243 lines
38 KiB
XML
243 lines
38 KiB
XML
<document id="511AE38FC3B3358A14173C9428C3EAF2" ID-CLB-Dataset="8450" ID-DOI="10.5281/zenodo.4504598" ID-GBIF-Dataset="5b2d651a-1c6b-445d-b1cc-62eb3ad7e85f" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504598" ID-ZooBank="932CC6CE-9BB3-40CD-B22F-691F70FE5475" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612466707376" checkinUser="carolina" docAuthor="Tumpeesuwan, Chanidaporn & Tumpeesuwan, Sakboworn" docDate="2018" docId="03B58780501B4B20FC1D0E81FD00FA62" docLanguage="en" docName="RafflesBZool.66.170-176.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Aenigmatoconcha sumonthai Tumpeesuwan & Tumpeesuwan 2018, new species" docType="treatment" docUuid="A222B4B6-65CA-49DE-B63F-FC2C57C6102E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="172" masterDocId="FF8CFFF8501A4B23FF8B0B3AFFB5FD05" masterDocTitle="Aenigmatoconcha sumonthai, a new helicarionid land snail from Chumphon Province, Southern Thailand (Helicarionidae: Durgellinae)" masterLastPageNumber="176" masterPageNumber="170" pageNumber="171" updateTime="1698927357985" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1ECC72D87ED3BF2E392277894380AC8B">Aenigmatoconcha sumonthai, a new helicarionid land snail from Chumphon Province, Southern Thailand (Helicarionidae: Durgellinae)</mods:title>
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<mods:namePart id="E646549520DFB136C582F7A50BD4D930">Tumpeesuwan, Chanidaporn</mods:namePart>
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<mods:namePart id="956D7E76F7DBCC02CED15825E5F2A299">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:title id="B2084DF29770885C2775553BF6E50003">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="0BAD6A22169418E16232620DBA98EB9E">2018</mods:date>
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<treatment id="03B58780501B4B20FC1D0E81FD00FA62" ID-DOI="http://doi.org/10.5281/zenodo.4504668" ID-GBIF-Taxon="177211889" ID-Zenodo-Dep="4504668" LSID="urn:lsid:zoobank.org:act:A222B4B6-65CA-49DE-B63F-FC2C57C6102E" httpUri="http://treatment.plazi.org/id/03B58780501B4B20FC1D0E81FD00FA62" lastPageId="3" lastPageNumber="172" pageId="1" pageNumber="171">
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<subSubSection id="C306651D501B4B22FC1D0E81FAC5F8D1" box="[918,1392,1467,1492]" pageId="1" pageNumber="171" type="nomenclature">
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<paragraph id="8BA33696501B4B22FC1D0E81FAC5F8D1" blockId="1.[918,1392,1467,1524]" box="[918,1392,1467,1492]" pageId="1" pageNumber="171">
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<heading id="D0EB81FA501B4B22FC1D0E81FAC5F8D1" bold="true" box="[918,1392,1467,1492]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="171" reason="2">
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<emphasis id="B968EA84501B4B22FC1D0E81FAC5F8D1" bold="true" box="[918,1392,1467,1492]" pageId="1" pageNumber="171">
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<taxonomicName id="4C1C4D15501B4B22FC1D0E81FB6EF8D6" ID-CoL="659WV" authority="Tumpeesuwan & Tumpeesuwan, 2018" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" box="[918,1243,1467,1491]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="171" phylum="Mollusca" rank="species" species="sumonthai" status="sp. nov.">
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<emphasis id="B968EA84501B4B22FC1D0E81FB6EF8D6" bold="true" box="[918,1243,1467,1491]" italics="true" pageId="1" pageNumber="171">Aenigmatoconcha sumonthai</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25B57FF501B4B22FB600E86FAC5F8D1" box="[1259,1392,1468,1492]" pageId="1" pageNumber="171" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306651D501B4B22FB940EE6FB53F8F1" box="[1055,1254,1500,1524]" pageId="1" pageNumber="171" type="description">
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<paragraph id="8BA33696501B4B22FB940EE6FB53F8F1" blockId="1.[918,1392,1467,1524]" box="[1055,1254,1500,1524]" pageId="1" pageNumber="171">
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<figureCitation id="13272A13501B4B22FB940EE6FB32F8F1" box="[1055,1159,1500,1524]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="2.[113,148,810,832]" captionStartId-1="2.[814,849,1433,1455]" captionStartId-2="3.[136,171,1544,1566]" captionStartId-3="4.[113,148,1273,1295]" captionStartId-4="4.[112,147,2064,2086]" captionTargetBox-0="[113,1451,188,794]" captionTargetBox-1="[814,1450,944,1417]" captionTargetBox-2="[136,1473,189,1529]" captionTargetBox-3="[113,753,420,1257]" captionTargetBox-4="[112,1451,1494,2049]" captionTargetId-0="figure-74@2.[113,1451,188,794]" captionTargetId-1="figure-24@2.[814,1450,944,1417]" captionTargetId-2="figure-8@3.[136,1473,189,1529]" captionTargetId-3="figure-70@4.[113,753,420,1257]" captionTargetId-4="figure-109@4.[112,1451,1494,2049]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionText-0="Fig. 2. Aenigmatoconcha sumonthai, new species, in natural habitat at the type locality; photographs taken on June 2017. A, two snails with the dorsal shell surface complately covered by the mantle shell lobes; B, two snails with mantle shell lobes retracted. (Photograph courtesy of Mr. Montri Sumontha)." captionText-1="Fig. 3. External shell morphology; Aenigmatoconcha sumonthai new species, holotype (NHMSU-0015)." captionText-2="Fig. 4. Aenigmatoconcha sumonthai new species, radula of paratype (NHMSU-0016). A, whole radula; B, close up view of central part of radula; C, close up view of central, 1st and 2nd lateral teeth; D, close up view of right side of central teeth; E, close up view of right latero-marginal teeth." captionText-3="Fig. 5. Aenigmatoconcha sumonthai, new species, genital system of paratype (NHMSU-0016). Abbreviations: ag = albumen gland, hd = hermaphroditic duct, ut = uterus, pro = prostate gland, vd = vas deferens, ep = epiphallus, erc = epiphallic retractor caecum, p = penis, gs = gametolytic sac, v = vagina, fo = free oviduct, at = atrium." captionText-4="Fig. 6. Living snails in natural habitat at type locality; photographs taken on June 2017. A, Aenigmatoconcha sumonthai, new species, on climbing plant near limestone wall; B, Sarika sp. on limestone (photographs courtesy of Mr. Montri Sumontha)." figureDoi-0="http://doi.org/10.5281/zenodo.4504604" figureDoi-1="http://doi.org/10.5281/zenodo.4504606" figureDoi-2="http://doi.org/10.5281/zenodo.4504608" figureDoi-3="http://doi.org/10.5281/zenodo.4504610" figureDoi-4="http://doi.org/10.5281/zenodo.4504612" httpUri-0="https://zenodo.org/record/4504604/files/figure.png" httpUri-1="https://zenodo.org/record/4504606/files/figure.png" httpUri-2="https://zenodo.org/record/4504608/files/figure.png" httpUri-3="https://zenodo.org/record/4504610/files/figure.png" httpUri-4="https://zenodo.org/record/4504612/files/figure.png" pageId="1" pageNumber="171">Figs. 2–6</figureCitation>
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;
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<tableCitation id="C69E032D501B4B22FB1F0EE6FB53F8F1" box="[1172,1254,1500,1524]" captionStart="Table 1" captionStartId="1.[136,191,189,211]" captionText="Table 1. Comparison of shells, radula and genital systems of Aenigmatoconcha clivicola and A. sumonthai, new species." pageId="1" pageNumber="171">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306651D501B4B22FCC80D26FA27FBF1" pageId="1" pageNumber="171" type="materials_examined">
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<paragraph id="8BA33696501B4B22FCC80D26FA27FBF1" blockId="1.[834,1475,1564,1780]" pageId="1" pageNumber="171">
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<emphasis id="B968EA84501B4B22FCC80D26FC46FB31" bold="true" box="[835,1011,1564,1588]" pageId="1" pageNumber="171">Type material.</emphasis>
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<materialsCitation id="3B743CCB501B4B22FC760D26FCCAFBB1" ID-GBIF-Occurrence="3029864302" collectingDate="2015-05" collectorName="Montri Sumontha" country="Thailand" location="Mueang district" pageId="1" pageNumber="171" specimenCount="1" stateProvince="Chumphon" typeStatus="holotype">
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<typeStatus id="54A78834501B4B22FC760D26FBD2FB31" box="[1021,1127,1564,1588]" pageId="1" pageNumber="171" type="holotype">Holotype</typeStatus>
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: NHMSU-0015 (
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<figureCitation id="13272A13501B4B22FAA10D26FAC4FB31" box="[1322,1393,1564,1588]" captionStart="Fig" captionStartId="2.[814,849,1433,1455]" captionTargetBox="[814,1450,944,1417]" captionTargetId="figure-24@2.[814,1450,944,1417]" captionTargetPageId="2" captionText="Fig. 3. External shell morphology; Aenigmatoconcha sumonthai new species, holotype (NHMSU-0015)." figureDoi="http://doi.org/10.5281/zenodo.4504606" httpUri="https://zenodo.org/record/4504606/files/figure.png" pageId="1" pageNumber="171">Fig. 3</figureCitation>
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). Shell height 8.0 mm, shell width
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<quantity id="4CE49B73501B4B22FBF30D06FB68FB56" box="[1144,1245,1596,1620]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="1" pageNumber="171" unit="mm" value="14.2">14.2 mm</quantity>
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; type locality: Tham Chang Phueak limestone range,
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<location id="8EC3604D501B4B22FB130D66FAF7FB71" LSID="urn:lsid:plazi:treatment:03B58780501B4B20FC1D0E81FD00FA62:8EC3604D501B4B22FB130D66FAF7FB71" box="[1176,1346,1628,1652]" country="Thailand" name="Mueang district" pageId="1" pageNumber="171" stateProvince="Chumphon">Mueang district</location>
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,
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<collectingRegion id="49D8F874501B4B22FAC70D66FC17FB91" country="Thailand" name="Chumphon" pageId="1" pageNumber="171">Chumphon province</collectingRegion>
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, southern
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<collectingCountry id="F30B7606501B4B22FB9C0D46FBCCFB91" box="[1047,1145,1660,1684]" name="Thailand" pageId="1" pageNumber="171">Thailand</collectingCountry>
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, coll.
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<collectorName id="26E95340501B4B22FB340D46FA37FB91" box="[1215,1410,1660,1684]" pageId="1" pageNumber="171">Montri Sumontha</collectorName>
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,
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<date id="FFA21056501B4B22FA1A0D46FCCAFBB1" pageId="1" pageNumber="171" value="2015-05">
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<collectingDate id="EFE6E9BE501B4B22FA1A0D46FCCAFBB1" pageId="1" pageNumber="171" value="2015-05">May 2015</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation id="3B743CCB501B4B22FC040DA6FA3AFBF1" ID-GBIF-Occurrence="3029864301" collectingDate="2015-05" collectionCode="ZRC" collectorName="Montri Sumontha" country="Thailand" location="Mueang district" pageId="1" pageNumber="171" specimenCount="1" stateProvince="Chumphon" typeStatus="paratype">
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<typeStatus id="54A78834501B4B22FC040DA6FBB6FBB1" box="[911,1027,1692,1716]" pageId="1" pageNumber="171" type="paratype">Paratypes</typeStatus>
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: NHMSU-0016 (empty shells, genital system and radula,
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<figureCitation id="13272A13501B4B22FB840D86FBEDFBD1" box="[1039,1112,1724,1748]" captionStart="Fig" captionStartId="3.[136,171,1544,1566]" captionTargetBox="[136,1473,189,1529]" captionTargetId="figure-8@3.[136,1473,189,1529]" captionTargetPageId="3" captionText="Fig. 4. Aenigmatoconcha sumonthai new species, radula of paratype (NHMSU-0016). A, whole radula; B, close up view of central part of radula; C, close up view of central, 1st and 2nd lateral teeth; D, close up view of right side of central teeth; E, close up view of right latero-marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.4504608" httpUri="https://zenodo.org/record/4504608/files/figure.png" pageId="1" pageNumber="171">Figs. 4</figureCitation>
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,
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<figureCitation id="13272A13501B4B22FBE80D86FBC7FBD1" box="[1123,1138,1724,1748]" captionStart="Fig" captionStartId="4.[113,148,1273,1295]" captionTargetBox="[113,753,420,1257]" captionTargetId="figure-70@4.[113,753,420,1257]" captionTargetPageId="4" captionText="Fig. 5. Aenigmatoconcha sumonthai, new species, genital system of paratype (NHMSU-0016). Abbreviations: ag = albumen gland, hd = hermaphroditic duct, ut = uterus, pro = prostate gland, vd = vas deferens, ep = epiphallus, erc = epiphallic retractor caecum, p = penis, gs = gametolytic sac, v = vagina, fo = free oviduct, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.4504610" httpUri="https://zenodo.org/record/4504610/files/figure.png" pageId="1" pageNumber="171">5</figureCitation>
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),
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<collectionCode id="ED0DAE53501B4B22FB0D0D86FB08FBD1" box="[1158,1213,1724,1748]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="171">ZRC</collectionCode>
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.MOL.11287 (one shell), THNHM-Iv-18004 (one shell), same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306651D501B4B22FCC80C26FBBAFA71" pageId="1" pageNumber="171" type="etymology">
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<paragraph id="8BA33696501B4B22FCC80C26FBBAFA71" blockId="1.[834,1475,1820,1908]" pageId="1" pageNumber="171">
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<emphasis id="B968EA84501B4B22FCC80C26FC70FA31" bold="true" box="[835,965,1820,1844]" pageId="1" pageNumber="171">Etymology.</emphasis>
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The species is named after the collector Montri Sumontha; the specific epithet “
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<taxonomicName id="4C1C4D15501B4B22FB070C06FB4CFA51" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" box="[1164,1273,1852,1876]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="171" phylum="Mollusca" rank="species" species="sumonthai">
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<emphasis id="B968EA84501B4B22FB070C06FB4CFA51" box="[1164,1273,1852,1876]" italics="true" pageId="1" pageNumber="171">sumonthai</emphasis>
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</taxonomicName>
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” is from the family name “Sumontha”.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306651D501B4B21FCC80CA6FD2FF9AD" lastPageId="2" lastPageNumber="172" pageId="1" pageNumber="171" type="diagnosis">
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<paragraph id="8BA33696501B4B21FCC80CA6FD2FF9AD" blockId="1.[835,1475,1948,2100]" lastBlockId="2.[113,754,944,1192]" lastPageId="2" lastPageNumber="172" pageId="1" pageNumber="171">
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<emphasis id="B968EA84501B4B22FCC80CA6FC0DFAB1" bold="true" box="[835,952,1948,1972]" pageId="1" pageNumber="171">Diagnosis.</emphasis>
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The characters distinguishing
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<taxonomicName id="4C1C4D15501B4B22FA890CA6FC01FAD1" authority="Tumpeesuwan & Tumpeesuwan, 2018" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="171" phylum="Mollusca" rank="species" species="sumonthai" status="sp. nov.">
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<emphasis id="B968EA84501B4B22FA890CA6FC01FAD1" italics="true" pageId="1" pageNumber="171">Aenigmatoconcha sumonthai</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25B57FF501B4B22FC4D0C86FBFBFAD1" box="[966,1102,1980,2004]" pageId="1" pageNumber="171" rank="species">new species</taxonomicNameLabel>
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, from
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<taxonomicName id="4C1C4D15501B4B22FB170C86FAA9FAD1" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2017" box="[1180,1308,1980,2004]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="171" phylum="Mollusca" rank="species" species="clivicola">
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<emphasis id="B968EA84501B4B22FB170C86FAA9FAD1" box="[1180,1308,1980,2004]" italics="true" pageId="1" pageNumber="171">A. clivicola</emphasis>
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</taxonomicName>
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are the colour patterns on mantle shell lobes and body, as well as morphology of radula and genital organs. Mantle shell lobes bear irregular black marbling pattern and numerous tiny white spots (
|
||
<figureCitation id="13272A1350184B21FEA5088AFE33FECD" box="[302,390,944,968]" captionStart="Fig" captionStartId="2.[113,148,810,832]" captionTargetBox="[113,1451,188,794]" captionTargetId="figure-74@2.[113,1451,188,794]" captionTargetPageId="2" captionText="Fig. 2. Aenigmatoconcha sumonthai, new species, in natural habitat at the type locality; photographs taken on June 2017. A, two snails with the dorsal shell surface complately covered by the mantle shell lobes; B, two snails with mantle shell lobes retracted. (Photograph courtesy of Mr. Montri Sumontha)." figureDoi="http://doi.org/10.5281/zenodo.4504604" httpUri="https://zenodo.org/record/4504604/files/figure.png" pageId="2" pageNumber="172">Fig. 2A</figureCitation>
|
||
). Two black bands, each running from the base of a long tentacle (eyestalk) to mantle edge, are present (
|
||
<figureCitation id="13272A1350184B21FF7D08CAFEF9F90D" box="[246,332,1008,1032]" captionStart="Fig" captionStartId="2.[113,148,810,832]" captionTargetBox="[113,1451,188,794]" captionTargetId="figure-74@2.[113,1451,188,794]" captionTargetPageId="2" captionText="Fig. 2. Aenigmatoconcha sumonthai, new species, in natural habitat at the type locality; photographs taken on June 2017. A, two snails with the dorsal shell surface complately covered by the mantle shell lobes; B, two snails with mantle shell lobes retracted. (Photograph courtesy of Mr. Montri Sumontha)." figureDoi="http://doi.org/10.5281/zenodo.4504604" httpUri="https://zenodo.org/record/4504604/files/figure.png" pageId="2" pageNumber="172">Fig. 2A</figureCitation>
|
||
). Central teeth of radula have a globose unicuspid cusp, with their size smaller than the 1
|
||
<superScript id="7C699BDE50184B21FD120F35FD16F918" attach="left" box="[665,675,1039,1053]" fontSize="6" pageId="2" pageNumber="172">st</superScript>
|
||
lateral teeth and only the distal halves of the globose cusps emerge from the 1
|
||
<superScript id="7C699BDE50184B21FF6A0F75FF5EF958" attach="left" box="[225,235,1103,1117]" fontSize="6" pageId="2" pageNumber="172">st</superScript>
|
||
lateral teeth (
|
||
<figureCitation id="13272A1350184B21FE0B0F6AFE60F96D" box="[384,469,1104,1128]" captionStart="Fig" captionStartId="3.[136,171,1544,1566]" captionTargetBox="[136,1473,189,1529]" captionTargetId="figure-8@3.[136,1473,189,1529]" captionTargetPageId="3" captionText="Fig. 4. Aenigmatoconcha sumonthai new species, radula of paratype (NHMSU-0016). A, whole radula; B, close up view of central part of radula; C, close up view of central, 1st and 2nd lateral teeth; D, close up view of right side of central teeth; E, close up view of right latero-marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.4504608" httpUri="https://zenodo.org/record/4504608/files/figure.png" pageId="2" pageNumber="172">Fig. 4C</figureCitation>
|
||
). The epiphallus is shorter than penis, and the middle part of the gametolytic sac in the new species is shorter than in
|
||
<taxonomicName id="4C1C4D1550184B21FE340FABFD8EF9AD" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2017" box="[447,571,1168,1192]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="clivicola">
|
||
<emphasis id="B968EA8450184B21FE340FABFD8EF9AD" box="[447,571,1168,1192]" italics="true" pageId="2" pageNumber="172">A. clivicola</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13272A1350184B21FDC20FAAFD39F9AD" box="[585,652,1168,1192]" captionStart="Fig" captionStartId="4.[113,148,1273,1295]" captionTargetBox="[113,753,420,1257]" captionTargetId="figure-70@4.[113,753,420,1257]" captionTargetPageId="4" captionText="Fig. 5. Aenigmatoconcha sumonthai, new species, genital system of paratype (NHMSU-0016). Abbreviations: ag = albumen gland, hd = hermaphroditic duct, ut = uterus, pro = prostate gland, vd = vas deferens, ep = epiphallus, erc = epiphallic retractor caecum, p = penis, gs = gametolytic sac, v = vagina, fo = free oviduct, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.4504610" httpUri="https://zenodo.org/record/4504610/files/figure.png" pageId="2" pageNumber="172">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF63661E50184B21FFFA0810FE7EFE7C" ID-DOI="http://doi.org/10.5281/zenodo.4504604" ID-Zenodo-Dep="4504604" httpUri="https://zenodo.org/record/4504604/files/figure.png" pageId="2" pageNumber="172" startId="2.[113,148,810,832]" targetBox="[113,1451,188,794]" targetPageId="2">
|
||
<paragraph id="8BA3369650184B21FFFA0810FE7EFE7C" blockId="2.[113,1451,810,889]" pageId="2" pageNumber="172">
|
||
Fig. 2.
|
||
<taxonomicName id="4C1C4D1550184B21FF320810FE67FE3A" authority="Tumpeesuwan & Tumpeesuwan, 2018" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" box="[185,466,810,831]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="sumonthai" status="sp. nov.">
|
||
<emphasis id="B968EA8450184B21FF320810FE67FE3A" box="[185,466,810,831]" italics="true" pageId="2" pageNumber="172">Aenigmatoconcha sumonthai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A25B57FF50184B21FE6A0810FDECFE45" box="[481,601,810,832]" pageId="2" pageNumber="172" rank="species">new species</taxonomicNameLabel>
|
||
, in natural habitat at the type locality; photographs taken on June 2017. A, two snails with the dorsal shell surface complately covered by the mantle shell lobes; B, two snails with mantle shell lobes retracted. (Photograph courtesy of Mr. Montri Sumontha).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C306651D50184B21FFFA0FEAFC09FA3E" pageId="2" pageNumber="172" type="description">
|
||
<paragraph id="8BA3369650184B21FFFA0FEAFF4FF8ED" blockId="2.[112,753,1232,1512]" pageId="2" pageNumber="172">
|
||
<emphasis id="B968EA8450184B21FFFA0FEAFF48F9ED" bold="true" box="[113,253,1232,1256]" pageId="2" pageNumber="172">Description.</emphasis>
|
||
Shell (
|
||
<figureCitation id="13272A1350184B21FEDB0FEAFE20F9ED" box="[336,405,1232,1256]" captionStart="Fig" captionStartId="2.[814,849,1433,1455]" captionTargetBox="[814,1450,944,1417]" captionTargetId="figure-24@2.[814,1450,944,1417]" captionTargetPageId="2" captionText="Fig. 3. External shell morphology; Aenigmatoconcha sumonthai new species, holotype (NHMSU-0015)." figureDoi="http://doi.org/10.5281/zenodo.4504606" httpUri="https://zenodo.org/record/4504606/files/figure.png" pageId="2" pageNumber="172">Fig. 3</figureCitation>
|
||
,
|
||
<tableCitation id="C69E032D50184B21FE2F0FEAFE4DF9ED" box="[420,504,1232,1256]" captionStart="Table 1" captionStartId="1.[136,191,189,211]" captionText="Table 1. Comparison of shells, radula and genital systems of Aenigmatoconcha clivicola and A. sumonthai, new species." pageId="2" pageNumber="172">Table 1</tableCitation>
|
||
): Dextral, flattened to depressed, with a rather wide and deep umbilicus. 6.80–8.00 mm high,
|
||
<quantity id="4CE49B7350184B21FF500E2AFE34F82D" box="[219,385,1296,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.285" metricValueMax="1.42" metricValueMin="1.15" pageId="2" pageNumber="172" unit="mm" value="12.85" valueMax="14.2" valueMin="11.5">11.50–14.2 mm</quantity>
|
||
wide, whorls 4¼–4½, thin, glossy, slightly convex, surface with irregular low axial ridges. Last whorl not descending in front, rounded at periphery. Colour pale brown, glass-like when fresh. Aperture large, ovately lunate, moderately oblique, with a simple and fragile margin, and a short umbilical keel at the umbilical side of the aperture.
|
||
</paragraph>
|
||
<caption id="DF63661E50184B21FCA50EA3FB06F8CE" ID-DOI="http://doi.org/10.5281/zenodo.4504606" ID-Zenodo-Dep="4504606" httpUri="https://zenodo.org/record/4504606/files/figure.png" pageId="2" pageNumber="172" startId="2.[814,849,1433,1455]" targetBox="[814,1450,944,1417]" targetPageId="2">
|
||
<paragraph id="8BA3369650184B21FCA50EA3FB06F8CE" blockId="2.[814,1449,1433,1483]" pageId="2" pageNumber="172">
|
||
Fig. 3. External shell morphology;
|
||
<taxonomicName id="4C1C4D1550184B21FB040EA3FA1CF8AB" authority="Tumpeesuwan & Tumpeesuwan, 2018" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" box="[1167,1449,1433,1454]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="sumonthai" status="sp. nov.">
|
||
<emphasis id="B968EA8450184B21FB040EA3FA1CF8AB" box="[1167,1449,1433,1454]" italics="true" pageId="2" pageNumber="172">Aenigmatoconcha sumonthai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57FF50184B21FCA50E8CFC13F8CE" box="[814,934,1461,1483]" pageId="2" pageNumber="172" rank="species">new species</taxonomicNameLabel>
|
||
, holotype (NHMSU-0015).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA3369650184B21FFFA0D2AFE13FBCD" blockId="2.[113,753,1552,1736]" pageId="2" pageNumber="172">
|
||
Body (
|
||
<figureCitation id="13272A1350184B21FF350D2AFEB8FB2D" box="[190,269,1552,1576]" captionStart="Fig" captionStartId="2.[113,148,810,832]" captionTargetBox="[113,1451,188,794]" captionTargetId="figure-74@2.[113,1451,188,794]" captionTargetPageId="2" captionText="Fig. 2. Aenigmatoconcha sumonthai, new species, in natural habitat at the type locality; photographs taken on June 2017. A, two snails with the dorsal shell surface complately covered by the mantle shell lobes; B, two snails with mantle shell lobes retracted. (Photograph courtesy of Mr. Montri Sumontha)." figureDoi="http://doi.org/10.5281/zenodo.4504604" httpUri="https://zenodo.org/record/4504604/files/figure.png" pageId="2" pageNumber="172">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="13272A1350184B21FE970D2AFEF5FB2D" box="[284,320,1552,1576]" captionStart="Fig" captionStartId="4.[112,147,2064,2086]" captionTargetBox="[112,1451,1494,2049]" captionTargetId="figure-109@4.[112,1451,1494,2049]" captionTargetPageId="4" captionText="Fig. 6. Living snails in natural habitat at type locality; photographs taken on June 2017. A, Aenigmatoconcha sumonthai, new species, on climbing plant near limestone wall; B, Sarika sp. on limestone (photographs courtesy of Mr. Montri Sumontha)." figureDoi="http://doi.org/10.5281/zenodo.4504612" httpUri="https://zenodo.org/record/4504612/files/figure.png" pageId="2" pageNumber="172">6A</figureCitation>
|
||
;
|
||
<tableCitation id="C69E032D50184B21FEC40D2AFE16FB2D" box="[335,419,1552,1576]" captionStart="Table 1" captionStartId="1.[136,191,189,211]" captionText="Table 1. Comparison of shells, radula and genital systems of Aenigmatoconcha clivicola and A. sumonthai, new species." pageId="2" pageNumber="172">Table 1</tableCitation>
|
||
): Living snails possess broad right and left transparent mantle shell lobes, with black marble pattern and numerous tiny white spots. Body pale brown to dark brown, with black bands behind the long tentacles. Foot dark brown, sole tripartite, caudal fossa and caudal horn well developed.
|
||
</paragraph>
|
||
<paragraph id="8BA3369650184B21FFFA0DCAFDFCFA8D" blockId="2.[112,754,1776,1929]" pageId="2" pageNumber="172">
|
||
Radula (
|
||
<figureCitation id="13272A1350184B21FF5E0DCAFEAAFA0D" box="[213,287,1776,1800]" captionStart="Fig" captionStartId="3.[136,171,1544,1566]" captionTargetBox="[136,1473,189,1529]" captionTargetId="figure-8@3.[136,1473,189,1529]" captionTargetPageId="3" captionText="Fig. 4. Aenigmatoconcha sumonthai new species, radula of paratype (NHMSU-0016). A, whole radula; B, close up view of central part of radula; C, close up view of central, 1st and 2nd lateral teeth; D, close up view of right side of central teeth; E, close up view of right latero-marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.4504608" httpUri="https://zenodo.org/record/4504608/files/figure.png" pageId="2" pageNumber="172">Fig. 4</figureCitation>
|
||
,
|
||
<tableCitation id="C69E032D50184B21FEBB0DCAFE3FFA0D" box="[304,394,1776,1800]" captionStart="Table 1" captionStartId="1.[136,191,189,211]" captionText="Table 1. Comparison of shells, radula and genital systems of Aenigmatoconcha clivicola and A. sumonthai, new species." pageId="2" pageNumber="172">Table 1</tableCitation>
|
||
): Rectangle, comprises 22–35 transverse v-shaped rows of teeth and 11–150 teeth in each row. All teeth unicuspid, of spatulate shape. Central teeth are at sharp angles in middle line. Half of cusp body of central teeth emerge from flanking 1
|
||
<superScript id="7C699BDE50184B21FE240C55FE0CFA78" attach="left" box="[431,441,1903,1917]" fontSize="6" pageId="2" pageNumber="172">st</superScript>
|
||
lateral teeth.
|
||
</paragraph>
|
||
<paragraph id="8BA3369650184B21FFFA0C8AFC09FA3E" blockId="2.[112,752,1968,2088]" lastBlockId="2.[811,1453,1539,1851]" pageId="2" pageNumber="172">
|
||
Genital system (
|
||
<figureCitation id="13272A1350184B21FE970C8AFEEEFACD" box="[284,347,1968,1992]" captionStart="Fig" captionStartId="4.[113,148,1273,1295]" captionTargetBox="[113,753,420,1257]" captionTargetId="figure-70@4.[113,753,420,1257]" captionTargetPageId="4" captionText="Fig. 5. Aenigmatoconcha sumonthai, new species, genital system of paratype (NHMSU-0016). Abbreviations: ag = albumen gland, hd = hermaphroditic duct, ut = uterus, pro = prostate gland, vd = vas deferens, ep = epiphallus, erc = epiphallic retractor caecum, p = penis, gs = gametolytic sac, v = vagina, fo = free oviduct, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.4504610" httpUri="https://zenodo.org/record/4504610/files/figure.png" pageId="2" pageNumber="172">Fig. 5</figureCitation>
|
||
,
|
||
<tableCitation id="C69E032D50184B21FEEE0C8AFE01FACD" box="[357,436,1968,1992]" captionStart="Table 1" captionStartId="1.[136,191,189,211]" captionText="Table 1. Comparison of shells, radula and genital systems of Aenigmatoconcha clivicola and A. sumonthai, new species." pageId="2" pageNumber="172">Table 1</tableCitation>
|
||
): Albumen gland of moderate size, slightly swollen. Hermaphroditic duct a rather thick tube. Uterus very long, large, swollen, of dark yellow colour. Vas deferens a long and slender tube, entering apically into epiphallus. Epiphallus a short slender tube, ephiphallic flagellum absent. Ephiphallic retractor caecum shorter and thicker than distal end epiphallus. Penis longer than epiphallus, sausage-shaped. Gametolytic sac comprises three portions, including swollen gland at distal end, slender tube in the middle, and robust tube at base. Vagina yellow in colour, cylindrical, and longer than free oviduct. Free oviduct divided into three parts, distal part short, orange, middle part constricted, and proximal part swollen. Amatorial organ absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306651D50184B20FCA60C59FD00FA62" lastPageId="3" lastPageNumber="173" pageId="2" pageNumber="172" type="discussion">
|
||
<paragraph id="8BA3369650184B20FCA60C59FD00FA62" blockId="2.[813,1454,1891,2107]" lastBlockId="3.[135,777,1679,1895]" lastPageId="3" lastPageNumber="173" pageId="2" pageNumber="172">
|
||
<emphasis id="B968EA8450184B21FCA60C59FC2AFA7E" bold="true" box="[813,927,1891,1915]" pageId="2" pageNumber="172">Remarks.</emphasis>
|
||
Although shell morphology and radula of
|
||
<taxonomicName id="4C1C4D1550184B21FA1F0C59FC29FA9E" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="sumonthai">
|
||
<emphasis id="B968EA8450184B21FA1F0C59FC29FA9E" italics="true" pageId="2" pageNumber="172">A. sumonthai</emphasis>
|
||
</taxonomicName>
|
||
are similar to those of
|
||
<taxonomicName id="4C1C4D1550184B21FB280CB9FAAAFA9E" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2017" box="[1187,1311,1923,1947]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="clivicola">
|
||
<emphasis id="B968EA8450184B21FB280CB9FAAAFA9E" box="[1187,1311,1923,1947]" italics="true" pageId="2" pageNumber="172">A. clivicola</emphasis>
|
||
</taxonomicName>
|
||
, the shell of
|
||
<taxonomicName id="4C1C4D1550184B21FCA60C99FC0EFABE" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" box="[813,955,1955,1979]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="sumonthai">
|
||
<emphasis id="B968EA8450184B21FCA60C99FC0EFABE" box="[813,955,1955,1979]" italics="true" pageId="2" pageNumber="172">A. sumonthai</emphasis>
|
||
</taxonomicName>
|
||
is significantly smaller and with less number of whorls (
|
||
<tableCitation id="C69E032D50184B21FC2F0CF9FC4DFADE" box="[932,1016,1987,2011]" captionStart="Table 1" captionStartId="1.[136,191,189,211]" captionText="Table 1. Comparison of shells, radula and genital systems of Aenigmatoconcha clivicola and A. sumonthai, new species." pageId="2" pageNumber="172">Table 1</tableCitation>
|
||
). The radula of
|
||
<taxonomicName id="4C1C4D1550184B21FB3B0CF9FA8BFADE" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" box="[1200,1342,1987,2011]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="sumonthai">
|
||
<emphasis id="B968EA8450184B21FB3B0CF9FA8BFADE" box="[1200,1342,1987,2011]" italics="true" pageId="2" pageNumber="172">A. sumonthai</emphasis>
|
||
</taxonomicName>
|
||
possesses transverse v-shaped rows of teeth, and the number of teeth in each row is not significantly different from
|
||
<taxonomicName id="4C1C4D1550184B21FAA20339FA10F51E" authorityName="C. Tumpeesuwan & S. Tumpeesuwan" authorityYear="2017" box="[1321,1445,2051,2075]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="clivicola">
|
||
<emphasis id="B968EA8450184B21FAA20339FA10F51E" box="[1321,1445,2051,2075]" italics="true" pageId="2" pageNumber="172">A. clivicola</emphasis>
|
||
</taxonomicName>
|
||
, when the central teeth are excluded.
|
||
<taxonomicName id="4C1C4D1550184B20FB6A0319FF42FBA2" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="3" lastPageNumber="173" order="Stylommatophora" pageId="2" pageNumber="172" phylum="Mollusca" rank="species" species="sumonthai">
|
||
<emphasis id="B968EA8450184B20FB6A0319FF42FBA2" italics="true" lastPageId="3" lastPageNumber="173" pageId="2" pageNumber="172">Aenigmatoconcha sumonthai</emphasis>
|
||
</taxonomicName>
|
||
possesses central teeth that have only half of its small globose unicuspid cusp emerging from the 1
|
||
<superScript id="7C699BDE50194B20FD3A0D94FD0EFBB9" attach="left" box="[689,699,1710,1724]" fontSize="6" pageId="3" pageNumber="173">st</superScript>
|
||
lateral teeth on both sides (
|
||
<figureCitation id="13272A1350194B20FED60DF5FE29FBE2" box="[349,412,1743,1767]" captionStart="Fig" captionStartId="3.[136,171,1544,1566]" captionTargetBox="[136,1473,189,1529]" captionTargetId="figure-8@3.[136,1473,189,1529]" captionTargetPageId="3" captionText="Fig. 4. Aenigmatoconcha sumonthai new species, radula of paratype (NHMSU-0016). A, whole radula; B, close up view of central part of radula; C, close up view of central, 1st and 2nd lateral teeth; D, close up view of right side of central teeth; E, close up view of right latero-marginal teeth." figureDoi="http://doi.org/10.5281/zenodo.4504608" httpUri="https://zenodo.org/record/4504608/files/figure.png" pageId="3" pageNumber="173">Fig. 4</figureCitation>
|
||
B–D). Genital systems are notably different, as the epiphallus of
|
||
<taxonomicName id="4C1C4D1550194B20FE580DCAFDD7FA02" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2018" box="[467,610,1775,1799]" class="Gastropoda" family="Helicarionidae" genus="Aenigmatoconcha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="173" phylum="Mollusca" rank="species" species="sumonthai">
|
||
<emphasis id="B968EA8450194B20FE580DCAFDD7FA02" box="[467,610,1775,1799]" italics="true" pageId="3" pageNumber="173">A. sumonthai</emphasis>
|
||
</taxonomicName>
|
||
is shorter than penis. The gametolytic sac comprises three parts: swollen gland at distal end, slender tube in the middle, and the base is a robust tube that is longer than the middle part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |