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<document id="B8BC49AA0931EF3A14F0D0577D4D4051" ID-CLB-Dataset="6089" ID-DOI="10.11646/zootaxa.4958.1.32" ID-GBIF-Dataset="f8b056ee-69fb-4585-8e6f-ec8a90f937b8" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692436" ID-ZooBank="4F2516D0-B025-4DA4-9C3D-F4E5093289C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618472120534" checkinUser="plazi" docAuthor="Roca-Cusachs, Marcos, Rider, Id. A. &amp; Jung, Sunghoon" docDate="2021" docId="038420420664C43FFF23FB9207D8FC57" docLanguage="en" docName="zootaxa.4958.1.32.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graziasternum Roca-Cusachs &amp; Rider &amp; Jung 2021, gen. nov." docType="treatment" docVersion="5" lastPageNumber="562" masterDocId="FFBD583A0665C43DFFB4FF890244FFC8" masterDocTitle="New genus and new species of Pentatomidae from Borneo (Hemiptera: Heteroptera)" masterLastPageNumber="569" masterPageNumber="560" pageNumber="561" updateTime="1698936910024" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BCAA5861C32AC2A4014043089562F270">New genus and new species of Pentatomidae from Borneo (Hemiptera: Heteroptera)</mods:title>
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<mods:name id="0963F51F3CFD2E75AFA0AB54F9382562" type="personal">
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<mods:namePart id="47824716B7F95A964D94B757501DF9EE">Roca-Cusachs, Marcos</mods:namePart>
<mods:affiliation id="D39DD85579FABDEA3C73C9E6E1427DD3">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture and life Sciences, Chungnam National University, Daejeon, Korea. marcosrocacusachs @ gmail. com &amp; IRBio. Institut de Recerca a la Biodiversitat, Universitat de Barcelona, Barcelona, Spain.</mods:affiliation>
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<mods:namePart id="8DA1C491BF23AC611B692B00F041F0EC">Rider, Id. A.</mods:namePart>
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<mods:namePart id="FB9B32D61BAA9BFBDFDD4C10E776657F">Jung, Sunghoon</mods:namePart>
<mods:affiliation id="79515A5AF4129B1FE4B9341D87A4F19F">Laboratory of Systematic Entomology, Department of Applied Biology, College of Agriculture and life Sciences, Chungnam National University, Daejeon, Korea. marcosrocacusachs @ gmail. com &amp; Department of Smart Agriculture Systems, College of Agriculture and life Sciences, Chungnam National University, Daejeon, Korea. Corresponding author. jung @ cnu. ac. kr</mods:affiliation>
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<mods:title id="419D9D45104C52912CEC6E8EDC9806D1">Zootaxa</mods:title>
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<mods:date id="8196EE6D32EFC9AF84E3437EB5B40CC6">2021</mods:date>
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<mods:number id="65CF461EC7D4022760EC2AE67252C7C9">2021-04-14</mods:number>
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<mods:number id="4EF0EA70069A7F0D0F27E0F4B1461E3F">4958</mods:number>
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<treatment id="038420420664C43FFF23FB9207D8FC57" ID-DOI="http://doi.org/10.5281/zenodo.4692438" ID-GBIF-Taxon="182742132" ID-Zenodo-Dep="4692438" LSID="urn:lsid:plazi:treatment:038420420664C43FFF23FB9207D8FC57" httpUri="http://treatment.plazi.org/id/038420420664C43FFF23FB9207D8FC57" lastPageId="2" lastPageNumber="562" pageId="1" pageNumber="561">
<subSubSection id="C337C2DF0664C43CFF23FB9200AAFBFD" box="[151,750,1051,1077]" pageId="1" pageNumber="561" type="nomenclature">
<paragraph id="8B9291540664C43CFF23FB9200AAFBFD" blockId="1.[151,750,1051,1077]" box="[151,750,1051,1077]" pageId="1" pageNumber="561">
<heading id="D0DA26380664C43CFF23FB9200AAFBFD" bold="true" box="[151,750,1051,1077]" fontSize="11" level="1" pageId="1" pageNumber="561" reason="1">
<emphasis id="B9594D460664C43CFF23FB9200AAFBFD" bold="true" box="[151,750,1051,1077]" pageId="1" pageNumber="561">
<taxonomicName id="4C2DEAD70664C43CFF23FB920037FBFD" authority="Roca-Cusachs &amp; Rider" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[151,627,1051,1077]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9594D460664C43CFF23FB920308FBFD" bold="true" box="[151,332,1051,1077]" italics="true" pageId="1" pageNumber="561">Graziasternum</emphasis>
Roca-Cusachs &amp; Rider
</taxonomicName>
,
<taxonomicNameLabel id="A26AF03D0664C43CFD34FB9200AAFBFD" box="[640,750,1051,1077]" pageId="1" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0664C43CFF23FBD3011BFBBA" box="[151,863,1114,1138]" pageId="1" pageNumber="561" type="type_taxon">
<paragraph id="8B9291540664C43CFF23FBD3011BFBBA" blockId="1.[151,863,1114,1138]" box="[151,863,1114,1138]" pageId="1" pageNumber="561">
<emphasis id="B9594D460664C43CFF23FBD30362FBBA" bold="true" box="[151,294,1114,1138]" pageId="1" pageNumber="561">
<typeStatus id="54962FF60664C43CFF23FBD30288FBBA" box="[151,204,1114,1138]" pageId="1" pageNumber="561">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C2DEAD70664C43CFE98FBD20052FBBA" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[300,534,1115,1138]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B9594D460664C43CFE98FBD20052FBBA" box="[300,534,1115,1138]" italics="true" pageId="1" pageNumber="561">Graziasternum graziae</emphasis>
</taxonomicName>
<emphasis id="B9594D460664C43CFDA9FBD30029FBBA" bold="true" box="[541,621,1114,1138]" pageId="1" pageNumber="561">
<taxonomicNameLabel id="A26AF03D0664C43CFDA9FBD30029FBBA" box="[541,621,1114,1138]" pageId="1" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0664C43CFF23FB120044FAEA" pageId="1" pageNumber="561" type="etymology">
<paragraph id="8B9291540664C43CFF23FB120044FAEA" blockId="1.[151,1437,1179,2034]" pageId="1" pageNumber="561">
<emphasis id="B9594D460664C43CFF23FB120366FB7D" bold="true" box="[151,290,1179,1205]" pageId="1" pageNumber="561">Etymology:</emphasis>
The genus
<taxonomicName id="4C2DEAD70664C43CFE1DFB150015FB7E" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[425,593,1180,1206]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFE1DFB150015FB7E" box="[425,593,1180,1206]" italics="true" pageId="1" pageNumber="561">Graziasternum</emphasis>
</taxonomicName>
is named after our friend and colleague Dr. Jocelia Grazia, to honour her 75th birthday, and in recognition of her prosperous career. The name
<taxonomicName id="4C2DEAD70664C43CFC3BFB490673FB12" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[911,1079,1216,1242]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFC3BFB490673FB12" box="[911,1079,1216,1242]" italics="true" pageId="1" pageNumber="561">Graziasternum</emphasis>
</taxonomicName>
is a combination of Dr. Grazias name, with the suffix
<emphasis id="B9594D460664C43CFE21FB6D03B9FB35" box="[405,509,1252,1277]" italics="true" pageId="1" pageNumber="561">osternum</emphasis>
, making reference to the flat, plate-like metasternum which it shares with the genus
<taxonomicName id="4C2DEAD70664C43CFF23FA8E0377FAE9" box="[151,307,1287,1313]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFF23FA8E0377FAE9" box="[151,307,1287,1313]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
. Gender is neuter.
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0664C43CFF73FAA201A0F966" pageId="1" pageNumber="561" type="diagnosis">
<paragraph id="8B9291540664C43CFF73FAA201A0F966" blockId="1.[151,1437,1179,2034]" pageId="1" pageNumber="561">
<emphasis id="B9594D460664C43CFF73FAA20306FA8D" bold="true" box="[199,322,1323,1349]" pageId="1" pageNumber="561">Diagnosis.</emphasis>
<taxonomicName id="4C2DEAD70664C43CFEFCFAA503B4FA8E" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[328,496,1324,1350]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9594D460664C43CFEFCFAA503B4FA8E" box="[328,496,1324,1350]" italics="true" pageId="1" pageNumber="561">Graziasternum</emphasis>
</taxonomicName>
<emphasis id="B9594D460664C43CFE43FAA20025FA8D" bold="true" box="[503,609,1323,1349]" pageId="1" pageNumber="561">
<taxonomicNameLabel id="A26AF03D0664C43CFE43FAA20025FA8D" box="[503,609,1323,1349]" pageId="1" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
reassembles the genus
<taxonomicName id="4C2DEAD70664C43CFCD3FAA20647FA8D" box="[871,1027,1323,1349]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFCD3FAA20647FA8D" box="[871,1027,1323,1349]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
in the following characters: the prosternum has a slightly elevated, medial carina; the medial carina of the mesosternum is more distinctly carinate; the metasternum is broad, prominent, plate-like, hexagonal, with posterior angles projecting slightly towards abdomen; the abdominal base has a small, broad, anteriorly-directed, ventrally-flattened tubercle; and the abdomen has a medial, longitudinal, shallow, but clearly depressed, sulcus. There are, however, several characters that will serve to separate
<taxonomicName id="4C2DEAD70664C43CFEE9FA690041FA32" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[349,517,1504,1530]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9594D460664C43CFEE9FA690041FA32" box="[349,517,1504,1530]" italics="true" pageId="1" pageNumber="561">Graziasternum</emphasis>
</taxonomicName>
<emphasis id="B9594D460664C43CFDBAFA56003CFA31" bold="true" box="[526,632,1503,1529]" pageId="1" pageNumber="561">
<taxonomicNameLabel id="A26AF03D0664C43CFDBAFA56003CFA31" box="[526,632,1503,1529]" pageId="1" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
from
<taxonomicName id="4C2DEAD70664C43CFD0BFA56011FFA31" box="[703,859,1503,1529]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFD0BFA56011FFA31" box="[703,859,1503,1529]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
. For example, species of
<taxonomicName id="4C2DEAD70664C43CFB35FA69076DFA32" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[1153,1321,1504,1530]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9594D460664C43CFB35FA69076DFA32" box="[1153,1321,1504,1530]" italics="true" pageId="1" pageNumber="561">Graziasternum</emphasis>
</taxonomicName>
<emphasis id="B9594D460664C43CFA86FA5607D8FA31" bold="true" box="[1330,1436,1503,1529]" pageId="1" pageNumber="561">
<taxonomicNameLabel id="A26AF03D0664C43CFA86FA5607D8FA31" box="[1330,1436,1503,1529]" pageId="1" pageNumber="561" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
are characterized by having concolorous punctures which do not form patterns (dark punctures forming patterns in
<taxonomicName id="4C2DEAD70664C43CFF23F9AE0377F989" box="[151,307,1575,1601]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFF23F9AE0377F989" box="[151,307,1575,1601]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
); by the absence of dark markings on the hemelytral membranes (dark hemelytral patterns present in
<taxonomicName id="4C2DEAD70664C43CFF23F9C20377F9AD" box="[151,307,1611,1637]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFF23F9C20377F9AD" box="[151,307,1611,1637]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
); by the entire, non-serrate, acute humeral angles (multiple, serrate, and blunt in
<taxonomicName id="4C2DEAD70664C43CFB78F9C2072CF9AD" box="[1228,1384,1611,1637]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFB78F9C2072CF9AD" box="[1228,1384,1611,1637]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
); by the pilosity on the legs (not pilose in
<taxonomicName id="4C2DEAD70664C43CFDF4F9E60098F941" box="[576,732,1647,1673]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFDF4F9E60098F941" box="[576,732,1647,1673]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
); and by the absence of black punctured markings on the legs (present in
<taxonomicName id="4C2DEAD70664C43CFEA1F91A03F5F965" box="[277,433,1683,1709]" class="Insecta" family="Pentatomidae" genus="Placosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="561" phylum="Arthropoda" rank="genus">
<emphasis id="B9594D460664C43CFEA1F91A03F5F965" box="[277,433,1683,1709]" italics="true" pageId="1" pageNumber="561">Placosternum</emphasis>
</taxonomicName>
); these characters clearly separate the two genera.
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0664C43FFF73F93E03EEFDA9" lastPageId="2" lastPageNumber="562" pageId="1" pageNumber="561" type="description">
<paragraph id="8B9291540664C43CFF73F93E01D0F93E" blockId="1.[151,1437,1179,2034]" pageId="1" pageNumber="561">
<emphasis id="B9594D460664C43CFF73F93E0315F919" bold="true" box="[199,337,1719,1745]" pageId="1" pageNumber="561">Description</emphasis>
. Body somewhat flattened dorsoventrally, compact, heavily punctured. Colouration not forming mottled pattern, brown or yellowish brown with some reddish areas.
</paragraph>
<paragraph id="8B9291540664C43CFF73F88903C7F84E" blockId="1.[151,1437,1179,2034]" pageId="1" pageNumber="561">Juga longer than tylus, rounded apically. Anteocular teeth absent. Compound eyes protruding beyond head profile. Each ocellus located near base of adjacent compound eye, nearly contiguous with compound eye. Antennae five-segmented, apex of segment one reaching nearly to apex of head. Apex of rostrum surpassing procoxae, not reaching mesocoxae.</paragraph>
<paragraph id="8B9291540664C43FFF73F806028BFEAD" blockId="1.[151,1437,1179,2034]" lastBlockId="2.[151,1437,151,609]" lastPageId="2" lastPageNumber="562" pageId="1" pageNumber="561">Anterior pronotal margins finely serrate; humeral angles, each slightly protruding beyond hemelytral base, acute, semi-spinose, entire (not toothed), slightly flattened towards apex. Scutellum distinctly longer than wide, basal angles, concolourous, foveate; scutellar apex narrow, clearly rounded. Hemelytra slightly covering mesial margins of connexiva; coria reddish, more or less contrasting with ground colouration; hemelytral membranes semi-hyaline, lacking black or coloured patterns; membranal veins marked. Peritremes elongate, slightly elevated, each rounded at apex, broadly L-shaped. Legs concolourous with ventral colouration, with hyaline, erect setae; tarsi 3-segmented. Prosternum broadly carinate; mesosternum distinctly carinate, not anteriorly produced; metasternum broad, prominent, ventrally flattened, hexagonal, with posterior angles slightly projecting caudad towards abdomen.</paragraph>
<paragraph id="8B9291540667C43FFF73FEE60769FE3D" blockId="2.[151,1437,151,609]" pageId="2" pageNumber="562">Connexivum uniformly coloured, finely punctate. Posterolateral angles of abdominal segments each slightly, but distinctly produced into minute, sharp denticle. Ventral surface of abdomen lightly coloured. Abdominal base directed anteriorly as broad, ventrally flattened tubercle apposed to notch in metasternal plate. Spiracles rounded, concolourous with abdomen. Abdomen with shallow, but clearly sulcate, medial longitudinal depression.</paragraph>
<paragraph id="8B9291540667C43FFF73FE7603EEFDA9" blockId="2.[151,1437,151,609]" pageId="2" pageNumber="562">Pygophore trapezoidal, slightly elongate. Genital cup well exposed dorsally. Posterolateral angles of pygophore rounded, blunt, produced caudally. Ventral margin U-shaped. Dorsal rim concave, broadly U-shaped. Parameres compact, robust, simple.</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0667C43FFF23FD3A0082FD05" box="[151,710,690,717]" pageId="2" pageNumber="562" type="multiple">
<paragraph id="8B9291540667C43FFF23FD3A0082FD05" blockId="2.[151,710,690,717]" box="[151,710,690,717]" pageId="2" pageNumber="562">
<heading id="D0DA26380667C43FFF23FD3A0082FD05" bold="true" box="[151,710,690,717]" fontSize="11" level="1" pageId="2" pageNumber="562" reason="1">
<emphasis id="B9594D460667C43FFF23FD3A0082FD05" bold="true" box="[151,710,690,717]" pageId="2" pageNumber="562">
Key to the species of
<emphasis id="B9594D460667C43FFE29FD3A0082FD05" bold="true" box="[413,710,691,717]" italics="true" pageId="2" pageNumber="562">
<taxonomicName id="4C2DEAD70667C43FFE29FD3A0016FD05" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[413,594,691,717]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="562" phylum="Arthropoda" rank="genus" status="gen. nov.">Graziasternum</taxonomicName>
<taxonomicNameLabel id="A26AF03D0667C43FFDEDFD3A0082FD05" box="[601,710,691,717]" pageId="2" pageNumber="562" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337C2DF0667C43FFF23FD7E07D8FC57" pageId="2" pageNumber="562" type="key">
<keyStep id="30D989F10667C43FFF23FD7E07D8FC57" pageId="2" pageNumber="562">
<paragraph id="8B9291540667C43FFF23FD7E07D8FC8F" blockId="2.[151,1437,759,928]" pageId="2" pageNumber="562">
<keyLead id="30DC32610667C43FFF23FD7E07D8FC8F" pageId="2" pageNumber="562">
<emphasis id="B9594D460667C43FFF23FD7E02E7FCC4" bold="true" box="[151,163,759,780]" pageId="2" pageNumber="562">1</emphasis>
General colouration brownish, head and antennal segments IV and IV concolour with ground colouration, with no black markings. Ventral surface of pronotum with a broad, lateral, contrasting dark-red band along each anterolateral margin, together resembling an inverted V-shape.................................................
<emphasis id="B9594D460667C43FFBD3FCBB07D8FC8F" bold="true" box="[1127,1436,818,839]" pageId="2" pageNumber="562">
<taxonomicName id="4C2DEAD70667C43FFBD3FCBB070EFC8F" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[1127,1354,818,839]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="562" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="B9594D460667C43FFBD3FCBB070EFC8F" bold="true" box="[1127,1354,818,839]" italics="true" pageId="2" pageNumber="562">Graziasternum graziae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26AF03D0667C43FFAE4FCBB07D8FC8F" box="[1360,1436,818,839]" pageId="2" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9291540667C43FFF23FCD907D8FC57" blockId="2.[151,1437,759,928]" pageId="2" pageNumber="562">
<keyLead id="30DC32610667C43FFF23FCD907D8FC57" pageId="2" pageNumber="562">
<emphasis id="B9594D460667C43FFF23FCD902DBFCAD" bold="true" box="[151,159,848,869]" pageId="2" pageNumber="562">-</emphasis>
General colouration yellowish brown with contrasting dark red hemelytra, head with black markings and antennal segments IV and IV with clear blackish, contrasted ring at apex. Ventral surface of pronotum uniformly coloured...............................................................................................
<emphasis id="B9594D460667C43FFBD0FC0307D8FC57" bold="true" box="[1124,1436,906,928]" pageId="2" pageNumber="562">
<taxonomicName id="4C2DEAD70667C43FFBD0FC03070EFC68" authority="Roca-Cusachs &amp; Rider &amp; Jung, 2021" authorityName="Roca-Cusachs &amp; Rider &amp; Jung" authorityYear="2021" box="[1124,1354,906,928]" class="Insecta" family="Pentatomidae" genus="Graziasternum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="562" phylum="Arthropoda" rank="species" species="joceliae" status="sp. nov.">
<emphasis id="B9594D460667C43FFBD0FC03070EFC68" bold="true" box="[1124,1354,906,928]" italics="true" pageId="2" pageNumber="562">Graziasternum joceliae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26AF03D0667C43FFAE4FC0307D8FC57" box="[1360,1436,906,927]" pageId="2" pageNumber="562" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>