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<document id="705C18BF0B53F791A7819791AC7F55C3" ID-CLB-Dataset="30595" ID-DOI="10.11646/zootaxa.4388.4.1" ID-GBIF-Dataset="a884362d-a9f8-4cee-8cd4-dcf59a73ede2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1189072" ID-ZooBank="7A57C29B-6B16-445E-A8D7-750C736E6B42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520318467094" checkinUser="plazi" docAuthor="Cupello, Mario" docDate="2018" docId="5C282C04FFCFFFEDFF2D57C81AD1FB87" docLanguage="en" docName="zootaxa.4388.4.1.pdf" docOrigin="Zootaxa 4388 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Canthidium (Canthidium) variolosum Howden &amp; Young 1981" docType="treatment" docVersion="10" lastPageNumber="465" masterDocId="A011547CFFC1FFE3FFBA541B1E1AFFFB" masterDocTitle="On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species" masterLastPageNumber="486" masterPageNumber="451" pageNumber="465" updateTime="1698528354521" updateUser="ExternalLinkService">
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<mods:title id="2931AF63E401617B20E4B168E26AA8EF">On the types species of the New World dung beetle genus Canthidium Erichson, 1847 (Coleoptera: Scarabaeidae: Scarabaeinae), with an annotated checklist of species</mods:title>
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<mods:namePart id="D438AE29DC2A3ADCAF250A343084646B">Cupello, Mario</mods:namePart>
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<mods:date id="C4F23B0455A73993C5B514D7550DBF81">2018</mods:date>
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<treatment id="5C282C04FFCFFFEDFF2D57C81AD1FB87" ID-DOI="http://doi.org/10.5281/zenodo.5994878" ID-GBIF-Taxon="141280777" ID-Zenodo-Dep="5994878" LSID="urn:lsid:plazi:treatment:5C282C04FFCFFFEDFF2D57C81AD1FB87" httpUri="http://treatment.plazi.org/id/5C282C04FFCFFFEDFF2D57C81AD1FB87" lastPageNumber="465" pageId="14" pageNumber="465">
<subSubSection id="9C9BCE99FFCFFFEDFF2D57C81D6AFC16" box="[151,880,979,1005]" pageId="14" pageNumber="465" type="nomenclature">
<paragraph id="D43E9D12FFCFFFEDFF2D57C81D6AFC16" blockId="14.[151,1437,979,1149]" box="[151,880,979,1005]" pageId="14" pageNumber="465">
<heading id="8F762A7EFFCFFFEDFF2D57C81D6AFC16" bold="true" box="[151,880,979,1005]" fontSize="10" level="2" pageId="14" pageNumber="465" reason="2">
<taxonomicName id="1381E691FFCFFFEDFF2D57C81D6AFC16" ID-CoL="68XF8" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,880,979,1005]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="variolosum" subGenus="Canthidium">
<emphasis id="E6F54100FFCFFFEDFF2D57C81D6AFC16" bold="true" box="[151,880,979,1005]" pageId="14" pageNumber="465">
<emphasis id="E6F54100FFCFFFEDFF2D57C81C57FC17" bold="true" box="[151,589,979,1004]" italics="true" pageId="14" pageNumber="465">Canthidium (Canthidium) variolosum</emphasis>
<bibRefCitation id="B010E0E3FFCFFFEDFDEE57CF1D6AFC16" box="[596,880,980,1005]" pageId="14" pageNumber="465" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="9C9BCE99FFCFFFEDFF7D57E31AD1FB87" pageId="14" pageNumber="465" type="reference_group">
<paragraph id="D43E9D12FFCFFFEDFF7D57E31AD1FB87" blockId="14.[151,1437,979,1149]" pageId="14" pageNumber="465">
<treatmentCitationGroup id="F491BA3CFFCFFFEDFF7D57E31AD1FB87" infered="true" pageId="14" pageNumber="465">
<taxonomicName id="1381E691FFCFFFEDFF7D57E31D36FBEB" ID-CoL="68XF8" authority="Howden &amp; Young, 1981: 88" authorityName="Howden &amp; Young" authorityPageNumber="88" authorityYear="1981" box="[199,812,1015,1040]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="variolosum">
<emphasis id="E6F54100FFCFFFEDFF7D57E31FCEFBEB" box="[199,468,1016,1040]" italics="true" pageId="14" pageNumber="465">Canthidium variolosum</emphasis>
<treatmentCitation id="5520BB03FFCFFFEDFE6557EC1D36FBEB" author="" box="[479,812,1015,1040]" page="88" pageId="14" pageNumber="465">
<bibRefCitation id="B010E0E3FFCFFFEDFE6557EC1D1BFBEB" box="[479,769,1015,1040]" pageId="14" pageNumber="465" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young, 1981</bibRefCitation>
: 88
</treatmentCitation>
</taxonomicName>
[here transferred to the subgenus
<taxonomicName id="1381E691FFCFFFEDFB7857E31B5CFBEB" ID-CoL="3GVY" authorityName="Erichson" authorityYear="1847" box="[1218,1350,1016,1040]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="E6F54100FFCFFFEDFB7857E31B5CFBEB" box="[1218,1350,1016,1040]" italics="true" pageId="14" pageNumber="465">Canthidium</emphasis>
</taxonomicName>
].
<emphasis id="E6F54100FFCFFFEDFADB57EC1EEDFBCE" bold="true" pageId="14" pageNumber="465">
<typeStatus id="0B3A23B0FFCFFFEDFADB57EC1B86FBEB" box="[1377,1436,1015,1040]" pageId="14" pageNumber="465">Type</typeStatus>
locality:
</emphasis>
<collectingRegion id="164553F0FFCFFFEDFF4750061F40FBCF" box="[253,346,1053,1076]" country="Panama" name="Panama" pageId="14" pageNumber="465">
<collectingCountry id="AC96DD82FFCFFFEDFF4750061F40FBCF" box="[253,346,1053,1076]" name="Panama" pageId="14" pageNumber="465">Panama</collectingCountry>
</collectingRegion>
:
<collectingRegion id="164553F0FFCFFFEDFEDF50061FD8FBCF" box="[357,450,1053,1076]" country="Panama" name="Panama" pageId="14" pageNumber="465">
<collectingCountry id="AC96DD82FFCFFFEDFEDF50061FD8FBCF" box="[357,450,1053,1076]" name="Panama" pageId="14" pageNumber="465">Panama</collectingCountry>
</collectingRegion>
: San Carlos:
<collectingRegion id="164553F0FFCFFFEDFDEE50061CD5FBCE" box="[596,719,1053,1077]" country="Philippines" name="Laguna" pageId="14" pageNumber="465">La Laguna</collectingRegion>
,
<quantity id="137930F7FFCFFFEDFD6050071D08FBCE" box="[730,786,1052,1077]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="465" unit="km" value="4.0">4 km</quantity>
west of Hato del Volcán (“
<collectingRegion id="164553F0FFCFFFEDFB8750071A86FBCF" box="[1085,1180,1052,1076]" country="Panama" name="Panama" pageId="14" pageNumber="465">
<collectingCountry id="AC96DD82FFCFFFEDFB8750071A86FBCF" box="[1085,1180,1052,1076]" name="Panama" pageId="14" pageNumber="465">Panamá</collectingCountry>
</collectingRegion>
, Las Lagunas,
<quantity id="137930F7FFCFFFEDFAF850071B61FBCE" box="[1346,1403,1052,1077]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="465" unit="km" value="4.0">4 km</quantity>
W Hato del Volcán”).
<emphasis id="E6F54100FFCFFFEDFECE50241C3FFBA3" bold="true" box="[372,549,1087,1112]" pageId="14" pageNumber="465">
<typeStatus id="0B3A23B0FFCFFFEDFECE50241FB5FBA3" box="[372,431,1087,1112]" pageId="14" pageNumber="465">Type</typeStatus>
material:
</emphasis>
<typeStatus id="0B3A23B0FFCFFFEDFD9550241C8DFBA3" box="[559,663,1087,1112]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
male (CMNC—ex Howden collection); see full list of
<typeStatus id="0B3A23B0FFCFFFEDFAA8505B1B67FBA3" box="[1298,1405,1088,1112]" pageId="14" pageNumber="465" type="paratype">paratypes</typeStatus>
in
<treatmentCitation id="5520BB03FFCFFFEDFF2D507F1FF8FB87" author="" box="[151,482,1124,1149]" page="88" pageId="14" pageNumber="465">
<bibRefCitation id="B010E0E3FFCFFFEDFF2D507F1FF8FB87" box="[151,482,1124,1149]" pageId="14" pageNumber="465" refId="ref27924" refString="Howden, H. F. &amp; Young, O. P. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article">Howden &amp; Young (1981: 88)</bibRefCitation>
</treatmentCitation>
.
<emphasis id="E6F54100FFCFFFEDFE57507F1C93FB86" bold="true" box="[493,649,1124,1149]" pageId="14" pageNumber="465">Distribution:</emphasis>
<collectingCountry id="AC96DD82FFCFFFEDFD2A507F1D10FB86" box="[656,778,1124,1149]" name="Costa Rica" pageId="14" pageNumber="465">Costa Rica</collectingCountry>
and
<collectingRegion id="164553F0FFCFFFEDFCFA507E1D83FB87" box="[832,921,1125,1148]" country="Panama" name="Panama" pageId="14" pageNumber="465">
<collectingCountry id="AC96DD82FFCFFFEDFCFA507E1D83FB87" box="[832,921,1125,1148]" name="Panama" pageId="14" pageNumber="465">Panama</collectingCountry>
</collectingRegion>
(Solís &amp; Kohlmann 2004).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>
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