageNumber="39" stateProvince="Assam">Majuli Island</location>
;
<collectingMethod id="04994ED3FFD089209C09F971FC5BF950" box="[776,930,1728,1754]" pageId="38" pageNumber="39">hand-picked</collectingMethod>
, cow dung;
<collectorName id="702D5312FFD089209B40F971FACCF950" box="[1089,1333,1728,1754]" pageId="38" pageNumber="39">Rajkamal Goswami</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD089209FBCF952FEF5F977" box="[189,268,1763,1789]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD089209E17F952FD93F974" box="[278,618,1763,1790]" pageId="38" pageNumber="39">Co/Sc1000492 to 1000495</specimenCode>
</materialsCitation>
•
<materialsCitation id="6DB03C99FFD089209D95F955FD97F8C9" collectingDate="2018-09-30" collectingMethod="cow dung baited pitfall trap" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="127" latitude="27.96913" location="Rani Village" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" pageId="38" pageNumber="39" specimenCode="Co/Sc1000496" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFD089209D95F955FD54F974" box="[660,685,1764,1790]" count="1" pageId="38" pageNumber="39" type="male">♂</specimenCount>
;
<collectingRegion id="1F1CF826FFD089209DC0F952FC4AF977" box="[705,947,1763,1789]" country="India" name="Arunachal Pradesh" pageId="38" pageNumber="39">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD089209CC6F952FBCBF977" box="[967,1074,1763,1789]" pageId="38" pageNumber="39">Pasighat</collectingMunicipality>
, community forest,
<location id="D807601FFFD089209A3CF952FEE0F8AA" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209A3CF952FEE0F8AA" country="India" latitude="27.96913" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" name="Rani Village" pageId="38" pageNumber="39" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD089209E2BF8B4FE05F8AB" box="[298,508,1797,1825]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="8.8644" value="27.96913">27°58ʹ8.8644ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD089209D0CF8B4FD14F8AB" box="[525,749,1797,1825]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="17.1804" value="95.32144">95°19ʹ17.1804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD089209C43F8B7FC20F8AA" box="[834,985,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">
<quantity id="1A209B21FFD089209C43F8B7FC6BF8AB" box="[834,914,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">127 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD089209CEAF8B7FB6CF8AA" box="[1003,1173,1798,1824]" pageId="38" pageNumber="39" value="2018-09-30">
<collectingDate id="B922E9ECFFD089209CEAF8B7FB6CF8AA" box="[1003,1173,1798,1824]" pageId="38" pageNumber="39" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD089209BA7F8B6FEB5F8CE" pageId="38" pageNumber="39">cow dung baited pitfall trap</collectingMethod>
;
<collectionCode id="BBC9AE01FFD089209E63F898FE48F8C9" box="[354,433,1833,1859]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD089209EBAF898FD97F8C9" box="[443,622,1833,1859]" pageId="38" pageNumber="39">Co/Sc1000496</specimenCode>
</materialsCitation>
•
<materialsCitation id="6DB03C99FFD089209D9CF89BFEBCF800" collectingDate="2018-09-30" collectingMethod="baited pitfall trap" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="131" latitude="27.969614" location="Rani Village" longLatPrecision="1" longitude="95.32392" pageId="38" pageNumber="39" specimenCode="Co/ Sc1000497" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFD089209D9CF89BFD4DF8CE" box="[669,692,1834,1860]" count="1" pageId="38" pageNumber="39" type="female">♀</specimenCount>
; community forest,
<location id="D807601FFFD089209CCAF898FB88F8C9" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209CCAF898FB88F8C9" box="[971,1137,1833,1859]" country="India" latitude="27.969614" longLatPrecision="1" longitude="95.32392" name="Rani Village" pageId="38" pageNumber="39">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD089209B88F899FA88F8CE" box="[1161,1393,1832,1860]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="10.6068" value="27.969614">27°58ʹ10.6068ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD089209FBCF8FAFE67F8ED" box="[189,414,1867,1895]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="26.1156" value="95.32392">95°19ʹ26.1156ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD089209EF9F8FCFD6DF8ED" box="[504,660,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="38" pageNumber="39" unit="m" value="131.0">
<quantity id="1A209B21FFD089209EF9F8FCFDB3F8ED" box="[504,586,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="38" pageNumber="39" unit="m" value="131.0">131 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD089209DA9F8FCFCA1F8ED" box="[680,856,1869,1895]" pageId="38" pageNumber="39" value="2018-09-30">
<collectingDate id="B922E9ECFFD089209DA9F8FCFCA1F8ED" box="[680,856,1869,1895]" pageId="38" pageNumber="39" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
; cow dung;
<collectingMethod id="04994ED3FFD089209CFFF8FCFB1BF8ED" box="[1022,1250,1869,1895]" pageId="38" pageNumber="39">baited pitfall trap</collectingMethod>
;
<collectionCode id="BBC9AE01FFD089209BF2F8FCFAB8F8ED" box="[1267,1345,1869,1895]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD089209A4AF8FCFEBCF800" pageId="38" pageNumber="39">Co/ Sc1000497</specimenCode>
</materialsCitation>
•
<materialsCitation id="6DB03C99FFD089209E63F8C1FE2EF85B" collectingDate="2018-09-29" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="127" latitude="27.96913" location="Rani Village" longLatPrecision="1" longitude="95.32144" pageId="38" pageNumber="39" specimenCode="Co/ Sc1000498 to 1000500" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFD089209E63F8C1FE83F800" box="[354,378,1904,1930]" count="1" pageId="38" pageNumber="39" type="male">♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD089209E88F8C1FE36F800" box="[393,463,1904,1930]" count="2" pageId="38" pageNumber="39" type="female">2 ♀♀</specimenCount>
; community forest,
<location id="D807601FFFD089209DC8F8C1FC9EF800" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209DC8F8C1FC9EF800" box="[713,871,1904,1930]" country="India" latitude="27.96913" longLatPrecision="1" longitude="95.32144" name="Rani Village" pageId="38" pageNumber="39">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD089209C75F8DFFBBCF800" box="[884,1093,1902,1930]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="8.8644" value="27.96913">27°58ʹ8.8644ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD089209B52F8DFFAC9F800" box="[1107,1328,1902,1930]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="38" pageNumber="39" precision="1" seconds="17.1804" value="95.32144">95°19ʹ17.1804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD089209FBCF822FEB2F827" box="[189,331,1939,1966]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">
<quantity id="1A209B21FFD089209FBCF822FEF0F824" box="[189,265,1939,1966]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="38" pageNumber="39" unit="m" value="127.0">127 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD089209E5BF822FE05F827" box="[346,508,1939,1965]" pageId="38" pageNumber="39" value="2018-09-29">
<collectingDate id="B922E9ECFFD089209E5BF822FE05F827" box="[346,508,1939,1965]" pageId="38" pageNumber="39" value="2018-09-29">29 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD089209D09F822FD66F827" box="[520,671,1939,1965]" pageId="38" pageNumber="39">hand-picked</collectingMethod>
from cow dung,
<collectorName id="702D5312FFD089209C6FF822FBC2F827" box="[878,1083,1939,1965]" pageId="38" pageNumber="39">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD089209BF2F822FAB8F827" box="[1267,1345,1939,1965]" pageId="38" pageNumber="39">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD089209A4AF822FE2EF85B" pageId="38" pageNumber="39">Co/ Sc1000498 to 1000500</specimenCode>
</materialsCitation>
•
<materialsCitation id="6DB03C99FFD0892F9EF0F806FDF2FECD" collectingDate="2019-02-18" collectingMethod="cow dung baited pitfall trap" collectionCode="AIMB" collectorName="Forest &amp; Harsha Malhotra" country="India" lastPageId="41" lastPageNumber="42" latitude="23.277756" location="South" longLatPrecision="1" longitude="91.40453" municipality="Trishna Wildlife Sanctuary" pageId="38" pageNumber="39" specimenCode="Co/Sc1000501" specimenCount="1" specimenCount-female="1" stateProvince="Tripura">
<specimenCount id="CBDEFD4DFFD089209EF0F806FDF1F85B" box="[497,520,1975,2001]" count="1" pageId="38" pageNumber="39" type="female">♀</specimenCount>
;
<collectingRegion id="1F1CF826FFD089209D19F807FD8FF85A" box="[536,630,1974,2000]" country="India" name="Tripura" pageId="38" pageNumber="39">Tripura</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD089209D80F807FC37F85A" box="[641,974,1974,2000]" pageId="38" pageNumber="39">Trishna Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFD089209CDBF807FBD8F85A" LSID="urn:lsid:plazi:treatment:557187D2FFD0892F9D34FC85FB77FB1B:D807601FFFD089209CDBF807FBD8F85A" box="[986,1057,1974,2000]" country="India" latitude="23.277756" longLatPrecision="1" longitude="91.40453" municipality="Trishna Wildlife Sanctuary" name="South" pageId="38" pageNumber="39" stateProvince="Tripura">South</location>
<collectingRegion id="1F1CF826FFD089209B29F807FB7EF85A" box="[1064,1159,1974,2000]" country="India" name="Tripura" pageId="38" pageNumber="39">Tripura</collectingRegion>
;
<geoCoordinate id="B8EC5003FFD089209B92F804FA88F85B" box="[1171,1393,1973,2001]" degrees="23" direction="north" minutes="16" orientation="latitude" pageId="38" pageNumber="39" precision="1" seconds="39.9216" value="23.277756">23°16ʹ39.9216ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFDF892F9FBCFEB6FE65FEA9" box="[189,412,263,291]" degrees="91" direction="east" minutes="24" orientation="longitude" pageId="41" pageNumber="42" precision="1" seconds="16.3152" value="91.40453">91°24ʹ16.3152ʺ E</geoCoordinate>
;
<collectorName id="702D5312FFDF892F9EABFEB8FE03FEA9" box="[426,506,265,291]" pageId="41" pageNumber="42">Forest</collectorName>
,
<date id="A9661004FFDF892F9D08FEB8FD48FEA9" box="[521,689,265,291]" pageId="41" pageNumber="42" value="2019-02-18">
<collectingDate id="B922E9ECFFDF892F9D08FEB8FD48FEA9" box="[521,689,265,291]" pageId="41" pageNumber="42" value="2019-02-18">18 Feb. 2019</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFDF892F9DBEFEB8FBE4FEA9" box="[703,1053,265,291]" pageId="41" pageNumber="42">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFDF892F9B2DFEB8FB05FEA9" box="[1068,1276,265,291]" pageId="41" pageNumber="42">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFDF892F9FFBFE9DFEB2FECD" box="[250,331,300,327]" pageId="41" pageNumber="42">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFDF892F9E54FE9DFDF2FECD" box="[341,523,300,327]" pageId="41" pageNumber="42">Co/Sc1000501</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="89A7664CFFD189219FBCF8F3FB9DF863" ID-DOI="http://doi.org/10.5281/zenodo.13742895" ID-Zenodo-Dep="13742895" httpUri="https://zenodo.org/record/13742895/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,233,1858,1884]" targetBox="[302,1276,327,1774]" targetPageId="39">
<paragraph id="DD6736C4FFD189219FBCF8F3FB9DF863" blockId="39.[189,1406,1858,2026]" pageId="39" pageNumber="40">
<emphasis id="EFACEAD6FFD189219FBCF8F3FEE7F8D6" bold="true" box="[189,286,1858,1884]" pageId="39" pageNumber="40">Fig. 15.</emphasis>
Pronotal basal foveae of male specimens.
<emphasis id="EFACEAD6FFD189219C2CF8F3FCBBF8D6" bold="true" box="[813,834,1858,1884]" pageId="39" pageNumber="40">A</emphasis>
.
<emphasis id="EFACEAD6FFD189219C52F8F3FBB8F8D6" box="[851,1089,1858,1884]" italics="true" pageId="39" pageNumber="40">Onitisbhomorensis</emphasis>
<taxonomicNameLabel id="F49F57ADFFD189219B4BF8F2FB50F8D7" box="[1098,1193,1859,1885]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NIM/ NBAIR/COL/ONIT/H-271023A.
<emphasis id="EFACEAD6FFD189219D59F8D4FD95F8F5" bold="true" box="[600,620,1893,1919]" pageId="39" pageNumber="40">B</emphasis>
.
<taxonomicName id="1AD84D47FFD189219D86F8D7FBAFF8F5" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[647,1110,1893,1920]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFD189219D86F8D7FC87F8F5" box="[647,894,1894,1920]" italics="true" pageId="39" pageNumber="40">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFD189219C93F8D7FBAFF8F5" author="Janssens A." box="[914,1110,1893,1920]" pageId="39" pageNumber="40" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000492.
<emphasis id="EFACEAD6FFD189219FBCF839FF2BF828" bold="true" box="[189,210,1928,1954]" pageId="39" pageNumber="40">C</emphasis>
.
<taxonomicName id="1AD84D47FFD189219FE7F838FEA3F828" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[230,346,1928,1955]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFD189219FE7F838FEA3F828" box="[230,346,1928,1955]" italics="true" pageId="39" pageNumber="40">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFD189219E67F838FE31F829" box="[358,456,1929,1955]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NIM/NBAIR/COL/ONIT/H-271023B.
<emphasis id="EFACEAD6FFD189219B2CF838FBBBF829" bold="true" box="[1069,1090,1929,1955]" pageId="39" pageNumber="40">D</emphasis>
.
<taxonomicName id="1AD84D47FFD189219B57F838FF04F84C" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFD189219B57F838FB09F828" box="[1110,1264,1928,1955]" italics="true" pageId="39" pageNumber="40">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFD189219BFCF839FF04F84C" author="Fabricius J. C." pageId="39" pageNumber="40" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000317.
<emphasis id="EFACEAD6FFD189219D19F81DFDD5F84C" bold="true" box="[536,556,1964,1990]" pageId="39" pageNumber="40">E</emphasis>
.
<taxonomicName id="1AD84D47FFD189219D3BF81DFD3BF84C" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[570,706,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFD189219D3BF81DFD3BF84C" box="[570,706,1964,1990]" italics="true" pageId="39" pageNumber="40">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFD189219DCBF81DFCDEF84C" box="[714,807,1964,1990]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, NIM/NBAIR/COL/ONIT/H-271023C.
<emphasis id="EFACEAD6FFD189219FBCF87EFF2FF863" bold="true" box="[189,214,1999,2025]" pageId="39" pageNumber="40">F.</emphasis>
<taxonomicName id="1AD84D47FFD189219FDEF87EFD8DF860" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" box="[223,628,1999,2026]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFD189219FDEF87EFE64F863" box="[223,413,1999,2025]" italics="true" pageId="39" pageNumber="40">O. singhalensis</emphasis>
<bibRefCitation id="B9494B35FFD189219EA4F87EFD8DF860" author="Lansberge G." box="[421,628,1999,2026]" pageId="39" pageNumber="40" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000684. (Images: AIMB).
</paragraph>
</caption>
<caption id="89A7664CFFDE892E9FBCF8F3FB1DF860" ID-DOI="http://doi.org/10.5281/zenodo.13742897" ID-Zenodo-Dep="13742897" httpUri="https://zenodo.org/record/13742897/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,229,1858,1884]" targetBox="[303,1269,309,1806]" targetPageId="40">
<paragraph id="DD6736C4FFDE892E9FBCF8F3FB1DF860" blockId="40.[189,1399,1858,2026]" pageId="40" pageNumber="41">
<emphasis id="EFACEAD6FFDE892E9FBCF8F3FED0F8D6" bold="true" box="[189,297,1858,1885]" pageId="40" pageNumber="41">Fig 16. A</emphasis>
. Protibia of
<taxonomicName id="1AD84D47FFDE892E9EB4F8F2FD6AF8D7" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[437,659,1859,1885]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFDE892E9EB4F8F2FD6AF8D7" box="[437,659,1859,1885]" italics="true" pageId="40" pageNumber="41">Onitis bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDE892E9D98F8F2FD08F8D7" box="[665,753,1859,1885]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
showing the position of spur, NIM/NBAIR/COL/ONIT/ H-271023A.
<emphasis id="EFACEAD6FFDE892E9E5BF8D7FE94F80A" bold="true" box="[346,365,1894,1920]" pageId="40" pageNumber="41">B</emphasis>
. Anterior tip of protibia of
<taxonomicName id="1AD84D47FFDE892E9DC3F8D7FB9BF80A" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[706,1122,1894,1920]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFDE892E9DC3F8D7FC5DF80A" box="[706,932,1894,1920]" italics="true" pageId="40" pageNumber="41">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFDE892E9CB1F8D7FB9BF80A" author="Janssens A." box="[944,1122,1894,1920]" pageId="40" pageNumber="41" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000492.
<emphasis id="EFACEAD6FFDE892E9FBCF838FF2BF829" bold="true" box="[189,210,1929,1955]" pageId="40" pageNumber="41">C</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9E74F83BFE1CF829" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[373,485,1929,1956]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov">
<emphasis id="EFACEAD6FFDE892E9E74F83BFE1CF829" box="[373,485,1929,1956]" italics="true" pageId="40" pageNumber="41">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDE892E9EF3F83BFDB5F82E" box="[498,588,1930,1956]" pageId="40" pageNumber="41" rank="species">sp. nov</taxonomicNameLabel>
, NIM/NBAIR/COL/ONIT/H-271023B.
<emphasis id="EFACEAD6FFDE892E9B2BF838FBC6F829" bold="true" box="[1066,1087,1929,1955]" pageId="40" pageNumber="41">D</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9BE3F83BFE8DF84C" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFDE892E9BE3F83BFA8FF829" box="[1250,1398,1929,1956]" italics="true" pageId="40" pageNumber="41">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFDE892E9FBCF81DFE8DF84C" author="Fabricius J. C." box="[189,372,1964,1990]" pageId="40" pageNumber="41" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
, AIMB/Co/Sc1000317.
<emphasis id="EFACEAD6FFDE892E9D8FF81DFD58F84C" bold="true" box="[654,673,1964,1990]" pageId="40" pageNumber="41">E</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9C39F81CFC45F84C" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[824,956,1964,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov">
<emphasis id="EFACEAD6FFDE892E9C39F81CFC45F84C" box="[824,956,1964,1991]" italics="true" pageId="40" pageNumber="41">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFDE892E9CC4F81CFBE3F84D" box="[965,1050,1965,1991]" pageId="40" pageNumber="41" rank="species">sp. nov</taxonomicNameLabel>
, NIM/NBAIR/COL/ONIT/H- 271023C.
<emphasis id="EFACEAD6FFDE892E9E34F861FEBEF860" bold="true" box="[309,327,2000,2026]" pageId="40" pageNumber="41">F</emphasis>
. Mid-leg of
<taxonomicName id="1AD84D47FFDE892E9ED6F861FCA7F863" authority="Lansberge, AIMB" authorityName="Lansberge, AIMB" authorityYear="1875" box="[471,862,1999,2026]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFDE892E9ED6F861FD71F863" box="[471,648,1999,2026]" italics="true" pageId="40" pageNumber="41">O. singhalensis</emphasis>
Lansberge, AIMB
</taxonomicName>
/Co/Sc1000684. (Images: AIMB).
</paragraph>
</caption>
<subSubSection id="95C2654FFFDF892F9FBCFEDCFE7EFC74" pageId="41" pageNumber="42" type="description">
<paragraph id="DD6736C4FFDF892F9FBCFEDCFEB7FE0D" blockId="41.[189,334,365,391]" box="[189,334,365,391]" pageId="41" pageNumber="42">
<heading id="862F81A8FFDF892F9FBCFEDCFEB7FE0D" bold="true" box="[189,334,365,391]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EFACEAD6FFDF892F9FBCFEDCFEB7FE0D" bold="true" box="[189,334,365,391]" pageId="41" pageNumber="42">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFE2DFCCBFE3D" blockId="41.[189,818,412,439]" box="[189,818,412,439]" pageId="41" pageNumber="42">MEASUREMENTS (in mm). TL=17.5–23; BW= 9–12.5.</paragraph>
<paragraph id="DD6736C4FFDF892F9FEBFE6FFED3FE72" blockId="41.[189,1399,478,785]" box="[234,298,478,504]" pageId="41" pageNumber="42">
<emphasis id="EFACEAD6FFDF892F9FEBFE6FFED3FE72" bold="true" box="[234,298,478,504]" pageId="41" pageNumber="42">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFDB0FC8CFC9B" blockId="41.[189,1399,478,785]" pageId="41" pageNumber="42">Black or reddish black, moderately shining; clypeal margin slightly reflexed; frontal carina broadly interrupted in the middle with an anterior short curved clypeal carina and transverse tubercle behind; pronotum strongly punctate anteriorly, basal foveae rounded, closer, lateral borders of pronotum slightly concave before anterior angles; elytral striae broad, shallow, interval imperceptibly punctured; pygidium flat, opaque; metasternum flat in the middle with an anterior weak longitudinal groove, metasternal shield smooth with scattered punctures, sides of the metasternum punctured with long rusty hairs. Clypeus evenly rugulose, pronotum with a broad smooth longitudinal median line, mid legs toothed, trochanters of mid leg long while that of hind legs serrated.</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFC88FB70FCFC" blockId="41.[189,1399,824,886]" pageId="41" pageNumber="42">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFDF892F9EADFC88FDD7FCD9" box="[428,558,825,851]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. E–H. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. I–L. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. M–P. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="41" pageNumber="42">Fig. 6M–P</figureCitation>
). LP=3, Lp= 2.5; BP=1.25, BpB=0.5, BpT =0.4.Parameres longer than the phallobase, symmetrical, straight above, rounded beneath and curved at the tip.
</paragraph>
<paragraph id="DD6736C4FFDF892F9FEBFC2CFEBCFC3D" blockId="41.[189,1398,925,1022]" box="[234,325,925,951]" pageId="41" pageNumber="42">
<emphasis id="EFACEAD6FFDF892F9FEBFC2CFEBCFC3D" bold="true" box="[234,325,925,951]" pageId="41" pageNumber="42">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFC71FE7EFC74" blockId="41.[189,1398,925,1022]" pageId="41" pageNumber="42">Clypeus parabolic, closely rugose, pronotal punctures stronger and closer than in male, protibia with four strong teeth.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDF892F9FBCFB94FB77FB1B" pageId="41" pageNumber="42" type="distribution">
<paragraph id="DD6736C4FFDF892F9FBCFB94FEAFFBB5" blockId="41.[189,342,1061,1087]" box="[189,342,1061,1087]" pageId="41" pageNumber="42">
<heading id="862F81A8FFDF892F9FBCFB94FEAFFBB5" bold="true" box="[189,342,1061,1087]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="EFACEAD6FFDF892F9FBCFB94FEAFFBB5" bold="true" box="[189,342,1061,1087]" pageId="41" pageNumber="42">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDF892F9FBCFBE5FB77FB1B" blockId="41.[189,1398,1108,1170]" pageId="41" pageNumber="42">
<collectingCountry id="A5CF7654FFDF892F9FBCFBE5FF07FBE4" box="[189,254,1108,1134]" name="India" pageId="41" pageNumber="42">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFDF892F9E0AFBE5FE09FBE4" box="[267,496,1108,1134]" country="India" name="Arunachal Pradesh" pageId="41" pageNumber="42">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFDF892F9EFCFBE5FDA9FBE4" box="[509,592,1108,1134]" country="India" name="Assam" pageId="41" pageNumber="42">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFDF892F9D5EFBE5FD40FBE4" box="[607,697,1108,1134]" country="India" name="Tripura" pageId="41" pageNumber="42">Tripura</collectingRegion>
,
<collectingRegion id="1F1CF826FFDF892F9DC6FBE5FC9AFBE4" box="[711,867,1108,1134]" country="India" name="West Bengal" pageId="41" pageNumber="42">West Bengal</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFDF892F9B3FFBE5FB58FBE4" box="[1086,1185,1108,1134]" name="Pakistan" pageId="41" pageNumber="42">Pakistan</collectingCountry>
(
<bibRefCitation id="B9494B35FFDF892F9BB5FBE5FF02FB1B" author="Bezdek A. &amp; Krell F. - T." pageId="41" pageNumber="42" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9E06FBC6FE0BFB1B" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[263,498,1143,1169]" pageId="41" pageNumber="42" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFDF892F9E73FBC9FE54FB1B" box="[370,429,1143,1169]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9EFFFBC6FD34FB18" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[510,717,1143,1170]" pageId="41" pageNumber="42" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFDF892F9D4CFBC9FD71FB1B" box="[589,648,1143,1169]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9DD8FBC6FC8EFB1B" author="Bezdek A." box="[729,887,1143,1169]" pageId="41" pageNumber="42" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
;
<bibRefCitation id="B9494B35FFDF892F9C82FBC6FB7BFB1B" author="Schoolmeesters P." box="[899,1154,1143,1169]" pageId="41" pageNumber="42" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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