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<document id="F0FB00741437861CADE5560363AE31A1" ID-CLB-Dataset="54828" ID-DOI="10.5733/afin.054.0101" ID-GBIF-Dataset="753510a6-cffa-4637-a1bd-e9ae7d5d1660" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670100" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639734503" checkinUser="felipe" docAuthor="Griffiths, O. L. &amp; Herbert, D. G." docDate="2013" docId="03BC1F3CE533D90C078AFB1D476DB3A9" docLanguage="en" docName="AfrInvertebr.54.1.1-48.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Ampelita milloti : Fischer-Piette 1952" docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="FF856744E529D917064DFF964621B743" masterDocTitle="New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics" masterLastPageNumber="48" masterPageNumber="1" pageNumber="26" updateTime="1699475339093" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1848810920D1AB180129F1434EC35D01">New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics</mods:title>
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<mods:namePart id="8CFB44187D299134EDFA48F9E49BE2BC">Griffiths, O. L.</mods:namePart>
<mods:affiliation id="1659E8D1DAC7637D12C558B059021CC0">Research associate of the Australian Museum, Sydney, Australia and Biodiversity Conservation Madagascar (BCM) Ltd, Rivière des Anguilles, Mauritius</mods:affiliation>
<mods:nameIdentifier id="5D0452F8EC12A95E40F530581F21E027" type="email">owen@bcm.intnet.mu</mods:nameIdentifier>
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<mods:namePart id="BF7CEB50173063EED0CF55A42B7F9F1F">Herbert, D. G.</mods:namePart>
<mods:affiliation id="5CC3D7266E5D072E188430AEE57F5E2E">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200 South Africa and School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, 3206 South Africa</mods:affiliation>
<mods:nameIdentifier id="1ABD2618049C880B77B76E860457FB89" type="email">dherbert@nmsa.org.za</mods:nameIdentifier>
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<mods:title id="B760A8C4198700B3B1BEEF1699101AEC">African Invertebrates</mods:title>
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<mods:date id="30442373C4C08ADB28FA47324B41E055">2013</mods:date>
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<mods:identifier id="3C7631A860F84CEF828096570E675AB6" type="DOI">10.5733/afin.054.0101</mods:identifier>
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<treatment id="03BC1F3CE533D90C078AFB1D476DB3A9" ID-DOI="http://doi.org/10.5281/zenodo.7649551" ID-GBIF-Taxon="209341527" ID-Zenodo-Dep="7649551" LSID="urn:lsid:plazi:treatment:03BC1F3CE533D90C078AFB1D476DB3A9" httpUri="http://treatment.plazi.org/id/03BC1F3CE533D90C078AFB1D476DB3A9" lastPageId="27" lastPageNumber="27" pageId="26" pageNumber="26">
<subSubSection id="C30FFDA1E533D90D078AFB1D4545B3E0" box="[455,868,1161,1187]" pageId="26" pageNumber="26" type="nomenclature">
<paragraph id="8BAAAE2AE533D90D078AFB1D4545B3E0" blockId="26.[455,868,1161,1187]" box="[455,868,1161,1187]" pageId="26" pageNumber="26">
<heading id="D0E21946E533D90D078AFB1D4545B3E0" box="[455,868,1161,1187]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="26" reason="3">
<taxonomicName id="4C15D5A9E533D90D078AFB1D4545B3E0" ID-CoL="66DHM" ID-ENA="1642594" authority="Fischer-Piette, 1952" authorityName=": Fischer-Piette" authorityYear="1952" box="[455,868,1161,1187]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="milloti">
<emphasis id="B9617238E533D90D078AFB1D445DB3E0" box="[455,636,1163,1187]" italics="true" pageId="26" pageNumber="26">Ampelita milloti</emphasis>
Fischer-Piette, 1952
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30FFDA1E533D90C045CFB27476DB3A9" lastPageId="27" lastPageNumber="27" pageId="26" pageNumber="26" type="description">
<paragraph id="8BAAAE2AE533D90D045CFB27453BB388" blockId="26.[181,1142,1201,1675]" box="[529,794,1201,1227]" pageId="26" pageNumber="26">
<figureCitation id="132EB2AFE533D90D045CFB274496B388" box="[529,695,1201,1227]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="26" pageNumber="26">Figs 17C, 17D</figureCitation>
,
<geoCoordinate id="EE21C8EDE533D90D048EFB27453BB388" box="[707,794,1201,1227]" degrees="18, 25" direction="east" orientation="longitude" pageId="26" pageNumber="26" precision="555" value="18.25">
<figureCitation id="132EB2AFE533D90D048EFB2744C0B388" box="[707,737,1201,1227]" captionStart="Fig" captionStartId="27.[144,177,509,530]" captionTargetBox="[144,1103,211,481]" captionTargetId="figure-359@27.[143,1103,204,488]" captionTargetPageId="27" captionText="Fig. 18. Ampelita milloti Fischer-Piette, 1952, Tsingy Beanka, st’n 16/06, max. diameter 24.8 mm, NMSA L7127." figureDoi="http://doi.org/10.5281/zenodo.7670144" httpUri="https://zenodo.org/record/7670144/files/figure.png" pageId="26" pageNumber="26">18</figureCitation>
,
<figureCitation id="132EB2AFE533D90D04A0FB27453BB388" box="[749,794,1201,1227]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, st’n 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., st’n 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette &amp; Bedoucha, 1965, st’n 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, st’n 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,st’n 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, st’n 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, st’n 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., st’n 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="26" pageNumber="26">25E</figureCitation>
</geoCoordinate>
</paragraph>
<paragraph id="8BAAAE2AE533D90D06F8FB4E4587B240" blockId="26.[181,1142,1201,1675]" pageId="26" pageNumber="26">
<taxonomicName id="4C15D5A9E533D90D06F8FB4E4409B3A8" authority=": Fischer-Piette 1952: 34" authorityName=": Fischer-Piette" authorityPageNumber="34" authorityYear="1952" box="[181,552,1238,1259]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="milloti">
<emphasis id="B9617238E533D90D06F8FB4E4769B3A8" box="[181,328,1240,1259]" italics="true" pageId="26" pageNumber="26">Ampelita milloti</emphasis>
:
<materialsCitation id="3B7DA477E533D90D071EFB404409B3A8" box="[339,552,1238,1259]" collectorName="de Salapango" country="Madagascar" location="Gorge de Salapango" pageId="26" pageNumber="26" specimenCount="1">Fischer-Piette 1952: 34</materialsCitation>
</taxonomicName>
, pl. 3, figs 31–34; Fischer-Piette
<emphasis id="B9617238E533D90D051DFB4E4559B3A8" box="[848,888,1240,1259]" italics="true" pageId="26" pageNumber="26">et al</emphasis>
. 1994: 123, pl. 15, figs 1–4.
<typeStatus id="54AE1088E533D90D0758FB784763B240" box="[277,322,1262,1283]" pageId="26" pageNumber="26">Type</typeStatus>
loc.: ‘
<location id="8ECAF8F1E533D90D0734FB784415B240" LSID="urn:lsid:plazi:treatment:03BC1F3CE533D90C078AFB1D476DB3A9:8ECAF8F1E533D90D0734FB784415B240" box="[377,564,1262,1283]" country="Madagascar" name="Gorge de Salapango" pageId="26" pageNumber="26">
Gorge
<collectorName id="26E0CBFCE533D90D07F6FB784415B240" box="[443,564,1262,1283]" pageId="26" pageNumber="26">de Salapango</collectorName>
</location>
[= Salapanga], Bemaraha’,
<collectingCountry id="F302EEBAE533D90D057EFB784583B240" box="[819,930,1262,1283]" name="Madagascar" pageId="26" pageNumber="26">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BAAAE2AE533D90D06F8FA844787B26E" blockId="26.[181,1142,1201,1675]" box="[181,422,1298,1325]" pageId="26" pageNumber="26">Morphological notes:</paragraph>
<paragraph id="8BAAAE2AE533D90D06F8FAAA4785B2F7" blockId="26.[181,1142,1201,1675]" pageId="26" pageNumber="26">
<emphasis id="B9617238E533D90D06F8FAAA4756B217" box="[181,375,1340,1364]" italics="true" pageId="26" pageNumber="26">External features</emphasis>
(
<figureCitation id="132EB2AFE533D90D07C5FAAC47D0B217" box="[392,497,1338,1364]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette &amp; Bedoucha, 1965, st’n 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., st’n 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette &amp; Bedoucha, 1965, st’n 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, st’n 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,st’n 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, st’n 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, st’n 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., st’n 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="26" pageNumber="26">Fig. 25E</figureCitation>
): Head-foot uniformly dark, charcoal-grey to black; skin texture uniformly granular throughout; lining of mantle cavity heavily pigmented with black and patterned with pale circular blotches (observations based on a single livecollected individual).
</paragraph>
<paragraph id="8BAAAE2AE533D90D06F8FA524462B17E" blockId="26.[181,1142,1201,1675]" pageId="26" pageNumber="26">
<emphasis id="B9617238E533D90D06F8FA524740B29F" box="[181,353,1476,1500]" italics="true" pageId="26" pageNumber="26">Distal genitalia</emphasis>
: The single preserved specimen available (shell diameter
<quantity id="4CED03CFE533D90D059BFA544218B29E" box="[982,1081,1474,1501]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="26" pageNumber="26" unit="mm" value="19.2">19.2 mm</quantity>
) possessed an immature reproductive tract. The distal genitalia were thus poorly developed and inadequate for comparative study. From what could be seen of the penis, it appeared similar to that of
<taxonomicName id="4C15D5A9E533D90D073BF9B3441DB17E" authorityName=": Fischer-Piette" authorityYear="1952" box="[374,572,1573,1597]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="namerokoensis">
<emphasis id="B9617238E533D90D073BF9B3441DB17E" box="[374,572,1573,1597]" italics="true" pageId="26" pageNumber="26">A. namerokoensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAAAE2AE533D90D06F8F9D045C6B1C8" blockId="26.[181,1142,1201,1675]" pageId="26" pageNumber="26">
Locality data:
<emphasis id="B9617238E533D90D0779F9DE4791B118" box="[308,432,1608,1627]" italics="true" pageId="26" pageNumber="26">Antsingimavo</emphasis>
: st’ns 4/06, 8/06 (all subfossil).
<taxonomicName id="4C15D5A9E533D90D0486F9DE456CB118" box="[715,845,1608,1627]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="beanka">
<emphasis id="B9617238E533D90D0486F9DE456CB118" box="[715,845,1608,1627]" italics="true" pageId="26" pageNumber="26">Tsingy Beanka</emphasis>
</taxonomicName>
: st’ns 12/06, 14/06, 16/06, 18/06, 02/09, 06/09, 07/09, 08/09, 01/10, 06/10, 07/10, 08/10, 09/10.
<emphasis id="B9617238E533D90D049AF9F645AAB130" box="[727,907,1632,1651]" italics="true" pageId="26" pageNumber="26">Tsingy de Bemaraha</emphasis>
: st’ns 07/95, 10/95, 14/95, 18/95, 19/95, 9/96, 10/96, 12/96.
<emphasis id="B9617238E533D90D07A8F9EE4520B1C8" box="[485,769,1656,1675]" italics="true" pageId="26" pageNumber="26">South bank of Tsiribihina River</emphasis>
: st’ns 01/99LP, 02/99LP.
</paragraph>
<caption id="DF6AFEA2E532D90C06DDFE6B4733B569" ID-DOI="http://doi.org/10.5281/zenodo.7670144" ID-Zenodo-Dep="7670144" httpUri="https://zenodo.org/record/7670144/files/figure.png" pageId="27" pageNumber="27" startId="27.[144,177,509,530]" targetBox="[144,1103,211,481]" targetPageId="27">
<paragraph id="8BAAAE2AE532D90C06DDFE6B4733B569" blockId="27.[144,1103,509,554]" pageId="27" pageNumber="27">
Fig. 18.
<taxonomicName id="4C15D5A9E532D90C0696FE69440AB551" authority="Fischer-Piette, 1952" authorityName=": Fischer-Piette" authorityYear="1952" box="[219,555,509,530]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="milloti">
<emphasis id="B9617238E532D90C0696FE69474CB551" box="[219,365,511,530]" italics="true" pageId="27" pageNumber="27">Ampelita milloti</emphasis>
Fischer-Piette, 1952
</taxonomicName>
,
<taxonomicName id="4C15D5A9E532D90C0479FE6B449CB551" box="[564,701,509,530]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="27" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, st’n 16/06, max. diameter 24.8 mm, NMSA L7127.
</paragraph>
</caption>
<paragraph id="8BAAAE2AE532D90C06DDFDF64531B5F9" blockId="27.[144,1104,608,1258]" pageId="27" pageNumber="27">
Distribution: Evidently a narrowly endemic species, currently known only from the south bank of Tsiribihina River, Tsingy de Bemaraha and
<taxonomicName id="4C15D5A9E532D90C055AFD1645E3B5D9" box="[791,962,640,666]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="27" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
. Subfossil at Antsingimavo where it now appears to be locally extinct.
</paragraph>
<paragraph id="8BAAAE2AE532D90C06DDFD5E4772B460" blockId="27.[144,1104,608,1258]" pageId="27" pageNumber="27">
Habitat: Living specimens are most commonly found amongst limestone rocks and in leaf-litter. In
<taxonomicName id="4C15D5A9E532D90C076BFD7E47F3B441" box="[294,466,744,770]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="27" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
, it occurs predominantly in the taller, moister forests of the southern part.
</paragraph>
<paragraph id="8BAAAE2AE532D90C06DDFCA644D6B488" blockId="27.[144,1104,608,1258]" pageId="27" pageNumber="27">
Remarks: Shell lenticular, of moderate size (max. diameter &lt;
<quantity id="4CED03CFE532D90C0577FCA645AFB408" box="[826,910,816,843]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="27" unit="mm" value="33.0">33 mm</quantity>
), with a very low spire and sharply angled periphery; last whorl descending at most slightly just prior to aperture; uniformly tan-brown with a yellow to white band around umbilicus. Specimens from
<taxonomicName id="4C15D5A9E532D90C0683FC064757B4E9" box="[206,374,912,938]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="27" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
are noticeably smaller than those from Bemaraha and Antsingimavo (maximum adult diameter 20–25
<emphasis id="B9617238E532D90C0400FC254445B488" box="[589,612,947,971]" italics="true" pageId="27" pageNumber="27">vs</emphasis>
<quantity id="4CED03CFE532D90C0421FC2644CAB488" box="[620,747,944,971]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="27" pageNumber="27" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">28–32 mm</quantity>
).
</paragraph>
<paragraph id="8BAAAE2AE532D90C06E7FC46476DB3A9" blockId="27.[144,1104,608,1258]" pageId="27" pageNumber="27">
Resembles
<taxonomicName id="4C15D5A9E532D90C0761FC4547D5B4A8" authorityName=": Fischer-Piette" authorityYear="1952" box="[300,500,979,1003]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="namerokoensis">
<emphasis id="B9617238E532D90C0761FC4547D5B4A8" box="[300,500,979,1003]" italics="true" pageId="27" pageNumber="27">A. namerokoensis</emphasis>
</taxonomicName>
(below) in size and shape, but easily separated from that species due to the almost complete absence of microscopic spiral striae (some traces may be visible on the base). Instead the microsculpture of
<taxonomicName id="4C15D5A9E532D90C0520FB8545F7B368" authorityName=": Fischer-Piette" authorityYear="1952" box="[877,982,1043,1067]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="milloti">
<emphasis id="B9617238E532D90C0520FB8545F7B368" box="[877,982,1043,1067]" italics="true" pageId="27" pageNumber="27">A. milloti</emphasis>
</taxonomicName>
comprises irregular, elongate granules aligned in a roughly radial (collabral) pattern (
<figureCitation id="132EB2AFE532D90C0597FBA64262B309" box="[986,1091,1072,1098]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="27" pageNumber="27">Fig. 17D</figureCitation>
). Furthermore,
<taxonomicName id="4C15D5A9E532D90C0765FBC547AFB328" authorityName=": Fischer-Piette" authorityYear="1952" box="[296,398,1107,1131]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="milloti">
<emphasis id="B9617238E532D90C0765FBC547AFB328" box="[296,398,1107,1131]" italics="true" pageId="27" pageNumber="27">A. milloti</emphasis>
</taxonomicName>
attains a somewhat larger size and has a wider umbilicus than
<taxonomicName id="4C15D5A9E532D90C0275FBC54716B3C8" authorityName=": Fischer-Piette" authorityYear="1952" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="namerokoensis">
<emphasis id="B9617238E532D90C0275FBC54716B3C8" italics="true" pageId="27" pageNumber="27">A. namerokoensis</emphasis>
</taxonomicName>
, and its protoconch is larger (diameter approx. 4.4
<emphasis id="B9617238E532D90C053BFBE545ACB3C8" box="[886,909,1139,1163]" italics="true" pageId="27" pageNumber="27">vs</emphasis>
approx.
<quantity id="4CED03CFE532D90C05BDFBE64269B3C8" box="[1008,1096,1136,1163]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="27" pageNumber="27" unit="mm" value="3.5">3.5 mm</quantity>
) and bears distinct close-set axial riblets (
<figureCitation id="132EB2AFE532D90C0417FB0644E1B3E9" box="[602,704,1168,1194]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="27" pageNumber="27">Fig. 17C</figureCitation>
), whereas that of
<taxonomicName id="4C15D5A9E532D90C05C4FB054271B3E8" authorityName=": Fischer-Piette" authorityYear="1952" box="[905,1104,1171,1195]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="namerokoensis">
<emphasis id="B9617238E532D90C05C4FB054271B3E8" box="[905,1104,1171,1195]" italics="true" pageId="27" pageNumber="27">A. namerokoensis</emphasis>
</taxonomicName>
is more or less smooth (
<figureCitation id="132EB2AFE532D90C07EEFB26442BB389" box="[419,522,1200,1226]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="27" pageNumber="27">Fig. 17E</figureCitation>
). Features separating
<taxonomicName id="4C15D5A9E532D90C054FFB25454DB388" authorityName=": Fischer-Piette" authorityYear="1952" box="[770,876,1203,1227]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="milloti">
<emphasis id="B9617238E532D90C054FFB25454DB388" box="[770,876,1203,1227]" italics="true" pageId="27" pageNumber="27">A. milloti</emphasis>
</taxonomicName>
from
<taxonomicName id="4C15D5A9E532D90C05FEFB254204B388" box="[947,1061,1203,1227]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="27" pageNumber="27" phylum="Mollusca" rank="species" species="beanka">
<emphasis id="B9617238E532D90C05FEFB254204B388" box="[947,1061,1203,1227]" italics="true" pageId="27" pageNumber="27">A. beanka</emphasis>
</taxonomicName>
are discussed above.
</paragraph>
</subSubSection>
</treatment>
</document>
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