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<document id="D078816D6FF8F7DC0616FDDC08889817" ID-DOI="10.5852/ejt.2018.396" ID-GBIF-Dataset="2b95a851-e1bf-4bdd-9cce-441ddc4135d2" ID-ISSN="2118-9773" ID-Zenodo-Dep="3787049" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1588696404769" checkinUser="valdenar" docAuthor="Camfield, Rebecca &amp; Hughes, Mark" docDate="2018" docId="0397A96F2F4DFFF6037BFD2AE5542D89" docLanguage="en" docName="ejt-396_camfield_hughes.pdf.imf" docOrigin="European Journal of Taxonomy 396" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia difformis W. C. Leong, C. I Peng &amp; K. F. Chung 2015" docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="FFAED1172F6DFFD7025FFFB1E00A2E5C" masterDocTitle="A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India" masterLastPageNumber="116" masterPageNumber="1" pageNumber="33" updateTime="1737367935407" updateUser="diego">
<mods:mods id="F12D7A1DEC845E258692FE2304376135" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5B8795A9D705122AF53F388D4586B233">
<mods:title id="FD6BD7583EEADCA8DDB612FCF8462725">A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India</mods:title>
</mods:titleInfo>
<mods:name id="647ACF193CA77EE50984EF2E34C2B293" type="personal">
<mods:role id="B2B344C09F453135BA2D8D1F5D51B302">
<mods:roleTerm id="BAB89D3F93F054B3D6D5B7757958D302">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DCEAEDC45E0C8731AA1F427F2C97986F">Camfield, Rebecca</mods:namePart>
<mods:affiliation id="570883F1A8CA6D9B28D72D983FA7C0AE">Royal Botanic Gardens Edinburgh, 20 a Inverleith Row, Edinburgh, EH 5 3 LR, United Kingdom.</mods:affiliation>
<mods:nameIdentifier id="980AA586754B16385E6F25ED8D00A907" type="email">RCamfield@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="D2666D382AC5C9968873C0F98C020B7E" type="personal">
<mods:role id="76F5F597D8407E0F9E6C843BA1BD49B5">
<mods:roleTerm id="7B1E762749CDE7267FE536F9D58558BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D1192282D634731C6820A0B858F21F2">Hughes, Mark</mods:namePart>
<mods:affiliation id="2124DBA125BCBF0E4656D90A3E0FD620">Royal Botanic Gardens Edinburgh, 20 a Inverleith Row, Edinburgh, EH 5 3 LR, United Kingdom.</mods:affiliation>
<mods:nameIdentifier id="D3A678AC53CFD4E29AF6275CE3B141C5" type="email">MHughes@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F2B9EC1A91DB5010BBF082F2F857AA07">text</mods:typeOfResource>
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<mods:titleInfo id="024B2347ED3DE7D0DF810640AD345E36">
<mods:title id="3F5C50A3A44780CB25A2865F3B2FEF2E">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="4EA4D4C1110395BA269997F6F0FC81C6">
<mods:date id="1D85355ECBDC28A9E03F7C7AF0968198">2018</mods:date>
<mods:detail id="66500AA295847507612E3D681A0FFF79" type="pubDate">
<mods:number id="FB4E010CEA59F7A6518E99E2A9B3411D">2018-01-19</mods:number>
</mods:detail>
<mods:detail id="FF517D11EF58BAD411CD6D4E0981E558" type="volume">
<mods:number id="E1C196DA9FD6DBEF905201F7CD483022">396</mods:number>
</mods:detail>
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<mods:start id="9F186E8D4CBAF44567C4B8680E7F3535">1</mods:start>
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<mods:classification id="AA70E855C2DB092D726BE0ACB153ACE5">journal article</mods:classification>
<mods:identifier id="2B322136D548B89C75AB23F5FF7E3F38" type="DOI">10.5852/ejt.2018.396</mods:identifier>
<mods:identifier id="AB53730FCA0ADFA8D0EF0325F49A5A69" type="GBIF-Dataset">2b95a851-e1bf-4bdd-9cce-441ddc4135d2</mods:identifier>
<mods:identifier id="5AC9C20ACDB8469909CCD18EA4BA2D87" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="1DEF09FA1902F28385F82D0252B2141F" type="Zenodo-Dep">3787049</mods:identifier>
</mods:mods>
<treatment id="0397A96F2F4DFFF6037BFD2AE5542D89" ID-DOI="http://doi.org/10.5281/zenodo.3794335" ID-GBIF-Taxon="163963839" ID-Zenodo-Dep="3794335" LSID="urn:lsid:plazi:treatment:0397A96F2F4DFFF6037BFD2AE5542D89" httpUri="http://treatment.plazi.org/id/0397A96F2F4DFFF6037BFD2AE5542D89" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" scope_class="Equisetopsida" scope_family="Begoniaceae" scope_order="Cucurbitales" scope_subClass="Magnoliidae" scope_superOrder="Rosanae">
<subSubSection id="C3244BF22F4DFFF7037BFD2AE51A2CE9" box="[292,1296,666,694]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8B8118792F4DFFF7037BFD2AE51A2CE9" blockId="32.[292,1296,666,728]" box="[292,1296,666,694]" pageId="32" pageNumber="33">
<taxonomicName id="4C3E63FA2F4DFFF7037BFD2AE4102CE9" ID-CoL="7SFDR" authority="(Irmsch.) W. C. Leong, C. I Peng &amp; K. F. Chung" authorityName="W. C. Leong, C. I Peng &amp; K. F. Chung" authorityYear="2015" baseAuthorityName="Irmsch." box="[292,1050,666,694]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="difformis">
<emphasis id="B94AC46B2F4DFFF7037BFD2AE1F42CE9" bold="true" box="[292,510,666,693]" italics="true" pageId="32" pageNumber="33">Begonia difformis</emphasis>
(Irmsch.) W.C.Leong, C.I Peng &amp; K.F.Chung
</taxonomicName>
[sect.
<taxonomicName id="4C3E63FA2F4DFFF70637FD2AE50C2CE9" box="[1128,1286,667,693]" class="Magnoliopsida" family="Begoniaceae" genus="Platycentrum" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F4DFFF70637FD2AE50C2CE9" box="[1128,1286,667,693]" italics="true" pageId="32" pageNumber="33">Platycentrum</emphasis>
</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F4DFFF700AFFD0FE3492C84" box="[752,835,702,728]" pageId="32" pageNumber="33" type="description">
<paragraph id="8B8118792F4DFFF700AFFD0FE3492C84" blockId="32.[292,1296,666,728]" box="[752,835,702,728]" pageId="32" pageNumber="33">
<figureCitation id="130504FC2F4DFFF700AFFD0FE3492C84" box="[752,835,702,728]" captionStart="Fig" captionStartId="33.[189,232,1958,1984]" captionTargetBox="[426,1161,1186,1917]" captionTargetId="figure@33.[416,1172,1174,1927]" captionTargetPageId="33" captionText="Fig. 21. Map showing the location of B. difformis (Irmsch.) W.C.Leong, C.I Peng &amp; K.F.Chung specimens." figureDoi="http://doi.org/10.5281/zenodo.3787081" httpUri="https://zenodo.org/record/3787081/files/figure.png" pageId="32" pageNumber="33">Fig. 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F4DFFF702E2FD4EE5552D3C" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="8B8118792F4DFFF702E2FD4EE5552D3C" blockId="32.[189,1399,767,935]" pageId="32" pageNumber="33">
<treatmentCitationGroup id="AB2E3F572F4DFFF702E2FD4EE5552D3C" pageId="32" pageNumber="33">
<emphasis id="B94AC46B2F4DFFF702E2FD4EE13E2D45" box="[189,308,767,793]" italics="true" pageId="32" pageNumber="33">Phytotaxa</emphasis>
227 (1): 86 (
<treatmentCitation id="0A9F3E682F4DFFF70398FCB1E2932D46" box="[455,665,767,794]" pageId="32" pageNumber="33">
<bibRefCitation id="EFAF65882F4DFFF70398FCB1E2932D46" author="Leong W. C. &amp; Deng T. &amp; Sun H. &amp; Peng C. &amp; Chung K. F." box="[455,665,767,794]" pageId="32" pageNumber="33" pagination="83 - 91" refId="ref55922" refString="Leong W. C., Deng T., Sun H., Peng C. - I &amp; Chung K. F. 2015. Begonia difformis comb. &amp; stat. nov. (sect. Platycentrum, Begoniaceae), a new species segregated from B. palmata D. Don. Phytotaxa 227: 83 - 91. https: // doi. org / 10.11646 / phytotaxa. 227.1.9" type="journal article" year="2015">
Leong
<emphasis id="B94AC46B2F4DFFF70045FCB1E2592D45" box="[538,595,767,793]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2015
</bibRefCitation>
</treatmentCitation>
). –
<taxonomicName id="4C3E63FA2F4DFFF700E0FCB1E57D2D45" ID-CoL="7SFDR" authority="Irmsch., Mitteilungen" authorityName="Irmsch., Mitteilungen" box="[703,1399,767,794]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="variety" species="lacinata" variety="difformis">
<emphasis id="B94AC46B2F4DFFF700E0FCB1E38D2D45" box="[703,903,767,793]" italics="true" pageId="32" pageNumber="33">Begonia lacinata</emphasis>
Roxb. var.
<emphasis id="B94AC46B2F4DFFF70654FD4EE47F2D45" box="[1035,1141,767,793]" italics="true" pageId="32" pageNumber="33">difformis</emphasis>
Irmsch.,
<emphasis id="B94AC46B2F4DFFF706BEFD4EE57D2D45" box="[1249,1399,767,793]" italics="true" pageId="32" pageNumber="33">Mitteilungen</emphasis>
</taxonomicName>
<emphasis id="B94AC46B2F4DFFF702E2FC92E31D2D61" box="[189,791,803,829]" italics="true" pageId="32" pageNumber="33">aus dem Institut für allgemeine Botanik in Hamburg</emphasis>
10: 531 (
<bibRefCitation id="EFAF65882F4DFFF701DCFC92E4382D61" author="Irmscher E." box="[899,1074,803,829]" pageId="32" pageNumber="33" pagination="525 - 534" refId="ref55230" refString="Irmscher E. 1939. Die Begoniaceen Chinas. Mitteilungen aus dem Institut fur allgemeine Botanik in Hamburg 10: 525 - 534." type="journal article" year="1939">Irmscher 1939</bibRefCitation>
). –
<taxonomicName id="4C3E63FA2F4DFFF70605FC92E34F2D3C" authority="(Irmsch.) Golding &amp; Kareg., Phytologia" authorityName="Golding &amp; Kareg., Phytologia" baseAuthorityName="Irmsch." class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="variety" species="palmata" variety="difformis">
<emphasis id="B94AC46B2F4DFFF70605FC92E52B2D61" box="[1114,1313,803,829]" italics="true" pageId="32" pageNumber="33">Begonia palmata</emphasis>
D.Don var.
<emphasis id="B94AC46B2F4DFFF702ADFCF7E1562D3C" box="[242,348,838,864]" italics="true" pageId="32" pageNumber="33">difformis</emphasis>
(Irmsch.) Golding &amp; Kareg.,
<emphasis id="B94AC46B2F4DFFF7009BFCF7E34F2D3C" box="[708,837,838,864]" italics="true" pageId="32" pageNumber="33">Phytologia</emphasis>
</taxonomicName>
54 (7): 495 (
<treatmentCitation id="0A9F3E682F4DFFF701B9FCF7E5582D3C" box="[998,1362,838,864]" pageId="32" pageNumber="33">
<bibRefCitation id="EFAF65882F4DFFF701B9FCF7E5582D3C" author="Golding J. &amp; Karegeannes C." box="[998,1362,838,864]" pageId="32" pageNumber="33" pagination="493 - 499" refId="ref53956" refString="Golding J. &amp; Karegeannes C. 1984. Begonia nomenclature Notes, 7. Phytologia 54 (7): 493 - 499. Available from http: // www. biodiversitylibrary. org / page / 12978493 # page / 513 / mode / 1 up [accessed Jun. 2009]." type="journal article" year="1984">Golding &amp; Karegeannes 1984</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F4DFFF70737FCF7E1D62AB6" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8B8118792F4DFFF70737FCF7E1832DFB" blockId="32.[189,1399,767,935]" pageId="32" pageNumber="33">
–
<typeStatus id="5485A6DB2F4DFFF702E2FCDBE0F62DD8" box="[189,252,874,900]" pageId="32" pageNumber="33">Type</typeStatus>
:
<materialsCitation id="3B5612242F4DFFF70357FCD8E5782DD8" box="[264,1394,873,900]" collectingDate="1912-09" collectionCode="E" collectorName="Forrest" country="China" location="Yunnan" pageId="32" pageNumber="33" specimenCode="E00299196" specimenCount="1" stateProvince="Yunnan" typeStatus="lectotype">
<collectingCountry id="F32958E92F4DFFF70357FCD8E15B2DDF" box="[264,337,873,899]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
,
<collectingRegion id="49FAD69B2F4DFFF70303FCDBE1B32DD8" box="[348,441,874,900]" country="China" name="Yunnan" pageId="32" pageNumber="33">Yunnan</collectingRegion>
,
<date id="FF803EB92F4DFFF70399FCD8E2342DDF" box="[454,574,873,899]" pageId="32" pageNumber="33" value="1912-09">
<collectingDate id="EFC4C7512F4DFFF70399FCD8E2342DDF" box="[454,574,873,899]" pageId="32" pageNumber="33" value="1912-09">Sep. 1912</collectingDate>
</date>
,
<emphasis id="B94AC46B2F4DFFF70013FCDBE2FE2DDF" box="[588,756,873,899]" italics="true" pageId="32" pageNumber="33">
<collectorName id="26CB7DAF2F4DFFF70013FCDBE2A92DDF" box="[588,675,874,899]" pageId="32" pageNumber="33">Forrest</collectorName>
16098
</emphasis>
(
<typeStatus id="5485A6DB2F4DFFF7015AFCD8E3442DDF" box="[773,846,873,899]" pageId="32" pageNumber="33">lecto-</typeStatus>
:
<specimenCode id="DB98B0022F4DFFF70105FCD8E3EB2DDF" box="[858,993,873,899]" pageId="32" pageNumber="33">E00299196</specimenCode>
, designated by
<bibRefCitation id="EFAF65882F4DFFF706C3FCDBE5782DD8" author="Leong W. C. &amp; Deng T. &amp; Sun H. &amp; Peng C. &amp; Chung K. F." box="[1180,1394,873,900]" pageId="32" pageNumber="33" pagination="83 - 91" refId="ref55922" refString="Leong W. C., Deng T., Sun H., Peng C. - I &amp; Chung K. F. 2015. Begonia difformis comb. &amp; stat. nov. (sect. Platycentrum, Begoniaceae), a new species segregated from B. palmata D. Don. Phytotaxa 227: 83 - 91. https: // doi. org / 10.11646 / phytotaxa. 227.1.9" type="journal article" year="2015">
Leong
<emphasis id="B94AC46B2F4DFFF706AFFCDBE5212DDF" box="[1264,1323,873,899]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2015
</bibRefCitation>
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF702E2FC3DE1712DFB" box="[189,379,908,935]" collectingDate="1912-09" collectionCode="K" collectorName="Forrest" country="China" location="Yunnan" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Yunnan" typeStatus="isolectotype">
<typeStatus id="5485A6DB2F4DFFF702E2FC3DE12D2DFA" box="[189,295,908,934]" pageId="32" pageNumber="33">isolecto-</typeStatus>
:
<collectionCode id="ED2F80BC2F4DFFF7036EFC3CE14C2DFB" box="[305,326,909,935]" pageId="32" pageNumber="33">K</collectionCode>
n.v.
</materialsCitation>
).
</paragraph>
<paragraph id="8B8118792F4DFFF702E2FC7CE1F32DBB" blockId="32.[189,505,973,999]" box="[189,505,973,999]" pageId="32" pageNumber="33">
<heading id="D0C9AF152F4DFFF702E2FC7CE1F32DBB" bold="true" box="[189,505,973,999]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B94AC46B2F4DFFF702E2FC7CE1F32DBB" bold="true" box="[189,505,973,999]" pageId="32" pageNumber="33">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F4DFFF702E2FC4DE1D62AB6" blockId="32.[189,1399,1020,1258]" pageId="32" pageNumber="33">
<materialsCitation id="3B5612242F4DFFF702E2FC4DE4762A4B" ID-GBIF-Occurrence="2610421311" box="[189,1148,1020,1047]" pageId="32" pageNumber="33">
<collectingCountry id="F32958E92F4DFFF702E2FC4DE11F2A4A" box="[189,277,1020,1046]" name="India" pageId="32" pageNumber="33">INDIA</collectingCountry>
:
<emphasis id="B94AC46B2F4DFFF70340FC4DE1782A4A" bold="true" box="[287,370,1020,1046]" pageId="32" pageNumber="33">
<collectingRegion id="49FAD69B2F4DFFF70340FC4DE1782A4A" box="[287,370,1020,1046]" country="India" name="Assam" pageId="32" pageNumber="33">Assam</collectingRegion>
</emphasis>
: Cachar, Boro Lakha,
<date id="FF803EB92F4DFFF70026FC4DE3022A4B" box="[633,776,1020,1047]" pageId="32" pageNumber="33" value="1951-05-01">1 May 1951</date>
,
<emphasis id="B94AC46B2F4DFFF7014BFC4DE3AD2A4A" box="[788,935,1020,1046]" italics="true" pageId="32" pageNumber="33">Koelz 27847</emphasis>
(MICH1225785).
</materialsCitation>
<materialsCitation id="3B5612242F4DFFF706DDFC4DE2A62A66" ID-GBIF-Occurrence="2610421311" pageId="32" pageNumber="33">
<emphasis id="B94AC46B2F4DFFF706DDFC4DE4F82A4A" bold="true" box="[1154,1266,1020,1046]" pageId="32" pageNumber="33">
<collectingRegion id="49FAD69B2F4DFFF706DDFC4DE4F82A4A" box="[1154,1266,1020,1046]" country="India" name="Manipur" pageId="32" pageNumber="33">Manipur</collectingRegion>
</emphasis>
: Karong,
<date id="FF803EB92F4DFFF70737FC4DE13E2A66" pageId="32" pageNumber="33" value="1950-10-08">8 Oct. 1950</date>
,
<emphasis id="B94AC46B2F4DFFF7031EFBAEE1DF2A65" box="[321,469,1055,1081]" italics="true" pageId="32" pageNumber="33">Koelz 26470</emphasis>
(MICH1225756).
</materialsCitation>
<materialsCitation id="3B5612242F4DFFF700ECFB91E1C02A01" ID-GBIF-Occurrence="2610421311" pageId="32" pageNumber="33">
<emphasis id="B94AC46B2F4DFFF700ECFB91E3342A66" bold="true" box="[691,830,1056,1082]" pageId="32" pageNumber="33">
<collectingRegion id="49FAD69B2F4DFFF700ECFB91E3342A66" box="[691,830,1056,1082]" country="India" name="Meghalaya" pageId="32" pageNumber="33">Meghalaya</collectingRegion>
</emphasis>
: Khasi Hills, Cherrapunji,
<date id="FF803EB92F4DFFF70620FB91E5112A66" box="[1151,1307,1055,1082]" pageId="32" pageNumber="33" value="1952-04-17">17 Apr. 1952</date>
,
<emphasis id="B94AC46B2F4DFFF70777FBAEE0F22A01" italics="true" pageId="32" pageNumber="33">Chand 5380</emphasis>
(MICH1225702)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF70389FBF2E4262A01" ID-GBIF-Occurrence="2610421311" box="[470,1068,1090,1117]" pageId="32" pageNumber="33">
ibid.,
<date id="FF803EB92F4DFFF70045FBF2E2B32A01" box="[538,697,1090,1117]" pageId="32" pageNumber="33" value="1952-05-14">14 May 1952</date>
,
<emphasis id="B94AC46B2F4DFFF70099FBF2E3532A01" box="[710,857,1091,1117]" italics="true" pageId="32" pageNumber="33">Koelz 29907</emphasis>
(MICH1225690)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF70667FBF2E1C72ADD" ID-GBIF-Occurrence="2610421311" pageId="32" pageNumber="33">
ibid.,
<date id="FF803EB92F4DFFF70623FBF2E5112A01" box="[1148,1307,1090,1117]" pageId="32" pageNumber="33" value="1952-05-29">29 May 1952</date>
,
<emphasis id="B94AC46B2F4DFFF70777FBF2E0F22ADC" italics="true" pageId="32" pageNumber="33">Chand 5779</emphasis>
(MICH1225812)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF70383FBD7E4252ADD" ID-GBIF-Occurrence="2610421311" box="[476,1071,1126,1153]" pageId="32" pageNumber="33">
ibid.,
<date id="FF803EB92F4DFFF7007CFBD7E2B92ADD" box="[547,691,1126,1153]" pageId="32" pageNumber="33" value="1952-06-02">2 Jun. 1952</date>
,
<emphasis id="B94AC46B2F4DFFF7009CFBD7E3532ADC" box="[707,857,1126,1152]" italics="true" pageId="32" pageNumber="33">Koelz 30159</emphasis>
(MICH1225720)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF70661FBD7E1D32AF8" ID-GBIF-Occurrence="2610421311" pageId="32" pageNumber="33">
ibid.,
<date id="FF803EB92F4DFFF706DAFBD7E52E2ADD" box="[1157,1316,1126,1153]" pageId="32" pageNumber="33" value="1952-06-18">18 Jun. 1952</date>
,
<emphasis id="B94AC46B2F4DFFF7076BFBD7E10E2AFF" italics="true" pageId="32" pageNumber="33">Koelz 30311</emphasis>
(MICH1225719)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF703B6FB38E4972AF8" ID-GBIF-Occurrence="2610421311" box="[489,1181,1161,1188]" pageId="32" pageNumber="33">
Mawphlang,
<date id="FF803EB92F4DFFF700D8FB38E32B2AF8" box="[647,801,1161,1188]" pageId="32" pageNumber="33" value="1953-07-15">15 Jul. 1953</date>
,
<emphasis id="B94AC46B2F4DFFF7016FFB38E3CD2AFF" box="[816,967,1161,1187]" italics="true" pageId="32" pageNumber="33">Koelz 33304</emphasis>
(MICH1225704)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF706F3FB38E27E2A9B" ID-GBIF-Occurrence="2610421311" pageId="32" pageNumber="33">
Pynursla,
<date id="FF803EB92F4DFFF7077AFB38E0F02A9A" pageId="32" pageNumber="33" value="1949-08-03">3 Aug. 1949</date>
,
<emphasis id="B94AC46B2F4DFFF70355FB1DE1952A9A" box="[266,415,1196,1222]" italics="true" pageId="32" pageNumber="33">Chand 2127</emphasis>
(MICH1225687)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF700DBFB1DE4D62A9B" ID-GBIF-Occurrence="2610421311" box="[644,1244,1196,1223]" pageId="32" pageNumber="33">
ibid.,
<date id="FF803EB92F4DFFF70093FB1DE3682A9A" box="[716,866,1196,1223]" pageId="32" pageNumber="33" value="1949-09-05">5 Sep. 1949</date>
,
<emphasis id="B94AC46B2F4DFFF7012DFB1DE40D2A9A" box="[882,1031,1196,1222]" italics="true" pageId="32" pageNumber="33">Chand 2170</emphasis>
(MICH1225688)
</materialsCitation>
;
<materialsCitation id="3B5612242F4DFFF706B3FB1DE1D62AB6" ID-GBIF-Occurrence="2610421311" pageId="32" pageNumber="33">
ibid.,
<emphasis id="B94AC46B2F4DFFF7076BFB1DE10C2AB6" italics="true" pageId="32" pageNumber="33">Koelz 23758</emphasis>
(MICH1225747).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F4DFFF602E2FAA0E21E2F15" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="8B8118792F4DFFF702E2FAA0E1442B77" blockId="32.[189,334,1297,1323]" box="[189,334,1297,1323]" pageId="32" pageNumber="33">
<heading id="D0C9AF152F4DFFF702E2FAA0E1442B77" bold="true" box="[189,334,1297,1323]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B94AC46B2F4DFFF702E2FAA0E1442B77" bold="true" box="[189,334,1297,1323]" pageId="32" pageNumber="33">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F4DFFF602E2FA8EE21E2F15" blockId="32.[189,1399,1343,2004]" lastBlockId="33.[189,1398,267,329]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
Rhizomatous, erect, monoecious herb,
<quantity id="4CC6B59C2F4DFFF700D3FA8EE3032B06" box="[652,777,1343,1370]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="32" pageNumber="33" unit="cm" value="50.0" valueMax="80.0" valueMin="20.0">20 –80 cm</quantity>
high. Rhizome: ca
<quantity id="4CC6B59C2F4DFFF701ABFA8EE4252B06" box="[1012,1071,1343,1370]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="cm" value="2.0">2 cm</quantity>
wide, internodes
<quantity id="4CC6B59C2F4DFFF7075AFA8EE57D2B06" box="[1285,1399,1343,1370]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="32" pageNumber="33" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">1–2.5 cm</quantity>
long. Stem:
<quantity id="4CC6B59C2F4DFFF7030FFAD2E1DD2B21" box="[336,471,1379,1405]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="32" pageNumber="33" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
wide, sparsely hirsute, internodes
<quantity id="4CC6B59C2F4DFFF70127FAD3E3E92B21" box="[888,995,1378,1405]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="32" pageNumber="33" unit="cm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 cm</quantity>
long. Stipules: narrowly ovate or ovate, 30 ×
<quantity id="4CC6B59C2F4DFFF7030EFA37E1A02BFC" box="[337,426,1414,1440]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="mm" value="20.0">20 mm</quantity>
, caduceus. Leaves: petiole
<quantity id="4CC6B59C2F4DFFF700AAFA37E3792BFC" box="[757,883,1414,1440]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="32" pageNumber="33" unit="cm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 cm</quantity>
long, sparsely to densely hirsute, ovate or oblate-orbicular, base cordate, lobes not overlapping, 7–17 ×
<quantity id="4CC6B59C2F4DFFF701E9FA18E42F2B98" box="[950,1061,1449,1476]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="32" pageNumber="33" unit="cm" value="8.0" valueMax="11.0" valueMin="5.0">5–11 cm</quantity>
, asymmetric, upper surface green, hispidulous along veins, lower surface paler green, hirsute; venation palmate, midrib
<quantity id="4CC6B59C2F4DFFF70753FA7DE57D2BBB" box="[1292,1399,1484,1511]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="14.0" metricValueMin="5.0" pageId="32" pageNumber="33" unit="cm" value="9.5" valueMax="14.0" valueMin="5.0">5–14 cm</quantity>
long; margin shallowly denticulate, usually divided to ¼ of the blade or more, apex acuminate to longacuminate. Inflorescences: cymose, protandrous; axillary, few; peduncle red pubescent, branching 1–2 times, primary
<quantity id="4CC6B59C2F4DFFF703F5F987E229280C" box="[426,547,1590,1617]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="32" pageNumber="33" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
long, secondary
<quantity id="4CC6B59C2F4DFFF700AFF987E341280C" box="[752,843,1590,1616]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="32" pageNumber="33" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
long, with 3–
<specimenCount id="9D38D3F02F4DFFF701A9F987E46A280C" box="[1014,1120,1590,1616]" pageId="32" pageNumber="33" type="female">4 female</specimenCount>
and 3–
<specimenCount id="9D38D3F02F4DFFF706E3F987E505280C" box="[1212,1295,1590,1616]" pageId="32" pageNumber="33" type="male">4 male</specimenCount>
flowers; bracts ovate, 10–15 ×
<quantity id="4CC6B59C2F4DFFF70399F9E8E23B2828" box="[454,561,1625,1652]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="32" pageNumber="33" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
, upper surface sparingly hispidulous to nearly glabrous, margin ciliate, caduceus. Male flowers: pedicel
<quantity id="4CC6B59C2F4DFFF70065F9CDE2B428CB" box="[570,702,1660,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="32" pageNumber="33" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
long, densely red puberulous; tepals 4, outer tepals obovate to orbicular, 12–23 ×
<quantity id="4CC6B59C2F4DFFF70390F911E24628E6" box="[463,588,1696,1722]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="2.0" metricValueMin="0.8" pageId="32" pageNumber="33" unit="mm" value="14.0" valueMax="20.0" valueMin="8.0">8–20 mm</quantity>
, white to pink, red setulose on reverse, margin entire, inner tepals oblanceolate to narrowly obovate, 10–23 ×
<quantity id="4CC6B59C2F4DFFF700E3F973E33A2881" box="[700,816,1730,1757]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="32" pageNumber="33" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
, white, glabrous, entire; androecium with 80–200 stamens, symmetric; filaments
<quantity id="4CC6B59C2F4DFFF7007CF957E291295C" box="[547,667,1766,1793]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="32" pageNumber="33" unit="mm" value="1.75" valueMax="2.5" valueMin="1.0">1–2.5 mm</quantity>
long, unequal, fused at base into short column; anther narrowly obovate,
<quantity id="4CC6B59C2F4DFFF70375F8B8E1652978" box="[298,367,1801,1828]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="mm" value="2.0">2 mm</quantity>
long, dehiscing through slits, not hooded, connective extended, acute. Female flowers: pedicel
<quantity id="4CC6B59C2F4DFFF70342F89DE1AF291B" box="[285,421,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="32" pageNumber="33" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
long, densely red puberulous; bracteoles absent; tepals 5, unequal, oblanceolate to orbicular, 8–24 ×
<quantity id="4CC6B59C2F4DFFF703E8F8E1E23B2936" box="[439,561,1872,1898]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.1" metricValueMin="0.4" pageId="32" pageNumber="33" unit="mm" value="12.5" valueMax="21.0" valueMin="4.0">4–21 mm</quantity>
, red setulose on reverse to almost glabrous, entire, inner tepals similar but smaller; ovary 2-locular, placentae bifid; capsule oblong-ellipsoid, 5–8 ×
<quantity id="4CC6B59C2F4DFFF7062EF8C2E4D529D1" box="[1137,1247,1907,1933]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="32" pageNumber="33" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, very sparse to dense red setulose, with one long wing and two short rounded wings; styles 2, forked once and twisted, caduceus. Fruit: recurved, capsule obovoid, ca 1.5 ×
<quantity id="4CC6B59C2F4DFFF701F4F808E3E12988" box="[939,1003,1977,2004]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="cm" value="1.0">1 cm</quantity>
, sparse red setulose to glabrous; wings extending along pedicel, unequal; longest wing obliquely oblong, 10–30 ×
<quantity id="4CC6B59C2F4CFFF606C1FEBAE51C2F7A" box="[1182,1302,267,294]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="33" pageNumber="34" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
; shorter wings oblong, ca 5 ×
<quantity id="4CC6B59C2F4CFFF603E5FE9EE2052F15" box="[442,527,303,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F4CFFF602E2FEC1E1B32F80" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8B8118792F4CFFF602E2FEC1E21F2FD6" blockId="33.[189,533,368,394]" box="[189,533,368,394]" pageId="33" pageNumber="34">
<heading id="D0C9AF152F4CFFF602E2FEC1E21F2FD6" bold="true" box="[189,533,368,394]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B94AC46B2F4CFFF602E2FEC1E21F2FD6" bold="true" box="[189,533,368,394]" pageId="33" pageNumber="34">Distribution and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F4CFFF602E2FE2EE1B32F80" blockId="33.[189,1399,414,476]" pageId="33" pageNumber="34">
Assam, Meghalaya and Manipur; also in
<collectingCountry id="F32958E92F4CFFF600EDFE2FE2F22FE4" box="[690,760,414,440]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
and
<collectingCountry id="F32958E92F4CFFF60168FE2EE3A12FE5" box="[823,939,415,441]" name="Myanmar" pageId="33" pageNumber="34">Myanmar</collectingCountry>
. Flowering: June to October; fruiting: August to November.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F4CFFF602E2FDB2E51C2C84" pageId="33" pageNumber="34" type="conservation">
<paragraph id="8B8118792F4CFFF602E2FDB2E1BF2C41" blockId="33.[189,437,515,541]" box="[189,437,515,541]" pageId="33" pageNumber="34">
<heading id="D0C9AF152F4CFFF602E2FDB2E1BF2C41" bold="true" box="[189,437,515,541]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B94AC46B2F4CFFF602E2FDB2E1BF2C41" bold="true" box="[189,437,515,541]" pageId="33" pageNumber="34">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F4CFFF602E2FD83E51C2C84" blockId="33.[189,1399,560,728]" pageId="33" pageNumber="34">
Least Concern. The species has an EOO of
<quantity id="4CC6B59C2F4CFFF60090FD83E3572C10" box="[719,861,562,588]" metricMagnitude="8" metricUnit="m" metricValue="1.7913" pageId="33" pageNumber="34" unit="km" value="179130.0">179,130 km</quantity>
<superScript id="7C4BB5312F4CFFF60102FD81E36C2C62" attach="left" box="[861,870,560,574]" fontSize="6" pageId="33" pageNumber="34">2</superScript>
and an AOO of
<quantity id="4CC6B59C2F4CFFF6066CFD83E4842C10" box="[1075,1166,562,588]" metricMagnitude="5" metricUnit="m" metricValue="1.08" pageId="33" pageNumber="34" unit="km" value="108.0">108 km</quantity>
<superScript id="7C4BB5312F4CFFF606D1FD81E49D2C62" attach="left" box="[1166,1175,560,574]" fontSize="6" pageId="33" pageNumber="34">2</superScript>
. The range of this recently described species could possibly be extended further, as it is likely more specimens will be found which are currently identified as the similar
<taxonomicName id="4C3E63FA2F4CFFF60143FDC8E3952CCE" authorityName="D. Don (Don" authorityYear="1825" box="[796,927,632,658]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F4CFFF60143FDC8E3952CCE" box="[796,927,632,658]" italics="true" pageId="33" pageNumber="34">B. palmata</emphasis>
</taxonomicName>
. The specimens we have cited are split between
<collectingRegion id="49FAD69B2F4CFFF60379FD2DE1892CEA" box="[294,387,668,694]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan</collectingRegion>
,
<collectingCountry id="F32958E92F4CFFF603CFFD2AE1DC2CE9" box="[400,470,667,693]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
and the study area which are linked by the Arakan Mountains and the eastern end of the Himalayas, providing suitable habitat which is currently under-explored botanically.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F4CFFF602E2FCB1E5542D89" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8B8118792F4CFFF602E2FCB1E1252D46" blockId="33.[189,303,768,794]" box="[189,303,768,794]" pageId="33" pageNumber="34">
<heading id="D0C9AF152F4CFFF602E2FCB1E1252D46" bold="true" box="[189,303,768,794]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B94AC46B2F4CFFF602E2FCB1E1252D46" bold="true" box="[189,303,768,794]" pageId="33" pageNumber="34">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F4CFFF602E2FC9EE5542D89" blockId="33.[189,1399,814,982]" pageId="33" pageNumber="34">
<taxonomicName id="4C3E63FA2F4CFFF602E2FC9EE19B2D14" authorityName="W. C. Leong, C. I Peng &amp; K. F. Chung" authorityYear="2015" baseAuthorityName="Irmsch." box="[189,401,814,840]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="difformis">
<emphasis id="B94AC46B2F4CFFF602E2FC9EE19B2D14" box="[189,401,814,840]" italics="true" pageId="33" pageNumber="34">Begonia difformis</emphasis>
</taxonomicName>
is very similar to
<taxonomicName id="4C3E63FA2F4CFFF60038FC9EE2E22D14" authorityName="D. Don (Don" authorityYear="1825" box="[615,744,814,840]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F4CFFF60038FC9EE2E22D14" box="[615,744,814,840]" italics="true" pageId="33" pageNumber="34">B. palmata</emphasis>
</taxonomicName>
but differs in having leaves with deeper lobes and with the hairs on the upper leaf surface being restricted to the veins (
<taxonomicName id="4C3E63FA2F4CFFF601FCFCE2E4292D30" authorityName="D. Don (Don" authorityYear="1825" box="[931,1059,850,876]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F4CFFF601FCFCE2E4292D30" box="[931,1059,850,876]" italics="true" pageId="33" pageNumber="34">B. palmata</emphasis>
</taxonomicName>
has hairs all over). The most notable difference are the red setose hairs on the flowers and fruit;
<taxonomicName id="4C3E63FA2F4CFFF601B5FCC7E4672DD3" authorityName="D. Don (Don" authorityYear="1825" box="[1002,1133,885,911]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F4CFFF601B5FCC7E4672DD3" box="[1002,1133,885,911]" italics="true" pageId="33" pageNumber="34">B. palmata</emphasis>
</taxonomicName>
has fine villose hairs. Also the largest wing of the fruit in
<taxonomicName id="4C3E63FA2F4CFFF60006FC28E2E92DEE" authorityName="W. C. Leong, C. I Peng &amp; K. F. Chung" authorityYear="2015" baseAuthorityName="Irmsch." box="[601,739,920,946]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="difformis">
<emphasis id="B94AC46B2F4CFFF60006FC28E2E92DEE" box="[601,739,920,946]" italics="true" pageId="33" pageNumber="34">B. difformis</emphasis>
</taxonomicName>
has a verrucose texture, unlike the smooth surface of the wings in the fruit of
<taxonomicName id="4C3E63FA2F4CFFF603F2FC0DE2242D89" authorityName="D. Don (Don" authorityYear="1825" box="[429,558,955,981]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="palmata">
<emphasis id="B94AC46B2F4CFFF603F2FC0DE2242D89" box="[429,558,955,981]" italics="true" pageId="33" pageNumber="34">B. palmata</emphasis>
</taxonomicName>
. The description is augmented with information from the protologue.
</paragraph>
</subSubSection>
</treatment>
</document>
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