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<document id="14727CCB43F76B917F3D508F1569C755" ID-CLB-Dataset="51393" ID-DOI="10.11646/zootaxa.4097.3.1" ID-GBIF-Dataset="4dee4a7f-ff4f-4c5a-8d5e-0044f5103e8b" ID-ISSN="1175-5326" ID-Zenodo-Dep="263937" ID-ZooBank="7DC7B6D8-6C7B-423C-A85E-EBE555FA3CCD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459493782774" checkinUser="plazi" docAuthor="Ariyama, Hiroyuki" docDate="2016" docId="0388D03FFFD2FFE682DFFC5EFD51FDBF" docLanguage="en" docName="zootaxa.4097.3.1.pdf.imf" docOrigin="Zootaxa 4097 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyproidea okinawensis Ariyama, 2016, sp. nov." docType="treatment" docVersion="9" lastPageNumber="313" masterDocId="FFB1A847FFDAFFEA8248FFB3FFDAFA69" masterDocTitle="Five species of the family Cyproideidae (Crustacea: Amphipoda) from Japan, with the description of a new genus and two new species" masterLastPageNumber="331" masterPageNumber="301" pageNumber="309" updateTime="1698202782737" updateUser="ExternalLinkService">
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<mods:title id="3E7B96FBF9EBC4095D2A5FF07E3A8923">Five species of the family Cyproideidae (Crustacea: Amphipoda) from Japan, with the description of a new genus and two new species</mods:title>
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<mods:namePart id="C437A98E64DEF1DC31C9793CD4FE3111">Ariyama, Hiroyuki</mods:namePart>
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<mods:date id="0B087424CF846A08A3E1DD51D9651017">2016</mods:date>
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<treatment id="0388D03FFFD2FFE682DFFC5EFD51FDBF" ID-DOI="http://doi.org/10.5281/zenodo.6056317" ID-GBIF-Taxon="119330794" ID-Zenodo-Dep="6056317" LSID="urn:lsid:plazi:treatment:0388D03FFFD2FFE682DFFC5EFD51FDBF" httpUri="http://treatment.plazi.org/id/0388D03FFFD2FFE682DFFC5EFD51FDBF" lastPageId="12" lastPageNumber="313" pageId="8" pageNumber="309">
<subSubSection id="C33B32A2FFD2FFE282DFFC5EFECDFE25" pageId="8" pageNumber="309" type="nomenclature">
<paragraph id="8B9E6129FFD2FFE282DFFC5EFDC8FE6E" blockId="8.[151,669,1005,1101]" box="[151,530,1005,1031]" pageId="8" pageNumber="309">
<heading id="D0D6D645FFD2FFE282DFFC5EFDC8FE6E" bold="true" box="[151,530,1005,1031]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<emphasis id="B955BD3BFFD2FFE282DFFC5EFDC8FE6E" bold="true" box="[151,530,1005,1031]" pageId="8" pageNumber="309">
<taxonomicName id="4C211AAAFFD2FFE282DFFC5EFE74FE6E" ID-CoL="33J95" box="[151,430,1005,1031]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="okinawensis" status="sp. nov.">
<emphasis id="B955BD3BFFD2FFE282DFFC5EFE74FE6E" bold="true" box="[151,430,1005,1031]" italics="true" pageId="8" pageNumber="309">Cyproidea okinawensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2660040FFD2FFE283FDFC5EFDC8FE6E" box="[437,530,1005,1031]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9E6129FFD2FFE282DFFBA3FECDFE25" blockId="8.[151,669,1005,1101]" pageId="8" pageNumber="309">
[Japanese name: Okinawa-hoteiyokoebi, new] (
<figureCitation id="131A7DACFFD2FFE282D7FB87FECAFE25" box="[159,272,1076,1101]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStartId-0="9.[151,250,1137,1159]" captionStartId-1="10.[151,250,1741,1763]" captionStartId-2="11.[151,250,1917,1939]" captionStartId-3="12.[151,250,1396,1418]" captionStartId-4="13.[151,250,1084,1106]" captionTargetBox-0="[202,1401,217,1084]" captionTargetBox-1="[151,1367,318,1695]" captionTargetBox-2="[193,1333,249,1826]" captionTargetBox-3="[152,1427,278,1395]" captionTargetBox-4="[151,1436,195,1062]" captionTargetId-0="figure@9.[151,1436,213,1115]" captionTargetId-1="figure@10.[151,1436,265,1719]" captionTargetId-2="figure@11.[193,1373,193,1883]" captionTargetId-3="figure@12.[151,1436,193,1414]" captionTargetId-4="figure@13.[151,1436,195,1062]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 7. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. Habitus, lateral view." captionText-1="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." captionText-2="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." captionText-3="FIGURE 10. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, anterior view; C, left pleopod 2, posterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsomedial view; F, left uropod 2, medial view; G, urosomite 3, telson and left uropod 3, dorsal view. Scale: A, E G, 0.69 mm; B D, 0.5 mm." captionText-4="FIGURE 11. Cyproidea okinawensis sp. nov. Not-collected individual. Type locality, 6 April 2015, photography by N. Shirakawa." httpUri-0="https://zenodo.org/record/263944/files/figure.png" httpUri-1="https://zenodo.org/record/263945/files/figure.png" httpUri-2="https://zenodo.org/record/263946/files/figure.png" httpUri-3="https://zenodo.org/record/263947/files/figure.png" httpUri-4="https://zenodo.org/record/263948/files/figure.png" pageId="8" pageNumber="309">Figs 711</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B32A2FFD2FFE282DFFBCFFAB5FF69" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="8B9E6129FFD2FFE282DFFBCFFAB5FF69" blockId="8.[151,1437,1148,2037]" pageId="8" pageNumber="309">
<emphasis id="B955BD3BFFD2FFE282DFFBCFFE5CFEFC" bold="true" box="[151,390,1148,1173]" pageId="8" pageNumber="309">Material examined.</emphasis>
<typeStatus id="549ADF8BFFD2FFE283DBFBCFFDDAFEFC" box="[403,512,1148,1173]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
: female (OMNH-Ar-9951), 5.0 mm, point &quot;Red beach&quot;, Kin Bay, Kin Town, Okinawa Prefecture,
<geoCoordinate id="EE1507EEFFD2FFE283C0FB2CFDD4FED1" box="[392,526,1183,1208]" direction="north" orientation="latitude" pageId="8" pageNumber="309" precision="15" value="26.439999">26°26'24&quot;N</geoCoordinate>
,
<geoCoordinate id="EE1507EEFFD2FFE28054FB2CFD77FED1" box="[540,685,1183,1208]" direction="east" orientation="longitude" pageId="8" pageNumber="309" precision="15" value="127.91">127°54'36&quot;E</geoCoordinate>
,
<quantity id="4CD9CCCCFFD2FFE280F3FB13FD32FEDE" box="[699,744,1184,1208]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="309" unit="m" value="3.0">3 m</quantity>
in depth, sandy bottom, using
<collectionCode id="ED30F9ECFFD2FFE28606FB13FB74FED1" box="[1102,1198,1184,1208]" pageId="8" pageNumber="309">SCUBA</collectionCode>
,
<date id="FF9F47E9FFD2FFE286F4FB13FA42FED1" box="[1212,1432,1183,1208]" pageId="8" pageNumber="309" value="2014-12-13">13 December 2014</date>
, coll. N. Shirakawa.
<typeStatus id="549ADF8BFFD2FFE2833BFB76FE3EFEB4" box="[371,484,1221,1245]" pageId="8" pageNumber="309" type="paratype">Paratypes</typeStatus>
: 2 ovigerous females (OMNH-Ar-99529953), 4.5,
<quantity id="4CD9CCCCFFD2FFE28661FB77FB5AFEB5" box="[1065,1152,1220,1244]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="309" unit="mm" value="4.3">4.3 mm</quantity>
, same data as
<typeStatus id="549ADF8BFFD2FFE28756FB77FA59FEB4" box="[1310,1411,1220,1245]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
; 2 ovigerous females (OMNH-Ar-99549955), 4.2, 4.0 mm,
<quantity id="4CD9CCCCFFD2FFE2816BFB54FC8AFE96" box="[803,848,1255,1280]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="309" unit="m" value="5.0">5 m</quantity>
,
<date id="FF9F47E9FFD2FFE28114FB54FBFCFF69" box="[860,1062,1255,1280]" pageId="8" pageNumber="309" value="2014-12-06">6 December 2014</date>
, other data same as
<typeStatus id="549ADF8BFFD2FFE2874EFB54FAB6FF69" box="[1286,1388,1255,1280]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B32A2FFD2FFE6828FFABFFE06FCFB" lastPageId="12" lastPageNumber="313" pageId="8" pageNumber="309" type="description">
<paragraph id="8B9E6129FFD2FFE2828FFABFFDA1FFDC" blockId="8.[151,1437,1148,2037]" pageId="8" pageNumber="309">
<emphasis id="B955BD3BFFD2FFE2828FFABFFE83FF4C" bold="true" box="[199,345,1292,1317]" pageId="8" pageNumber="309">Description.</emphasis>
Female [based on
<typeStatus id="549ADF8BFFD2FFE28065FABFFD48FF4C" box="[557,658,1292,1317]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
, 5.0 mm (OMNH-Ar-9951),
<typeStatus id="549ADF8BFFD2FFE28190FABEFBE1FF4C" box="[984,1083,1293,1317]" pageId="8" pageNumber="309" type="paratype">paratype</typeStatus>
,
<quantity id="4CD9CCCCFFD2FFE2860FFABFFB44FF4D" box="[1095,1182,1292,1317]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="309" unit="mm" value="4.5">4.5 mm</quantity>
(OMNH-Ar-9952) for lower lip, and
<typeStatus id="549ADF8BFFD2FFE28373FA83FE44FF21" box="[315,414,1328,1352]" pageId="8" pageNumber="309" type="paratype">paratype</typeStatus>
,
<quantity id="4CD9CCCCFFD2FFE283E2FA83FDDBFF2E" box="[426,513,1328,1352]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="309" unit="mm" value="4.3">4.3 mm</quantity>
(OMNH-Ar-9953) for mandibles].
<emphasis id="B955BD3BFFD2FFE281DBFA9CFC17FF21" box="[915,973,1327,1352]" italics="true" pageId="8" pageNumber="309">Body</emphasis>
(
<figureCitation id="131A7DACFFD2FFE28194FA9CFBFAFF21" box="[988,1056,1327,1352]" captionStart="FIGURE 7" captionStartId="9.[151,250,1137,1159]" captionTargetBox="[202,1401,217,1084]" captionTargetId="figure@9.[151,1436,213,1115]" captionTargetPageId="9" captionText="FIGURE 7. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. Habitus, lateral view." httpUri="https://zenodo.org/record/263944/files/figure.png" pageId="8" pageNumber="309">Fig. 7</figureCitation>
) oval; head longer than pereonite 1, eyes relatively small, diameter 35% length of head; pereonites 45 long, posteroventral corners of pereonites 5 7 protruded; coxa 5 barely reaching posteroventral corner of pereonite 6; pleonites 13 each with dorsolateral ridge extending to urosomite 1; urosome slender.
</paragraph>
<paragraph id="8B9E6129FFD2FFE2828FFA73FC96FC2C" blockId="8.[151,1437,1148,2037]" pageId="8" pageNumber="309">
<emphasis id="B955BD3BFFD2FFE2828FFA73FEE6FFB1" box="[199,316,1472,1496]" italics="true" pageId="8" pageNumber="309">Antenna 1</emphasis>
(
<figureCitation id="131A7DACFFD2FFE28306FA0CFE49FFB1" box="[334,403,1471,1496]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
A, A1) short, with ratio of lengths of peduncular articles 13 1:1.4:0.6, tooth of article 2 occupying quarter of whole length; accessory flagellum vestigial, with 3 setae; primary flagellum with 7 articles, articles 2, 3, 5, 6 each with 1 aesthetasc at distal end, respectively.
<emphasis id="B955BD3BFFD2FFE281C0F9BBFC21FC49" box="[904,1019,1544,1568]" italics="true" pageId="8" pageNumber="309">Antenna 2</emphasis>
(
<figureCitation id="131A7DACFFD2FFE28643F9B4FB97FC49" box="[1035,1101,1543,1568]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
B) short, with ratio of lengths of peduncular articles 35 1:1.8:1.5; flagellum with 3 articles.
</paragraph>
<paragraph id="8B9E6129FFD2FFE2828FF9E2FD17FD9C" blockId="8.[151,1437,1148,2037]" pageId="8" pageNumber="309">
<emphasis id="B955BD3BFFD2FFE2828FF9E2FEE9FC01" box="[199,307,1615,1640]" italics="true" pageId="8" pageNumber="309">Upper lip</emphasis>
(
<figureCitation id="131A7DACFFD2FFE2830AF9FCFE5EFC01" box="[322,388,1615,1640]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
C), ventrolateral margins with short setae.
<emphasis id="B955BD3BFFD2FFE28128F9FCFC16FC01" box="[864,972,1615,1640]" italics="true" pageId="8" pageNumber="309">Mandible</emphasis>
(
<figureCitation id="131A7DACFFD2FFE28193F9FCFBC6FC01" box="[987,1052,1615,1640]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
D, D1, E, E1), incisor bearing 67 cusps, left lacinia mobilis fan-shaped, distal margin serrated, right lacinia mobilis lacking, number of both accessory blades 6 (
<typeStatus id="549ADF8BFFD2FFE2833DF92BFE00FCD9" box="[373,474,1688,1712]" pageId="8" pageNumber="309" type="paratype">paratype</typeStatus>
,
<quantity id="4CD9CCCCFFD2FFE283ACF92BFDE6FCD9" box="[484,572,1688,1712]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="309" unit="mm" value="4.3">4.3 mm</quantity>
) or 10 (
<typeStatus id="549ADF8BFFD2FFE280D9F924FD2DFCD9" box="[657,759,1687,1712]" pageId="8" pageNumber="309" type="holotype">holotype</typeStatus>
, 5.0 mm); palp article length ratio 1:1.3:
<quantity id="4CD9CCCCFFD2FFE286FDF92BFB23FCD9" box="[1205,1273,1687,1712]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" pageId="8" pageNumber="309" unit="in" value="1.1">1.1 in</quantity>
right, article 2 without setae, tip of article 3 with a long seta.
<emphasis id="B955BD3BFFD2FFE28081F90EFCE3FCBC" box="[713,825,1724,1749]" italics="true" pageId="8" pageNumber="309">Lower lip</emphasis>
(
<figureCitation id="131A7DACFFD2FFE28106F90FFC4EFCBD" box="[846,916,1724,1749]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
F) wide; inner lobes indistinct; anteromedial surfaces of outer lobes setose, each with 1 round and 1 truncate projections on corner, mandibular processes small.
<emphasis id="B955BD3BFFD2FFE282DFF8B7FEDFFD75" box="[151,261,1796,1821]" italics="true" pageId="8" pageNumber="309">Maxilla 1</emphasis>
(
<figureCitation id="131A7DACFFD2FFE2835FF8B7FE83FD75" box="[279,345,1796,1821]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
G, G1
<emphasis id="B955BD3BFFD2FFE283D7F8B6FE77FD75" box="[415,429,1797,1820]" italics="true" pageId="8" pageNumber="309"></emphasis>
2), inner plate rounded, with 4 setae on tip; mediodistal corner of outer plate projected, distal margin with 7 robust setae, medial margin bearing 5 robust setae; distal end of palp projected in middle, with 3 minutely-serrated robust setae and 3 robust setae (+1 additional spine in left).
<emphasis id="B955BD3BFFD2FFE28679F8FFFB45FD0D" box="[1073,1183,1868,1893]" italics="true" pageId="8" pageNumber="309">Maxilla 2</emphasis>
(
<figureCitation id="131A7DACFFD2FFE286F9F8FFFB2EFD0D" box="[1201,1268,1868,1893]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">Fig. 8</figureCitation>
H), inner plate shorter and wider than outer, terminal ends of both plates each with a few setae, dorsodistal surfaces of both plates setose.
<emphasis id="B955BD3BFFD2FFE282A2F827FEBEFDC4" box="[234,356,1940,1965]" italics="true" pageId="8" pageNumber="309">Maxilliped</emphasis>
(
<figureCitation id="131A7DACFFD2FFE2833BF827FE6CFDC5" box="[371,438,1940,1965]" captionStart="FIGURE 8" captionStartId="10.[151,250,1741,1763]" captionTargetBox="[151,1367,318,1695]" captionTargetId="figure@10.[151,1436,265,1719]" captionTargetPageId="10" captionText="FIGURE 8. Cyproidea okinawensis sp. nov. All except for D 1, E 1 and F, holotype, female (OMNH-Ar- 9951), 5.0 mm; D 1, E 1, paratype, female (OMNH-Ar- 9953), 4.3 mm; F, paratype, female (OMNH-Ar- 9952), 4.5 mm. A, left antenna 1, lateral view; A 1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D 1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E 1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G 1, distal part of right maxilla 1, ventral view, normal setae omitted; G 2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I 1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I 2, distal part of inner plate of right maxilliped, ventral view. Scale: A B, 0.25 mm; C I, 0.22 mm; A 1, 0.125 mm; D 1, E 1, G 1 2, I 1 2, 0.11 mm." httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="8" pageNumber="309">
Fig.
<date id="FF9F47E9FFD2FFE283E0F827FE6CFDC5" box="[424,438,1940,1964]" pageId="8" pageNumber="309">8</date>
</figureCitation>
I,
<date id="FF9F47E9FFD2FFE28387F827FDDEFDC5" box="[463,516,1940,1964]" pageId="8" pageNumber="309">
I1
<emphasis id="B955BD3BFFD2FFE283AEF826FE2EFDC5" box="[486,500,1941,1964]" italics="true" pageId="8" pageNumber="309"></emphasis>
2
</date>
), inner plate truncate, mediodistal corner with 2 robust setae dorsally; outer plate with 2 distal robust setae, mediodistal margin serrated; palp articles 1
<emphasis id="B955BD3BFFD2FFE281E2F80AFC62FDB9" box="[938,952,1977,2000]" italics="true" pageId="8" pageNumber="309"></emphasis>
2 with 3 and 6 setae, respectively, article 3 covered with short setae dorsally, article 4 slender.
</paragraph>
<caption id="DF5E31A1FFD3FFE382DFFBC2FAC9FEEE" box="[151,1299,1137,1159]" httpUri="https://zenodo.org/record/263944/files/figure.png" pageId="9" pageNumber="310" targetBox="[202,1401,217,1084]" targetPageId="9">
<paragraph id="8B9E6129FFD3FFE382DFFBC2FAC9FEEE" blockId="9.[151,1299,1137,1159]" box="[151,1299,1137,1159]" pageId="9" pageNumber="310">
<emphasis id="B955BD3BFFD3FFE382DFFBC2FEC8FEEF" bold="true" box="[151,274,1137,1159]" pageId="9" pageNumber="310">FIGURE 7.</emphasis>
<taxonomicName id="4C211AAAFFD3FFE38351FBC2FDDFFEEE" box="[281,517,1137,1159]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="okinawensis" status="sp. nov.">
<emphasis id="B955BD3BFFD3FFE38351FBC2FDDFFEEE" box="[281,517,1137,1159]" italics="true" pageId="9" pageNumber="310">Cyproidea okinawensis</emphasis>
</taxonomicName>
<emphasis id="B955BD3BFFD3FFE38043FBC2FD86FEEF" bold="true" box="[523,604,1137,1158]" pageId="9" pageNumber="310">
<taxonomicNameLabel id="A2660040FFD3FFE38043FBC2FD86FEEF" box="[523,604,1137,1158]" pageId="9" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female (OMNH-Ar-9951), 5.0 mm. Habitus, lateral view.
</paragraph>
</caption>
<paragraph id="8B9E6129FFD3FFE3828FFB1DFCA2FF8E" blockId="9.[151,1437,1198,2015]" pageId="9" pageNumber="310">
<emphasis id="B955BD3BFFD3FFE3828FFB1DFE86FEAE" box="[199,348,1198,1223]" italics="true" pageId="9" pageNumber="310">Gnathopod 1</emphasis>
(
<figureCitation id="131A7DACFFD3FFE38326FB1DFE69FEAE" box="[366,435,1198,1223]" captionStart="FIGURE 9" captionStartId="11.[151,250,1917,1939]" captionTargetBox="[193,1333,249,1826]" captionTargetId="figure@11.[193,1373,193,1883]" captionTargetPageId="11" captionText="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
A, A1), coxa roundish pentagonal; basis long, slightly curved posteriorly; posterodistal corner of ischium not projected; merus short; carpus long, posterodistal corner slightly protruded, bearing 3 long setae, posteromedial surface covered with thin setae; propodus long, about 1.5 times as long as carpus, palm with many acute denticles, anteromedial surface covered with thin setae; dactylus long, slightly curved, posteroproximal margin bearing 1 long and 6 short acute spines.
<emphasis id="B955BD3BFFD3FFE38095FA8DFCACFF3E" box="[733,886,1342,1367]" italics="true" pageId="9" pageNumber="310">Gnathopod 2</emphasis>
(
<figureCitation id="131A7DACFFD3FFE381C2FA8DFC09FF3E" box="[906,979,1342,1367]" captionStart="FIGURE 9" captionStartId="11.[151,250,1917,1939]" captionTargetBox="[193,1333,249,1826]" captionTargetId="figure@11.[193,1373,193,1883]" captionTargetPageId="11" captionText="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
B, B1) stouter than gnathopod 1; coxa trapezoidal; posterodistal corner of ischium strongly projected, with several setae; merus short; posterodistal projection of carpus longer than carpus proper, bearing 4 medial setae, posteromedial surface covered with thin setae; propodus relatively wide, palm with many blunt denticles; dactylus strongly flexed at a point of one fourth, gradually narrower, posteroproximal margin bearing 10 denticles.
</paragraph>
<paragraph id="8B9E6129FFD3FFE3828FFA41FAF3FD6E" blockId="9.[151,1437,1198,2015]" pageId="9" pageNumber="310">
<emphasis id="B955BD3BFFD3FFE3828FFA41FE89FC62" box="[199,339,1522,1547]" italics="true" pageId="9" pageNumber="310">Pereopod 3</emphasis>
(
<figureCitation id="131A7DACFFD3FFE38325FA41FE66FC62" box="[365,444,1522,1547]" captionStart="FIGURE 9" captionStartId="11.[151,250,1917,1939]" captionTargetBox="[193,1333,249,1826]" captionTargetId="figure@11.[193,1373,193,1883]" captionTargetPageId="11" captionText="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
C), coxa triangular, anterior margin rounded, posterior margin slightly concave, posteroproximal corner with small projection, posteroventral corner with small medial hollow; basis and merus
<emphasis id="B955BD3BFFD3FFE387C6F9A4FA46FC47" box="[1422,1436,1559,1582]" italics="true" pageId="9" pageNumber="310"></emphasis>
propodus long.
<emphasis id="B955BD3BFFD3FFE38301F989FE11FC3A" box="[329,459,1594,1619]" italics="true" pageId="9" pageNumber="310">Pereopod 4</emphasis>
(
<figureCitation id="131A7DACFFD3FFE38394F989FDFBFC3A" box="[476,545,1594,1619]" captionStart="FIGURE 9" captionStartId="11.[151,250,1917,1939]" captionTargetBox="[193,1333,249,1826]" captionTargetId="figure@11.[193,1373,193,1883]" captionTargetPageId="11" captionText="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
D), coxa extremely wide, swollen posteriorly, anterior margin slightly convex, with small projection on anteroventral corner, ventral margin also slightly convex, posteroproximal corner a little projected; basis and merus
<emphasis id="B955BD3BFFD3FFE38380F930FE0CFCF3" box="[456,470,1667,1690]" italics="true" pageId="9" pageNumber="310"></emphasis>
propodus long.
<emphasis id="B955BD3BFFD3FFE380C3F931FCD4FCF3" box="[651,782,1666,1691]" italics="true" pageId="9" pageNumber="310">Pereopod 5</emphasis>
(
<figureCitation id="131A7DACFFD3FFE38168F931FCBFFCF2" box="[800,869,1666,1691]" captionStart="FIGURE 9" captionStartId="11.[151,250,1917,1939]" captionTargetBox="[193,1333,249,1826]" captionTargetId="figure@11.[193,1373,193,1883]" captionTargetPageId="11" captionText="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
E, E1), coxa lanceolate, width about 2.8 times of length, anterior margin with small hollow, posterior end pointed; basis and merus
<emphasis id="B955BD3BFFD3FFE3866AF914FBEAFCD7" box="[1058,1072,1703,1726]" italics="true" pageId="9" pageNumber="310"></emphasis>
propodus long.
<emphasis id="B955BD3BFFD3FFE38697F915FA85FCD6" box="[1247,1375,1702,1727]" italics="true" pageId="9" pageNumber="310">Pereopod 6</emphasis>
(
<figureCitation id="131A7DACFFD3FFE38726F915FF7EFC8A" captionStart="FIGURE 9" captionStartId="11.[151,250,1917,1939]" captionTargetBox="[193,1333,249,1826]" captionTargetId="figure@11.[193,1373,193,1883]" captionTargetPageId="11" captionText="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
F, F1) slightly shorter than pereopod 5; coxa roundish trapezoidal; basis and merus
<emphasis id="B955BD3BFFD3FFE38611F978FBBDFC8B" box="[1113,1127,1739,1762]" italics="true" pageId="9" pageNumber="310"></emphasis>
propodus long.
<emphasis id="B955BD3BFFD3FFE38751F979FA46FC8B" box="[1305,1436,1738,1763]" italics="true" pageId="9" pageNumber="310">Pereopod 7</emphasis>
(
<figureCitation id="131A7DACFFD3FFE382D7F95DFF38FD6E" box="[159,226,1774,1799]" captionStart="FIGURE 9" captionStartId="11.[151,250,1917,1939]" captionTargetBox="[193,1333,249,1826]" captionTargetId="figure@11.[193,1373,193,1883]" captionTargetPageId="11" captionText="FIGURE 9. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A B, left gnathopods 1 2, lateral views; A 1, palm and dactylus of left gnathopod 1, lateral view; B 1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; C G, left pereopods 3 7, lateral views; E 1, left gill and oostegite on coxa 5, lateral view; F 1, left gill on coxa 6, lateral view. Scale: A G, E 1, F 1, 0.5 mm; A 1, B 1, 0.25 mm." httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="9" pageNumber="310">Fig. 9</figureCitation>
G) almost same length as pereopod 6; coxa roundish trapezoidal; basis and merus
<emphasis id="B955BD3BFFD3FFE3863AF95CFB5AFD6F" box="[1138,1152,1775,1798]" italics="true" pageId="9" pageNumber="310"></emphasis>
propodus long.
</paragraph>
<paragraph id="8B9E6129FFD3FFE08299F8A1FEADFA91" blockId="9.[151,1437,1198,2015]" lastBlockId="10.[151,1437,151,248]" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310">
<emphasis id="B955BD3BFFD3FFE38299F8A1FE55FD43" box="[209,399,1810,1835]" italics="true" pageId="9" pageNumber="310">Pleonal epimera</emphasis>
(
<figureCitation id="131A7DACFFD3FFE383E9F8A1FE2CFD42" box="[417,502,1810,1835]" captionStart="FIGURE 10" captionStartId="12.[151,250,1396,1418]" captionTargetBox="[152,1427,278,1395]" captionTargetId="figure@12.[151,1436,193,1414]" captionTargetPageId="12" captionText="FIGURE 10. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, anterior view; C, left pleopod 2, posterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsomedial view; F, left uropod 2, medial view; G, urosomite 3, telson and left uropod 3, dorsal view. Scale: A, E G, 0.69 mm; B D, 0.5 mm." httpUri="https://zenodo.org/record/263947/files/figure.png" pageId="9" pageNumber="310">Fig. 10</figureCitation>
A), each with oblique lateral ridge (short in epimeron 1); epimera 12 projected bluntly and epimeron 3 rounded each on posteroventral corner; all ventral margins bare.
<emphasis id="B955BD3BFFD3FFE386C4F885FB2EFD26" box="[1164,1268,1846,1871]" italics="true" pageId="9" pageNumber="310">Pleopods</emphasis>
(
<figureCitation id="131A7DACFFD3FFE3874DF885FA8DFD26" box="[1285,1367,1846,1871]" captionStart="FIGURE 10" captionStartId="12.[151,250,1396,1418]" captionTargetBox="[152,1427,278,1395]" captionTargetId="figure@12.[151,1436,193,1414]" captionTargetPageId="12" captionText="FIGURE 10. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, anterior view; C, left pleopod 2, posterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsomedial view; F, left uropod 2, medial view; G, urosomite 3, telson and left uropod 3, dorsal view. Scale: A, E G, 0.69 mm; B D, 0.5 mm." httpUri="https://zenodo.org/record/263947/files/figure.png" pageId="9" pageNumber="310">Fig. 10</figureCitation>
B
<emphasis id="B955BD3BFFD3FFE38721F884FAADFD27" box="[1385,1399,1847,1870]" italics="true" pageId="9" pageNumber="310"></emphasis>
D), pleopod 3 a little shorter than pleopods 12; peduncles of pleopods 23 bearing a few setae; inner and outer rami with 10, 9, 9 and 11, 11, 12 articles in pleopods 1
<emphasis id="B955BD3BFFD3FFE3808EF8CCFD0EFDFF" box="[710,724,1919,1942]" italics="true" pageId="9" pageNumber="310"></emphasis>
3, respectively.
<emphasis id="B955BD3BFFD3FFE381CCF8CDFC3FFDFE" box="[900,997,1918,1943]" italics="true" pageId="9" pageNumber="310">Uropods</emphasis>
(
<figureCitation id="131A7DACFFD3FFE381BEF8CDFB9CFDFE" box="[1014,1094,1918,1943]" captionStart="FIGURE 10" captionStartId="12.[151,250,1396,1418]" captionTargetBox="[152,1427,278,1395]" captionTargetId="figure@12.[151,1436,193,1414]" captionTargetPageId="12" captionText="FIGURE 10. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, anterior view; C, left pleopod 2, posterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsomedial view; F, left uropod 2, medial view; G, urosomite 3, telson and left uropod 3, dorsal view. Scale: A, E G, 0.69 mm; B D, 0.5 mm." httpUri="https://zenodo.org/record/263947/files/figure.png" pageId="9" pageNumber="310">Fig. 10</figureCitation>
E
<emphasis id="B955BD3BFFD3FFE38610F8CCFBBCFDFF" box="[1112,1126,1919,1942]" italics="true" pageId="9" pageNumber="310"></emphasis>
G), peduncle and both rami without robust setae, but bearing minute denticles; peduncle of uropod 1 long, both rami about 0.8 times as long as peduncle; uropod 2 86% length of uropod 1, both rami 94% as long as peduncle; uropod 3 short, 64% length of uropod 1, peduncle relatively short, inner ramus about 1.3 times as long as peduncle, outer ramus almost same length as peduncle.
<emphasis id="B955BD3BFFD0FFE0833DFF0FFE61FABC" box="[373,443,188,213]" italics="true" pageId="10" pageNumber="311">Telson</emphasis>
(
<figureCitation id="131A7DACFFD0FFE08385FF0FFDC7FABD" box="[461,541,188,213]" captionStart="FIGURE 10" captionStartId="12.[151,250,1396,1418]" captionTargetBox="[152,1427,278,1395]" captionTargetId="figure@12.[151,1436,193,1414]" captionTargetPageId="12" captionText="FIGURE 10. Cyproidea okinawensis sp. nov. Holotype, female (OMNH-Ar- 9951), 5.0 mm. A, right pleonal epimera 1 3, lateral view; B, left pleopod 1, anterior view; C, left pleopod 2, posterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsomedial view; F, left uropod 2, medial view; G, urosomite 3, telson and left uropod 3, dorsal view. Scale: A, E G, 0.69 mm; B D, 0.5 mm." httpUri="https://zenodo.org/record/263947/files/figure.png" pageId="10" pageNumber="311">Fig. 10</figureCitation>
G) ovoid, breadth about 0.6 times of length, without setae. Male unknown.
</paragraph>
<caption id="DF5E31A1FFD0FFE082DFF97EFCE0FDB1" httpUri="https://zenodo.org/record/263945/files/figure.png" pageId="10" pageNumber="311" targetBox="[151,1367,318,1695]" targetPageId="10">
<paragraph id="8B9E6129FFD0FFE082DFF97EFCE0FDB1" blockId="10.[151,1436,1740,2008]" pageId="10" pageNumber="311">
<emphasis id="B955BD3BFFD0FFE082DFF97EFECEFC8B" bold="true" box="[151,276,1741,1763]" pageId="10" pageNumber="311">FIGURE 8.</emphasis>
<taxonomicName id="4C211AAAFFD0FFE08354F97FFDD0FC8B" box="[284,522,1740,1762]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="okinawensis" status="sp. nov.">
<emphasis id="B955BD3BFFD0FFE08354F97FFDD0FC8B" box="[284,522,1740,1762]" italics="true" pageId="10" pageNumber="311">Cyproidea okinawensis</emphasis>
</taxonomicName>
<emphasis id="B955BD3BFFD0FFE08059F97EFDB9FC8B" bold="true" box="[529,611,1741,1762]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A2660040FFD0FFE08059F97EFDB9FC8B" box="[529,611,1741,1762]" pageId="10" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
All except for D1, E1 and F, holotype, female (OMNH-Ar-9951), 5.0 mm; D1, E1, paratype, female (OMNH-Ar-9953), 4.3 mm; F, paratype, female (OMNH-Ar-9952), 4.5 mm. A, left antenna 1, lateral view; A1, accessory flagellum of left antenna 1, medial view; B, left antenna 2, lateral view; C, upper lip, posterior view; D, left mandible, ventral view; D1, incisor, lacinia mobilis and accessory blades of left mandible, medial view; E, right mandible, dorsal view; E1, incisor and accessory blades of right mandible, medial view; F, lower lip, ventral view; G, right maxilla 1, ventral view; G1, distal part of right maxilla 1, ventral view, normal setae omitted; G2, distal part of left maxilla 1 palp, dorsal view, normal setae omitted; H, left maxilla 2, dorsal view; I, right maxilliped, ventral view; I1, distal part of outer plate of right maxilliped, ventral view, normal setae omitted; I2, distal part of inner plate of right maxilliped, ventral view. Scale: AB, 0.25 mm; CI, 0.22 mm; A1, 0.125 mm; D1, E1, G12, I12, 0.11 mm.
</paragraph>
</caption>
<caption id="DF5E31A1FFD1FFE182DFF8CEFBC4FD86" httpUri="https://zenodo.org/record/263946/files/figure.png" pageId="11" pageNumber="312" targetBox="[193,1333,249,1826]" targetPageId="11">
<paragraph id="8B9E6129FFD1FFE182DFF8CEFBC4FD86" blockId="11.[151,1436,1917,2031]" pageId="11" pageNumber="312">
<emphasis id="B955BD3BFFD1FFE182DFF8CEFEC9FDFB" bold="true" box="[151,275,1917,1939]" pageId="11" pageNumber="312">FIGURE 9.</emphasis>
<taxonomicName id="4C211AAAFFD1FFE18353F8CEFDD2FDFA" box="[283,520,1917,1939]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="okinawensis" status="sp. nov.">
<emphasis id="B955BD3BFFD1FFE18353F8CEFDD2FDFA" box="[283,520,1917,1939]" italics="true" pageId="11" pageNumber="312">Cyproidea okinawensis</emphasis>
</taxonomicName>
<emphasis id="B955BD3BFFD1FFE18047F8CEFDBBFDFB" bold="true" box="[527,609,1917,1938]" pageId="11" pageNumber="312">
<taxonomicNameLabel id="A2660040FFD1FFE18047F8CEFDBBFDFB" box="[527,609,1917,1938]" pageId="11" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female (OMNH-Ar-9951), 5.0 mm. AB, left gnathopods 12, lateral views; A1, palm and dactylus of left gnathopod 1, lateral view; B1, posterodistal projection of carpus, palm and dactylus of left gnathopod 2, lateral view, thin setae omitted; CG, left pereopods 37, lateral views; E1, left gill and oostegite on coxa 5, lateral view; F1, left gill on coxa 6, lateral view. Scale: AG, E1, F1, 0.5 mm; A1, B1, 0.25 mm.
</paragraph>
</caption>
<caption id="DF5E31A1FFD6FFE682DFFAC7FE87FF8F" httpUri="https://zenodo.org/record/263947/files/figure.png" pageId="12" pageNumber="313" targetBox="[152,1427,278,1395]" targetPageId="12">
<paragraph id="8B9E6129FFD6FFE682DFFAC7FE87FF8F" blockId="12.[151,1436,1396,1510]" pageId="12" pageNumber="313">
<emphasis id="B955BD3BFFD6FFE682DFFAC7FEF8FFE0" bold="true" box="[151,290,1396,1418]" pageId="12" pageNumber="313">FIGURE 10.</emphasis>
<taxonomicName id="4C211AAAFFD6FFE68363FAC7FDC0FFE3" box="[299,538,1396,1418]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="okinawensis" status="sp. nov.">
<emphasis id="B955BD3BFFD6FFE68363FAC7FDC0FFE3" box="[299,538,1396,1418]" italics="true" pageId="12" pageNumber="313">Cyproidea okinawensis</emphasis>
</taxonomicName>
<emphasis id="B955BD3BFFD6FFE6806BFAC7FDACFFE0" bold="true" box="[547,630,1396,1417]" pageId="12" pageNumber="313">
<taxonomicNameLabel id="A2660040FFD6FFE6806BFAC7FDACFFE0" box="[547,630,1396,1417]" pageId="12" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, female (OMNH-Ar-9951), 5.0 mm. A, right pleonal epimera 13, lateral view; B, left pleopod 1, anterior view; C, left pleopod 2, posterior view; D, left pleopod 3, posterior view; E, left uropod 1, dorsomedial view; F, left uropod 2, medial view; G, urosomite 3, telson and left uropod 3, dorsal view. Scale: A, EG, 0.69 mm; BD, 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9E6129FFD6FFE6828FF9BEFE06FCFB" blockId="12.[151,1436,1549,2006]" pageId="12" pageNumber="313">
<emphasis id="B955BD3BFFD6FFE6828FF9BEFE4DFC4F" bold="true" box="[199,407,1549,1574]" pageId="12" pageNumber="313">Coloration in life</emphasis>
(
<figureCitation id="131A7DACFFD6FFE683E1F9BEFE20FC4F" box="[425,506,1549,1574]" captionStart="FIGURE 11" captionStartId="13.[151,250,1084,1106]" captionTargetBox="[151,1436,195,1062]" captionTargetId="figure@13.[151,1436,195,1062]" captionTargetPageId="13" captionText="FIGURE 11. Cyproidea okinawensis sp. nov. Not-collected individual. Type locality, 6 April 2015, photography by N. Shirakawa." httpUri="https://zenodo.org/record/263948/files/figure.png" pageId="12" pageNumber="313">Fig. 11</figureCitation>
). Eyes red; dorsolateral surfaces from head to pereonite 7 with many brown and white pigments, ventral surfaces of pereonites and whole of pleonite 1 white; anteroproximal part of coxa 3 white, with several red pigments, anteroproximal and anteroventral parts of coxa 4 white, posteroproximal part of coxa 4 with black and red pigments.
</paragraph>
</subSubSection>
<subSubSection id="C33B32A2FFD6FFE6828FF92EFD44FCDF" box="[199,670,1693,1718]" pageId="12" pageNumber="313" type="etymology">
<paragraph id="8B9E6129FFD6FFE6828FF92EFD44FCDF" blockId="12.[151,1436,1549,2006]" box="[199,670,1693,1718]" pageId="12" pageNumber="313">
<emphasis id="B955BD3BFFD6FFE6828FF92EFE94FCDF" bold="true" box="[199,334,1693,1718]" pageId="12" pageNumber="313">Etymology.</emphasis>
Referring to the
<typeStatus id="549ADF8BFFD6FFE68044F92DFDE6FCDF" box="[524,572,1694,1718]" pageId="12" pageNumber="313">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33B32A2FFD6FFE6828FF972FE47FDE7" pageId="12" pageNumber="313" type="discussion">
<paragraph id="8B9E6129FFD6FFE6828FF972FE47FDE7" blockId="12.[151,1436,1549,2006]" pageId="12" pageNumber="313">
<emphasis id="B955BD3BFFD6FFE6828FF972FEE1FCB3" bold="true" box="[199,315,1729,1754]" pageId="12" pageNumber="313">Remarks.</emphasis>
<taxonomicName id="4C211AAAFFD6FFE6830FF972FD89FCB3" box="[327,595,1729,1754]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="okinawensis" status="sp. nov.">
<emphasis id="B955BD3BFFD6FFE6830FF972FD89FCB3" box="[327,595,1729,1754]" italics="true" pageId="12" pageNumber="313">Cyproidea okinawensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2660040FFD6FFE68017F972FD65FCB3" box="[607,703,1729,1754]" pageId="12" pageNumber="313" rank="species">
<emphasis id="B955BD3BFFD6FFE68017F972FD65FCB3" bold="true" box="[607,703,1729,1754]" pageId="12" pageNumber="313">sp. nov.</emphasis>
</taxonomicNameLabel>
has the gnathopod 2 with both of a posterodistally-projected ischium and a long flexed dactylus. Only two species of
<taxonomicName id="4C211AAAFFD6FFE68109F956FC6CFC97" box="[833,950,1765,1790]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
<emphasis id="B955BD3BFFD6FFE68109F956FC6CFC97" box="[833,950,1765,1790]" italics="true" pageId="12" pageNumber="313">Cyproidea</emphasis>
</taxonomicName>
,
<taxonomicName id="4C211AAAFFD6FFE68181F955FB84FC97" box="[969,1118,1765,1790]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="liodactyla">
<emphasis id="B955BD3BFFD6FFE68181F955FB84FC97" box="[969,1118,1765,1790]" italics="true" pageId="12" pageNumber="313">C. liodactyla</emphasis>
</taxonomicName>
and
<taxonomicName id="4C211AAAFFD6FFE686D6F955FA46FC97" authority="Ren, 2006" authorityName="Ren" authorityYear="2006" box="[1182,1436,1765,1790]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B955BD3BFFD6FFE686D6F955FAC3FC97" box="[1182,1305,1765,1790]" italics="true" pageId="12" pageNumber="313">C. robusta</emphasis>
<bibRefCitation id="EFB01CD8FFD6FFE6876CF955FA46FC97" author="Ren" box="[1316,1436,1765,1790]" pageId="12" pageNumber="331" refString="Ren, X. (2006) Fauna Sinica, Invertebrata Vol. 41. Crustacea: Amphipoda: Gammaridea (I). Science Press, Beijing, 588 pp. [in Chinese with English abstract and descriptions of new species]" type="book" year="2006">Ren, 2006</bibRefCitation>
</taxonomicName>
collected from
<collectingCountry id="F33621B9FFD6FFE6830CF8BAFE50FD4B" box="[324,394,1801,1826]" name="China" pageId="12" pageNumber="313">China</collectingCountry>
, have such a gnathopod 2. However, this new species is different from
<taxonomicName id="4C211AAAFFD6FFE686F7F8B9FA89FD4B" box="[1215,1363,1801,1826]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="liodactyla">
<emphasis id="B955BD3BFFD6FFE686F7F8B9FA89FD4B" box="[1215,1363,1801,1826]" italics="true" pageId="12" pageNumber="313">C. liodactyla</emphasis>
</taxonomicName>
in the smaller eyes, the narrower coxa 5 and the coloration. Although the coloration of
<taxonomicName id="4C211AAAFFD6FFE68668F89DFB4FFD2F" box="[1056,1173,1837,1862]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B955BD3BFFD6FFE68668F89DFB4FFD2F" box="[1056,1173,1837,1862]" italics="true" pageId="12" pageNumber="313">C. robusta</emphasis>
</taxonomicName>
resembles (milk white, with points of darkish-brown pigments), this species can be distinguished from
<taxonomicName id="4C211AAAFFD6FFE68670F8E1FB6BFD03" box="[1080,1201,1873,1898]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B955BD3BFFD6FFE68670F8E1FB6BFD03" box="[1080,1201,1873,1898]" italics="true" pageId="12" pageNumber="313">C. robusta</emphasis>
</taxonomicName>
by the ovoid telson (pointed in
<taxonomicName id="4C211AAAFFD6FFE68350F8C5FE56FDE7" box="[280,396,1909,1934]" class="Malacostraca" family="Cyproideidae" genus="Cyproidea" kingdom="Animalia" order="Amphipoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B955BD3BFFD6FFE68350F8C5FE56FDE7" box="[280,396,1909,1934]" italics="true" pageId="12" pageNumber="313">C. robusta</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33B32A2FFD6FFE6828FF82AFD51FDBF" pageId="12" pageNumber="313" type="distribution">
<paragraph id="8B9E6129FFD6FFE6828FF82AFC19FDDB" blockId="12.[151,1436,1549,2006]" box="[199,963,1945,1970]" pageId="12" pageNumber="313">
<emphasis id="B955BD3BFFD6FFE6828FF82AFEBAFDDB" bold="true" box="[199,352,1945,1970]" pageId="12" pageNumber="313">Distribution.</emphasis>
<collectingCountry id="F33621B9FFD6FFE6832FF829FE71FDD8" box="[359,427,1946,1969]" name="Japan" pageId="12" pageNumber="313">Japan</collectingCountry>
: only Kin Town in Okinawa Prefecture (
<figureCitation id="131A7DACFFD6FFE68139F82AFC68FDDB" box="[881,946,1945,1970]" captionStart="FIGURE 1" captionStartId="1.[151,250,1553,1575]" captionTargetBox="[152,1362,189,1552]" captionTargetId="figure@1.[151,1436,113,1606]" captionTargetPageId="1" captionText="FIGURE 1. Map showing collecting sites of the examined specimens." httpUri="https://zenodo.org/record/263938/files/figure.png" pageId="12" pageNumber="313">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B9E6129FFD6FFE6828FF80EFD51FDBF" blockId="12.[151,1436,1549,2006]" box="[199,651,1981,2006]" pageId="12" pageNumber="313">
<emphasis id="B955BD3BFFD6FFE6828FF80EFEF0FDBF" bold="true" box="[199,298,1981,2006]" pageId="12" pageNumber="313">Habitat.</emphasis>
Sandy bottom,
<quantity id="4CD9CCCCFFD6FFE68393F80EFDF8FDBC" box="[475,546,1981,2006]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="12" pageNumber="313" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">35 m</quantity>
in depth.
</paragraph>
</subSubSection>
</treatment>
</document>