286 lines
36 KiB
XML
286 lines
36 KiB
XML
<document id="87087E1C3FFFC9DB3661930A7CE99D2A" ID-DOI="10.1111/j.1096-3642.2008.00436.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445858" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630459740808" checkinUser="felipe" docAuthor="Skinner, Adam" docDate="2009" docId="03B587C2FF87FFACD86BBFE695A6F8A3" docLanguage="en" docName="j.1096-3642.2008.00436.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00436.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Pseudonaja aspidorhyncha" docType="treatment" docVersion="4" lastPageNumber="189" masterDocId="FF8CFFBAFF97FFBEDB3BBA22946BFFCD" masterDocTitle="A multivariate morphometric analysis and systematic review of Pseudonaja (Serpentes, Elapidae, Hydrophiinae)" masterLastPageNumber="197" masterPageNumber="171" pageNumber="187" updateTime="1699278810533" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="59F26BC35BED4D8D112AAEFEADB59B06">A multivariate morphometric analysis and systematic review of Pseudonaja (Serpentes, Elapidae, Hydrophiinae)</mods:title>
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<mods:namePart id="4FE0A65DD23584DD0441C7FC3526FC38">Skinner, Adam</mods:namePart>
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<mods:title id="0550F94550F8A652F28D356D152AD346">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="B812FB2B54A0D430616A7C1BA07280E9">
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<mods:date id="EE4CD7BA79F941F663A00F312938981F">2009</mods:date>
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<mods:detail id="DDC19D35E6EBF60105FF22E24C3C5598" type="pubDate">
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<mods:number id="E5342419A58E6618BA81E61FB9987E01">2009-01-31</mods:number>
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<mods:detail id="76ADC3B9926B05041BF79FB45F02984A" type="volume">
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<mods:number id="46030B20221B45BFE7F68CA4627F568B">155</mods:number>
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<mods:identifier id="FC59466A43A7523E03B0359B6160CF9F" type="DOI">10.1111/j.1096-3642.2008.00436.x</mods:identifier>
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<mods:identifier id="85398B2995086ED311E54D4CEF13FEEB" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03B587C2FF87FFACD86BBFE695A6F8A3" ID-DOI="http://doi.org/10.5281/zenodo.5492447" ID-Zenodo-Dep="5492447" LSID="urn:lsid:plazi:treatment:03B587C2FF87FFACD86BBFE695A6F8A3" httpUri="http://treatment.plazi.org/id/03B587C2FF87FFACD86BBFE695A6F8A3" lastPageId="18" lastPageNumber="189" pageId="16" pageNumber="187">
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<subSubSection id="C306655FFF87FFAED86BBFE69108FA14" box="[848,1379,1475,1497]" pageId="16" pageNumber="187" type="nomenclature">
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<paragraph id="8BA336D4FF87FFAED86BBFE69108FA14" blockId="16.[848,1379,1475,1497]" box="[848,1379,1475,1497]" pageId="16" pageNumber="187">
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<heading id="D0EB81B8FF87FFAED86BBFE69108FA14" box="[848,1379,1475,1497]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="187" reason="2">
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<taxonomicName id="4C1C4D57FF87FFAED86BBFE69108FA14" authority="(MCCOY, 1879)" baseAuthorityName="MCCOY" baseAuthorityYear="1879" box="[848,1379,1475,1497]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="187" phylum="Chordata" rank="species" species="aspidorhyncha">
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<emphasis id="B968EAC6FF87FFAED86BBFE690C2FA1A" box="[848,1193,1476,1497]" italics="true" pageId="16" pageNumber="187">PSEUDONAJA ASPIDORHYNCHA</emphasis>
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(MCCOY, 1879)
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C306655FFF87FFAFD81DBFC89580FB9E" lastPageId="17" lastPageNumber="188" pageId="16" pageNumber="187" type="reference_group">
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<paragraph id="8BA336D4FF87FFAED81DBFC8978CF991" blockId="16.[806,1421,1514,1903]" pageId="16" pageNumber="187">
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<taxonomicName id="4C1C4D57FF87FFAED81DBFC8917AFA32" authority="McCoy, 1879: 13" authorityName="McCoy" authorityPageNumber="13" authorityYear="1879" box="[806,1297,1514,1535]" class="Reptilia" family="Elapidae" genus="Diemenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="187" phylum="Chordata" rank="species" species="aspidorhyncha">
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<emphasis id="B968EAC6FF87FFAED81DBFC8902EFA32" box="[806,1093,1514,1535]" italics="true" pageId="16" pageNumber="187">Diemenia aspidorhyncha</emphasis>
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<materialsCitation id="3B743C89FF87FFAEDF6BBFC8917AFA32" accessionNumber="D12352" box="[1104,1297,1514,1535]" collectionCode="NMV" location="Darling River" municipality="Murray River" pageId="16" pageNumber="187" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">McCoy, 1879: 13</materialsCitation>
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</taxonomicName>
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||
.
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||
<typeStatus id="54A78876FF87FFAEDE1BBFC891E2F9CD" box="[1312,1417,1514,1536]" pageId="16" pageNumber="187" type="holotype">Holotype</typeStatus>
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||
, NMV D12352; type locality, restricted to junction of
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<collectingMunicipality id="6BC7ACAEFF87FFAED876BC059798F9F0" box="[845,1011,1575,1597]" pageId="16" pageNumber="187">Murray River</collectingMunicipality>
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and
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<location id="8EC3600FFF87FFAEDF79BC059081F9F0" LSID="urn:lsid:plazi:treatment:03B587C2FF87FFACD86BBFE695A6F8A3:8EC3600FFF87FFAEDF79BC059081F9F0" box="[1090,1258,1575,1597]" municipality="Murray River" name="Darling River" pageId="16" pageNumber="187" stateProvince="Victoria">Darling River</location>
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,
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<collectingRegion id="49D8F836FF87FFAEDE3BBC059137F9F0" box="[1280,1372,1575,1597]" country="Australia" name="Victoria" pageId="16" pageNumber="187">Victoria</collectingRegion>
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, by
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<bibRefCitation id="EF8D4B25FF87FFAED81DBC649788F991" author="Coventry AJ" box="[806,995,1606,1628]" pageId="16" pageNumber="187" pagination="115 - 124" refId="ref15180" refString="Coventry AJ. 1970. Reptile and amphibian type specimens housed in the National Museum of Victoria. Memoirs of the National Museum of Victoria 31: 115 - 124." type="journal article" year="1970">Coventry (1970)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BA336D4FF87FFAED81DBC479060F97A" blockId="16.[806,1421,1514,1903]" pageId="16" pageNumber="187">
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<taxonomicName id="4C1C4D57FF87FFAED81DBC47908AF9B4" authority="Longman, 1915: 31" authorityName="Longman" authorityPageNumber="31" authorityYear="1915" box="[806,1249,1636,1658]" class="Reptilia" family="Elapidae" genus="Diemenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="187" phylum="Chordata" rank="species" species="carinata">
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<emphasis id="B968EAC6FF87FFAED81DBC479790F9B7" box="[806,1019,1637,1658]" italics="true" pageId="16" pageNumber="187">Diemenia carinata</emphasis>
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<materialsCitation id="3B743C89FF87FFAEDF38BC46908AF9B4" box="[1027,1249,1636,1658]" collectionCode="QM" location="Charleville" municipality="Cane Grass Station" pageId="16" pageNumber="187" specimenCode="J1508" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">Longman, 1915: 31</materialsCitation>
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</taxonomicName>
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||
.
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||
<typeStatus id="54A78876FF87FFAEDFD5BC46913CF9B7" box="[1262,1367,1636,1658]" pageId="16" pageNumber="187" type="holotype">Holotype</typeStatus>
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, QM
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<specimenCode id="DBBA9EAFFF87FFAED81DBCA09706F955" box="[806,877,1666,1688]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="16" pageNumber="187" type="Museum">J1508</specimenCode>
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; type locality,
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<collectingMunicipality id="6BC7ACAEFF87FFAEDF24BCA19167F954" box="[1055,1292,1667,1689]" pageId="16" pageNumber="187">Cane Grass Station</collectingMunicipality>
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, via
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<location id="8EC3600FFF87FFAEDE77BCA19705F97A" LSID="urn:lsid:plazi:treatment:03B587C2FF87FFACD86BBFE695A6F8A3:8EC3600FFF87FFAEDE77BCA19705F97A" municipality="Cane Grass Station" name="Charleville" pageId="16" pageNumber="187" stateProvince="Queensland">Charleville</location>
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,
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<collectingRegion id="49D8F836FF87FFAED847BC83906CF97A" box="[892,1031,1697,1719]" country="Australia" name="Queensland" pageId="16" pageNumber="187">Queensland</collectingRegion>
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.
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</paragraph>
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<paragraph id="8BA336D4FF87FFAED81DBCE297F2F8DE" blockId="16.[806,1421,1514,1903]" pageId="16" pageNumber="187">
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<taxonomicName id="4C1C4D57FF87FFAED81DBCE29153F91B" authority="Mitchell, 1951: 547" authorityName="Mitchell" authorityPageNumber="547" authorityYear="1951" box="[806,1336,1728,1750]" class="Reptilia" family="Elapidae" genus="Demansia" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="187" phylum="Chordata" rank="species" species="acutirostris">
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<emphasis id="B968EAC6FF87FFAED81DBCE29045F918" box="[806,1070,1728,1749]" italics="true" pageId="16" pageNumber="187">Demansia acutirostris</emphasis>
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<materialsCitation id="3B743C89FF87FFAEDF79BCE29153F91B" box="[1090,1336,1728,1750]" collectionCode="SAMA" location="Lake Eyre" pageId="16" pageNumber="187" specimenCode="R3133" specimenCount="1" stateProvince="South Australia" typeStatus="holotype">Mitchell, 1951: 547</materialsCitation>
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</taxonomicName>
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.
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<typeStatus id="54A78876FF87FFAEDE6BBCE29731F939" pageId="16" pageNumber="187" type="holotype">Holotype</typeStatus>
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, SAMA
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<specimenCode id="DBBA9EAFFF87FFAED884BCFD9061F939" box="[959,1034,1759,1780]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="187">R3133</specimenCode>
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; type locality,
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<location id="8EC3600FFF87FFAEDF80BCFC9152F939" LSID="urn:lsid:plazi:treatment:03B587C2FF87FFACD86BBFE695A6F8A3:8EC3600FFF87FFAEDF80BCFC9152F939" box="[1211,1337,1758,1780]" name="Lake Eyre" pageId="16" pageNumber="187" stateProvince="South Australia">Lake Eyre</location>
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,
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<collectingRegion id="49D8F836FF87FFAEDE71BCFC97FEF8DE" country="Australia" name="South Australia" pageId="16" pageNumber="187">South Australia</collectingRegion>
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||
.
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</paragraph>
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||
<paragraph id="8BA336D4FF87FFAFD81DBD3E9580FB9E" blockId="16.[806,1421,1514,1903]" lastBlockId="17.[164,779,1055,1107]" lastPageId="17" lastPageNumber="188" pageId="16" pageNumber="187">
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||
<treatmentCitationGroup id="AB0C11FAFF87FFAFD81DBD3E9580FB9E" lastPageId="17" lastPageNumber="188" pageId="16" pageNumber="187">
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||
<treatmentCitation id="0ABD10C5FF87FFAED81DBD3E90FBF8FF" author="Wells RW" box="[806,1168,1820,1842]" page="6" pageId="16" pageNumber="187" year="2002">
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||
<taxonomicName id="4C1C4D57FF87FFAED81DBD3E90FBF8FF" authority="Wells, 2002: 6" authorityName="Wells" authorityPageNumber="6" authorityYear="2002" box="[806,1168,1820,1842]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="187" phylum="Chordata" rank="species" species="gowi">
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<emphasis id="B968EAC6FF87FFAED81DBD3E978CF8FC" box="[806,999,1820,1841]" italics="true" pageId="16" pageNumber="187">Pseudonaja gowi</emphasis>
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<bibRefCitation id="EF8D4B25FF87FFAED8D5BD3E90FBF8FF" author="Wells RW" box="[1006,1168,1820,1842]" pageId="16" pageNumber="187" pagination="1 - 48" refId="ref16337" refString="Wells RW. 2002. Taxonomy of the genus Pseudonaja (Reptilia: Elapidae) in Australia. Australian Biodiversity Record 2002: 1 - 48." type="journal article" year="2002">Wells, 2002: 6</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
<typeStatus id="54A78876FF87FFAEDFA6BD3E916DF8FF" box="[1181,1286,1820,1842]" pageId="16" pageNumber="187" type="holotype">Holotype</typeStatus>
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||
, designated as ‘the largest specimen of this species from the vicinity of Lyndhurst, SA in the South Australian Museum collection’ (
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<treatmentCitation id="0ABD10C5FF86FFAFDAA5BE3D9638FBF8" author="Wells RW" box="[414,595,1055,1077]" page="6" pageId="17" pageNumber="188" year="2002">
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||
<bibRefCitation id="EF8D4B25FF86FFAFDAA5BE3D9638FBF8" author="Wells RW" box="[414,595,1055,1077]" pageId="17" pageNumber="188" pagination="1 - 48" refId="ref16337" refString="Wells RW. 2002. Taxonomy of the genus Pseudonaja (Reptilia: Elapidae) in Australia. Australian Biodiversity Record 2002: 1 - 48." type="journal article" year="2002">Wells, 2002: 6</bibRefCitation>
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||
</treatmentCitation>
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||
); type locality, Lyndhurst,
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<collectingRegion id="49D8F836FF86FFAFDA17BE1F958CFB9E" box="[300,487,1085,1107]" country="Australia" name="South Australia" pageId="17" pageNumber="188">South Australia</collectingRegion>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF63665CFF86FFAFDB9FB9B89079FC25" ID-DOI="http://doi.org/10.5281/zenodo.5445882" ID-Zenodo-Dep="5445882" httpUri="https://zenodo.org/record/5445882/files/figure.png" pageId="17" pageNumber="188" startId="17.[164,243,922,941]" targetBox="[164,1444,196,892]" targetPageId="17">
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||
<paragraph id="8BA336D4FF86FFAFDB9FB9B89079FC25" blockId="17.[164,1441,922,1000]" pageId="17" pageNumber="188">
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||
<emphasis id="B968EAC6FF86FFAFDB9FB9B89572FC60" bold="true" box="[164,281,922,941]" pageId="17" pageNumber="188">Figure 10.</emphasis>
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Geographical distribution of
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<taxonomicName id="4C1C4D57FF86FFAFD959B9B8974CFC60" authorityName=", Gunther" authorityYear="1872" box="[610,807,922,941]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="affinis">
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||
<emphasis id="B968EAC6FF86FFAFD959B9B8974CFC60" box="[610,807,922,941]" italics="true" pageId="17" pageNumber="188">Pseudonaja affinis</emphasis>
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||
</taxonomicName>
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||
specimens examined in this study (closed circles) and/or included in Skinner
|
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<emphasis id="B968EAC6FF86FFAFDABAB99A95DEFC07" box="[385,437,951,970]" italics="true" pageId="17" pageNumber="188">et al.</emphasis>
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||
’s (2005) phylogenetic analysis (open circles represent specimens not included in the morphometric analyses). Codes for geographical groups correspond with those in Table 2.
|
||
</paragraph>
|
||
</caption>
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||
<subSubSection id="C306655FFF86FFACDB9FBE5995A6F8A3" lastPageId="18" lastPageNumber="189" pageId="17" pageNumber="188" type="diagnosis">
|
||
<paragraph id="8BA336D4FF86FFAFDB9FBE599678F918" blockId="17.[164,780,1146,1750]" pageId="17" pageNumber="188">
|
||
<emphasis id="B968EAC6FF86FFAFDB9FBE599637FB42" box="[164,604,1146,1168]" italics="true" pageId="17" pageNumber="188">
|
||
Diagnosis:
|
||
<taxonomicName id="4C1C4D57FF86FFAFDA1EBE589637FB42" baseAuthorityName="MCCOY" baseAuthorityYear="1879" box="[293,604,1146,1167]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="aspidorhyncha">Pseudonaja aspidorhyncha</taxonomicName>
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||
</emphasis>
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||
may be differentiated from the remaining species of
|
||
<taxonomicName id="4C1C4D57FF86FFAFD9BCBEBB9760FB63" authorityName="Gunther" authorityYear="1858" box="[647,779,1177,1198]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="genus">
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||
<emphasis id="B968EAC6FF86FFAFD9BCBEBB9760FB63" box="[647,779,1177,1198]" italics="true" pageId="17" pageNumber="188">Pseudonaja</emphasis>
|
||
</taxonomicName>
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||
redescribed here except
|
||
<taxonomicName id="4C1C4D57FF86FFAFDAFCBE9A9623FB00" box="[455,584,1208,1229]" class="Reptilia" family="Elapidae" genus="Demansia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="nuchalis">
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||
<emphasis id="B968EAC6FF86FFAFDAFCBE9A9623FB00" box="[455,584,1208,1229]" italics="true" pageId="17" pageNumber="188">P. nuchalis</emphasis>
|
||
</taxonomicName>
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||
in possessing a strap-like rostral that imparts a chisel shape to the snout from a dorsal perspective (see
|
||
<figureCitation id="13272A51FF86FFAFD976BED796E6FAC6" box="[589,653,1269,1291]" captionStart="Figure 1" captionStartId="2.[144,223,954,973]" captionTargetBox="[144,1424,199,924]" captionTargetId="figure-75@2.[144,1424,196,924]" captionTargetPageId="2" captionText="Figure 1. Pseudonaja aspidorhyncha (SAMA R18600), dorsal perspective of head (left); P. mengdeni (SAMA R20981), dorsal perspective of head (right); see Table 1 for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5445860" httpUri="https://zenodo.org/record/5445860/files/figure.png" pageId="17" pageNumber="188">Fig. 1</figureCitation>
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||
). A diploid chromosome number of 34 serves to distinguish
|
||
<taxonomicName id="4C1C4D57FF86FFAFD9CCBF369527FA8A" baseAuthorityName="MCCOY" baseAuthorityYear="1879" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="aspidorhyncha">
|
||
<emphasis id="B968EAC6FF86FFAFD9CCBF369527FA8A" italics="true" pageId="17" pageNumber="188">P. aspidorhyncha</emphasis>
|
||
</taxonomicName>
|
||
from the remaining species of
|
||
<taxonomicName id="4C1C4D57FF86FFAFD985BF109488FAAB" authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="genus">
|
||
<emphasis id="B968EAC6FF86FFAFD985BF109488FAAB" italics="true" pageId="17" pageNumber="188">Pseudonaja</emphasis>
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C1C4D57FF86FFAFDA7BBF7395CFFAAB" authorityName=", Gunther" authorityYear="1872" box="[320,420,1361,1382]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="affinis">
|
||
<emphasis id="B968EAC6FF86FFAFDA7BBF7395CFFAAB" box="[320,420,1361,1382]" italics="true" pageId="17" pageNumber="188">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
and possibly
|
||
<taxonomicName id="4C1C4D57FF86FFAFD973BF7396BFFAAB" authorityName="Wells & Wellington" authorityYear="1985" box="[584,724,1361,1382]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="mengdeni">
|
||
<emphasis id="B968EAC6FF86FFAFD973BF7396BFFAAB" box="[584,724,1361,1382]" italics="true" pageId="17" pageNumber="188">P. mengdeni</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF8D4B25FF86FFAFDB9FBF4D95E0FA48" author="Skinner A & Donnellan SC & Hutchinson MN & Hutchinson RG" box="[164,395,1391,1413]" pageId="17" pageNumber="188" pagination="558 - 571" refId="ref16126" refString="Skinner A, Donnellan SC, Hutchinson MN, Hutchinson RG. 2005. A phylogenetic analysis of Pseudonaja (Hydrophiinae, Elapidae, Serpentes) based on mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 37: 558 - 571." type="journal article" year="2005">
|
||
Skinner
|
||
<emphasis id="B968EAC6FF86FFAFDA32BF52952AFA49" box="[265,321,1391,1413]" italics="true" pageId="17" pageNumber="188">et al.</emphasis>
|
||
, 2005
|
||
</bibRefCitation>
|
||
, and below). The relative length of the prefrontal suture (i.e. the length of the prefrontal suture divided by head length) is consistently lower than for
|
||
<taxonomicName id="4C1C4D57FF86FFAFDA63BFEE95AFFA2D" baseAuthorityName="DUMERIL, BIBRON & DUMERIL" baseAuthorityYear="1854" box="[344,452,1483,1505]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="textilis">
|
||
<emphasis id="B968EAC6FF86FFAFDA63BFEE95AFFA2D" box="[344,452,1483,1505]" italics="true" pageId="17" pageNumber="188">P. textilis</emphasis>
|
||
</taxonomicName>
|
||
specimens (
|
||
<geoCoordinate id="EE285013FF86FFAFD96DBFE996FFFA2D" box="[598,660,1483,1504]" degrees="0.049" direction="north" orientation="latitude" pageId="17" pageNumber="188" precision="55" value="0.049">0.049</geoCoordinate>
|
||
<geoCoordinate id="EE285013FF86FFAFD9AFBFE9968BFA2D" box="[660,736,1483,1504]" degrees="0.092" direction="west" orientation="longitude" pageId="17" pageNumber="188" precision="55" value="-0.092">–0.092</geoCoordinate>
|
||
vs.
|
||
<geoCoordinate id="EE285013FF86FFAFDB9FBFC89488F9CD" box="[164,227,1514,1536]" degrees="0.096" direction="north" orientation="latitude" pageId="17" pageNumber="188" precision="55" value="0.096">0.096</geoCoordinate>
|
||
<geoCoordinate id="EE285013FF86FFAFDBD8BFC8955BF9CD" box="[227,304,1514,1536]" degrees="0.136" direction="west" orientation="longitude" pageId="17" pageNumber="188" precision="55" value="-0.136">–0.136</geoCoordinate>
|
||
; see
|
||
<tableCitation id="C69E036FFF86FFAFDA51BFC895D5F9CD" box="[362,446,1514,1536]" captionStart="Table 9" captionStartId="13.[164,228,200,219]" captionText="Table 9. Descriptive statistics for morphometric variables; see Table 1 for abbreviations" pageId="17" pageNumber="188">Table 9</tableCitation>
|
||
).
|
||
<taxonomicName id="4C1C4D57FF86FFAFDAEEBFC89761FA32" baseAuthorityName="MCCOY" baseAuthorityYear="1879" box="[469,778,1514,1535]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="aspidorhyncha">
|
||
<emphasis id="B968EAC6FF86FFAFDAEEBFC89761FA32" box="[469,778,1514,1535]" italics="true" pageId="17" pageNumber="188">Pseudonaja aspidorhyncha</emphasis>
|
||
</taxonomicName>
|
||
is separable from
|
||
<taxonomicName id="4C1C4D57FF86FFAFDABCBC2B9673F9D0" authorityName="Wells & Wellington" authorityYear="1985" box="[391,536,1544,1566]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="mengdeni">
|
||
<emphasis id="B968EAC6FF86FFAFDABCBC2B9673F9D0" box="[391,536,1544,1566]" italics="true" pageId="17" pageNumber="188">P. mengdeni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D57FF86FFAFD95ABC2B968FF9D0" box="[609,740,1544,1566]" class="Reptilia" family="Elapidae" genus="Demansia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="nuchalis">
|
||
<emphasis id="B968EAC6FF86FFAFD95ABC2B968FF9D0" box="[609,740,1544,1566]" italics="true" pageId="17" pageNumber="188">P. nuchalis</emphasis>
|
||
</taxonomicName>
|
||
in displaying (when preserved) a whitish to pale grey (as opposed to predominantly dark bluish grey) buccal epithelium (in live specimens, the buccal epithelium is never black as in
|
||
<taxonomicName id="4C1C4D57FF86FFAFDAB2BCA19679F955" authorityName="Wells & Wellington" authorityYear="1985" box="[393,530,1667,1688]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="mengdeni">
|
||
<emphasis id="B968EAC6FF86FFAFDAB2BCA19679F955" box="[393,530,1667,1688]" italics="true" pageId="17" pageNumber="188">P. mengdeni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D57FF86FFAFD977BCA196A2F955" box="[588,713,1667,1688]" class="Reptilia" family="Elapidae" genus="Demansia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="nuchalis">
|
||
<emphasis id="B968EAC6FF86FFAFD977BCA196A2F955" box="[588,713,1667,1688]" italics="true" pageId="17" pageNumber="188">P. nuchalis</emphasis>
|
||
</taxonomicName>
|
||
), and usually exhibits a greater number of ventrals than
|
||
<taxonomicName id="4C1C4D57FF86FFAFDB9FBCE29524F918" baseAuthorityName="WAITE" baseAuthorityYear="1925" box="[164,335,1728,1749]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="inframacula">
|
||
<emphasis id="B968EAC6FF86FFAFDB9FBCE29524F918" box="[164,335,1728,1749]" italics="true" pageId="17" pageNumber="188">P. inframacula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1C4D57FF86FFAFDAB6BCE29667F918" box="[397,524,1728,1749]" class="Reptilia" family="Elapidae" genus="Demansia" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="188" phylum="Chordata" rank="species" species="nuchalis">
|
||
<emphasis id="B968EAC6FF86FFAFDAB6BCE29667F918" box="[397,524,1728,1749]" italics="true" pageId="17" pageNumber="188">P. nuchalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA336D4FF86FFAFDB9FBCDC9099FA48" blockId="17.[164,779,1789,1903]" lastBlockId="17.[826,1441,1055,1903]" pageId="17" pageNumber="188">
|
||
<emphasis id="B968EAC6FF86FFAFDB9FBCDC9547F8DE" box="[164,300,1790,1811]" italics="true" pageId="17" pageNumber="188">Description:</emphasis>
|
||
Ventrals 207–226; subcaudals 47–63; rostral strap like, snout appearing chisel shaped from dorsal perspective; nasal undivided, contacting preocular; two postoculars; parietal contacting (64 of 72 cases) or separated from (eight cases) lower postocular; six (70 of 72 cases) or five (two cases) supralabials; seven (71 of 72 cases) or eight (one case) infralabials; temporals 1 + 2 + 3 (61 of 71 cases), 1 + 2 + 4 (eight cases), 1 + 3 + 3 (one case) or 2 + 2 + 4 (one case); 5–8 nuchals contacting parietals; dorsals in 21–27 rows at first ventral, 17–20 rows one head length posterior to occiput, 17 rows midbody, 13–15 rows one head length anterior to vent, 14–18 rows at last ventral; anal divided (35 of 36 cases) or undivided (one case); snout– vent length
|
||
<quantity id="4CE49B31FF86FFAFD8E8BF73901EFAAB" box="[979,1141,1361,1383]" metricMagnitude="-1" metricUnit="m" metricValue="8.084999999999999" metricValueMax="12.76" metricValueMin="3.41" pageId="17" pageNumber="188" unit="mm" value="808.5" valueMax="1276.0" valueMin="341.0">341–1276 mm</quantity>
|
||
; tail length
|
||
<quantity id="4CE49B31FF86FFAFDE2EBF7391F7FAAB" box="[1301,1436,1361,1382]" metricMagnitude="-1" metricUnit="m" metricValue="1.24" metricValueMax="1.9899999999999998" metricValueMin="0.49" pageId="17" pageNumber="188" unit="mm" value="124.0" valueMax="199.0" valueMin="49.0">49–199 mm</quantity>
|
||
,
|
||
<geoCoordinate id="EE285013FF86FFAFD801BF4D97EFFA48" box="[826,900,1391,1413]" degrees="12.618" direction="north" orientation="latitude" pageId="17" pageNumber="188" precision="55" value="12.618">12.618</geoCoordinate>
|
||
<geoCoordinate id="EE285013FF86FFAFD8BEBF4D97B5FA48" box="[901,990,1391,1413]" degrees="18.862" direction="west" orientation="longitude" pageId="17" pageNumber="188" precision="55" value="-18.862">–18.862</geoCoordinate>
|
||
% of snout–vent length.
|
||
</paragraph>
|
||
<paragraph id="8BA336D4FF86FFACD868BFAC95D6FABF" blockId="17.[826,1441,1055,1903]" lastBlockId="18.[144,759,1311,1517]" lastPageId="18" lastPageNumber="189" pageId="17" pageNumber="188">Dorsum pale to medium brown; indistinct pattern of darker brown or greyish brown, oblique bands, approximately one dorsal in width, often evident (usually more pronounced laterally and posteriorly); some specimens exhibit a series of indistinct to distinct, broad, darker brown or black bands along body, occasionally with interposed narrower bands; not uncommonly, a broad, dark brown or black band is present on neck; body and neck commonly with scattered dark brown or black scales; frontal, supraoculars and subocular (i.e. 3rd and 4th) supralabials often darker brown; occasionally, entire head dark brown; venter dirty cream or yellow, or medium brown; ventrals commonly exhibit a darker brown sectorial marking laterally; posterior margins of ventrals of some specimens darker brown; chin typically cream or pale yellow; buccal epithelium whitish to pale grey (preserved specimens); iris dark with light orange ring around pupil.</paragraph>
|
||
<caption id="DF63665CFF85FFACDBABBE459026FB78" ID-DOI="http://doi.org/10.5281/zenodo.5445884" ID-Zenodo-Dep="5445884" httpUri="https://zenodo.org/record/5445884/files/figure.png" pageId="18" pageNumber="189" startId="18.[144,223,1127,1146]" targetBox="[144,1424,195,1097]" targetPageId="18">
|
||
<paragraph id="8BA336D4FF85FFACDBABBE459026FB78" blockId="18.[144,1421,1127,1205]" pageId="18" pageNumber="189">
|
||
<emphasis id="B968EAC6FF85FFACDBABBE45956FFBB7" bold="true" box="[144,260,1127,1146]" pageId="18" pageNumber="189">Figure 11.</emphasis>
|
||
Geographical distribution of
|
||
<taxonomicName id="4C1C4D57FF85FFACD971BE459703FBB7" baseAuthorityName="MCCOY" baseAuthorityYear="1879" box="[586,872,1127,1146]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="189" phylum="Chordata" rank="species" species="aspidorhyncha">
|
||
<emphasis id="B968EAC6FF85FFACD971BE459703FBB7" box="[586,872,1127,1146]" italics="true" pageId="18" pageNumber="189">Pseudonaja aspidorhyncha</emphasis>
|
||
</taxonomicName>
|
||
specimens examined in this study (closed circles) and/or included in Skinner
|
||
<emphasis id="B968EAC6FF85FFACDA84BEA79599FB5A" box="[447,498,1156,1175]" italics="true" pageId="18" pageNumber="189">et al.</emphasis>
|
||
’s (2005) phylogenetic analysis (open circles represent specimens not included in the morphometric analyses). Codes for geographical groups correspond with those in Table 2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA336D4FF85FFACDB93BF59958BFA20" blockId="18.[144,759,1311,1517]" pageId="18" pageNumber="189">
|
||
<bibRefCitation id="EF8D4B25FF85FFACDB93BF599501FA5C" author="Mengden GA" box="[168,362,1403,1425]" pageId="18" pageNumber="189" pagination="193 - 208" refId="ref15779" refString="Mengden GA. 1985. A chromosomal and electrophoretic analysis of the genus Pseudonaja. In: Grigg G, Shine R, Ehmann H, eds. Biology of Australasian frogs and reptiles. Sydney: Royal Zoological Society of New South Wales, 193 - 208." type="book chapter" year="1985">Mengden (1985)</bibRefCitation>
|
||
reported a diploid chromosome number of 34, with autosome pairs 4–16 being separable into two distinct size classes, and sex chromosomes that are of equal size.
|
||
</paragraph>
|
||
<paragraph id="8BA336D4FF85FFACDBABBC0F96EFF94D" blockId="18.[144,759,1581,1664]" pageId="18" pageNumber="189">
|
||
<emphasis id="B968EAC6FF85FFACDBABBC0F954AF98F" box="[144,289,1581,1602]" italics="true" pageId="18" pageNumber="189">Distribution:</emphasis>
|
||
Southern central
|
||
<collectingCountry id="F30B7644FF85FFACD935BC0F9617F98E" box="[526,636,1581,1603]" name="Australia" pageId="18" pageNumber="189">Australia</collectingCountry>
|
||
, south of
|
||
<geoCoordinate id="EE285013FF85FFACDBABBC6994ACF9AD" box="[144,199,1611,1632]" degrees="27" direction="south" orientation="latitude" pageId="18" pageNumber="189" precision="55555" value="-27.0">27°S</geoCoordinate>
|
||
, from Penong, western Eyre Peninsula, to Hermidale, central
|
||
<collectingRegion id="49D8F836FF85FFACDA73BC48967EF94D" box="[328,533,1642,1664]" country="Australia" name="New South Wales" pageId="18" pageNumber="189">New South Wales</collectingRegion>
|
||
(
|
||
<figureCitation id="13272A51FF85FFACD913BC48961CF94D" box="[552,631,1642,1664]" captionStart="Figure 11" captionStartId="18.[144,223,1127,1146]" captionTargetBox="[144,1424,195,1097]" captionTargetId="figure-250@18.[144,1424,195,1097]" captionTargetPageId="18" captionText="Figure 11. Geographical distribution of Pseudonaja aspidorhyncha specimens examined in this study (closed circles) and/or included in Skinner et al.’s (2005) phylogenetic analysis (open circles represent specimens not included in the morphometric analyses). Codes for geographical groups correspond with those in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5445884" httpUri="https://zenodo.org/record/5445884/files/figure.png" pageId="18" pageNumber="189">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA336D4FF85FFACDBABBCE295A6F8A3" blockId="18.[144,759,1728,1903]" pageId="18" pageNumber="189">
|
||
<emphasis id="B968EAC6FF85FFACDBABBCE294BEF918" box="[144,213,1728,1749]" italics="true" pageId="18" pageNumber="189">Notes:</emphasis>
|
||
<bibRefCitation id="EF8D4B25FF85FFACDBDBBCE295FBF91B" author="Wells RW" box="[224,400,1728,1750]" pageId="18" pageNumber="189" pagination="1 - 48" refId="ref16337" refString="Wells RW. 2002. Taxonomy of the genus Pseudonaja (Reptilia: Elapidae) in Australia. Australian Biodiversity Record 2002: 1 - 48." type="journal article" year="2002">Wells (2002: 6)</bibRefCitation>
|
||
erected the name
|
||
<taxonomicName id="4C1C4D57FF85FFACD948BCE294AEF939" authorityName="Wells" authorityYear="2002" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="189" phylum="Chordata" rank="species" species="gowi">
|
||
<emphasis id="B968EAC6FF85FFACD948BCE294AEF939" italics="true" pageId="18" pageNumber="189">Pseudonaja gowi</emphasis>
|
||
</taxonomicName>
|
||
for
|
||
<bibRefCitation id="EF8D4B25FF85FFACDA31BCFC95B5F939" author="Mengden GA" box="[266,478,1758,1780]" pageId="18" pageNumber="189" pagination="193 - 208" refId="ref15779" refString="Mengden GA. 1985. A chromosomal and electrophoretic analysis of the genus Pseudonaja. In: Grigg G, Shine R, Ehmann H, eds. Biology of Australasian frogs and reptiles. Sydney: Royal Zoological Society of New South Wales, 193 - 208." type="book chapter" year="1985">Mengden’s (1985)</bibRefCitation>
|
||
‘Southern with black nuchal band’ morph. As discussed by
|
||
<bibRefCitation id="EF8D4B25FF85FFACD96CBCDF957EF8FC" author="Skinner A & Donnellan SC & Hutchinson MN & Hutchinson RG" pageId="18" pageNumber="189" pagination="558 - 571" refId="ref16126" refString="Skinner A, Donnellan SC, Hutchinson MN, Hutchinson RG. 2005. A phylogenetic analysis of Pseudonaja (Hydrophiinae, Elapidae, Serpentes) based on mitochondrial DNA sequences. Molecular Phylogenetics and Evolution 37: 558 - 571." type="journal article" year="2005">
|
||
Skinner
|
||
<emphasis id="B968EAC6FF85FFACD984BCDC969CF8DF" box="[703,759,1789,1811]" italics="true" pageId="18" pageNumber="189">et al.</emphasis>
|
||
(2005: 569)
|
||
</bibRefCitation>
|
||
, specimens of this morph are part of
|
||
<taxonomicName id="4C1C4D57FF85FFACD9D9BD3E949AF882" class="Reptilia" family="Elapidae" genus="Demansia" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="189" phylum="Chordata" rank="species" species="nuchalis">
|
||
<emphasis id="B968EAC6FF85FFACD9D9BD3E949AF882" italics="true" pageId="18" pageNumber="189">P. nuchalis</emphasis>
|
||
</taxonomicName>
|
||
‘Southern’, so that the name
|
||
<taxonomicName id="4C1C4D57FF85FFACD97EBD199611F89D" authorityName="Wells" authorityYear="2002" box="[581,634,1851,1872]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="189" phylum="Chordata" rank="species" species="gowi">
|
||
<emphasis id="B968EAC6FF85FFACD97EBD199611F89D" box="[581,634,1851,1872]" italics="true" pageId="18" pageNumber="189">gowi</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="4C1C4D57FF85FFACDA25BD7B95ADF8A3" baseAuthorityName="MCCOY" baseAuthorityYear="1879" box="[286,454,1881,1902]" class="Reptilia" family="Elapidae" genus="Pseudonaja" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="189" phylum="Chordata" rank="species" species="aspidorhyncha">
|
||
<emphasis id="B968EAC6FF85FFACDA25BD7B95ADF8A3" box="[286,454,1881,1902]" italics="true" pageId="18" pageNumber="189">aspidorhyncha</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |