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<document id="4233D51E2ABBCCCA103B863DFE21229C" ID-CLB-Dataset="53451" ID-DOI="10.11646/palaeoentomology.6.2.3" ID-GBIF-Dataset="43be93d6-0de1-47bd-aa74-16ee12d54c4e" ID-ISSN="2624-2834" ID-Zenodo-Dep="7942954" ID-ZooBank="0071B62F-E205-4EBD-BBD2-63776364F2E6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1683888063692" checkinUser="plazi" docAuthor="SUKATSHEVA, IRINA D. &amp; SINITSHENKOVA, NINA D." docDate="2023" docId="485087B820104B23FCCCA10CFC57F90E" docLanguage="en" docName="palaeoentomology.6.2.3.pdf" docOrigin="Palaeoentomology 6 (2)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.2.3" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Kamopanorpa aristovi SUKATSHEVA &amp; SINITSHENKOVA 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="121" masterDocId="B469FFC020104B22FFE6A414FF9CFF92" masterDocTitle="A new caddisfly species Kamopanorpa aristovi sp. nov. of the family Microptysmatidae (Insecta, Trichoptera) from the Middle Permian of Udmurtya, Russia" masterLastPageNumber="123" masterPageNumber="120" pageNumber="120" updateTime="1685384549411" updateUser="ExternalLinkService">
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<mods:title id="06A8AE3D4835F965E503F09715C15496">A new caddisfly species Kamopanorpa aristovi sp. nov. of the family Microptysmatidae (Insecta, Trichoptera) from the Middle Permian of Udmurtya, Russia</mods:title>
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<mods:name id="EFC40A38D72627EB83E9467A9278241E" type="personal">
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<mods:namePart id="220E39B6047323EDBABB08C4925C4988">SUKATSHEVA, IRINA D.</mods:namePart>
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<mods:namePart id="9F4833D0E763FCE231381D170C18FE1B">SINITSHENKOVA, NINA D.</mods:namePart>
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<mods:title id="C15FCB9FAF04C861815FCDA23E62B67F">Palaeoentomology</mods:title>
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<mods:date id="9AC3F3DBE8A0A13B22B38B7C9C955985">2023</mods:date>
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<mods:identifier id="00E2B120BA45C0B2D219F3B57287AC4D" type="CLB-Dataset">53451</mods:identifier>
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<treatment id="485087B820104B23FCCCA10CFC57F90E" ID-DOI="http://doi.org/10.5281/zenodo.7982760" ID-Zenodo-Dep="7982760" LSID="urn:lsid:plazi:treatment:485087B820104B23FCCCA10CFC57F90E" httpUri="http://treatment.plazi.org/id/485087B820104B23FCCCA10CFC57F90E" lastPageId="1" lastPageNumber="121" pageId="0" pageNumber="120">
<subSubSection id="88E3652520104B22FCCCA10CFCC7FAC5" pageId="0" pageNumber="120" type="nomenclature">
<paragraph id="C04636AE20104B22FCCCA10CFCC7FAC5" blockId="0.[810,1451,1304,1403]" pageId="0" pageNumber="120">
<heading id="9B0E81C220104B22FCCCA10CFCC7FAC5" bold="true" centered="true" fontSize="10" level="2" pageId="0" pageNumber="120" reason="3">
<emphasis id="F28DEABC20104B22FCCCA10CFCC7FAC5" bold="true" pageId="0" pageNumber="120">
<taxonomicName id="07F94D2D20104B22FCCCA10CFA18FAA0" authority="Sukatsheva &amp; Sinitshenkova" authorityName="SUKATSHEVA &amp; SINITSHENKOVA" authorityYear="2023" box="[810,1412,1304,1330]" class="Insecta" family="Microptysmatidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="0" pageNumber="120" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
<emphasis id="F28DEABC20104B22FCCCA10CFBB7FAA0" bold="true" box="[810,1067,1304,1330]" italics="true" pageId="0" pageNumber="120">Kamopanorpa aristovi</emphasis>
Sukatsheva &amp; Sinitshenkova
</taxonomicName>
<taxonomicNameLabel id="E9BE57C720104B22FA6CA10CFCC7FAC5" pageId="0" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="88E3652520104B22FCCCA177FCE6FAE9" box="[810,890,1379,1403]" pageId="0" pageNumber="120" type="description">
<paragraph id="C04636AE20104B22FCCCA177FCE6FAE9" blockId="0.[810,1451,1304,1403]" box="[810,890,1379,1403]" pageId="0" pageNumber="120">
(
<figureCitation id="58C22A2B20104B22FCD4A177FCF3FAE9" box="[818,879,1379,1403]" captionStart="FIGURE 1" captionStartId="1.[136,235,1279,1303]" captionTargetBox="[215,1371,189,1256]" captionTargetId="figure-380@1.[215,1371,189,1256]" captionTargetPageId="1" captionText="FIGURE 1. Kamopanorpa aristovi sp. nov., holotype PIN, no 3695/141, locality Kostovaty (Middle Ural, Udmurtya, Russia; Middle Permian, Lower Urzhum Formation, Urzhum Horizon). A, Photograph. B, Fore wing. Scale bars = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7942956" httpUri="https://zenodo.org/record/7942956/files/figure.png" pageId="0" pageNumber="120">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="88E3652520104B22FCCCA1BFFB47FA75" pageId="0" pageNumber="120" type="materials_examined">
<paragraph id="C04636AE20104B22FCCCA1BFFB47FA75" blockId="0.[810,1452,1451,2092]" pageId="0" pageNumber="120">
<materialsCitation id="70913CF320104B22FCCCA1BFFB47FA75" collectionCode="PIN" pageId="0" pageNumber="120" specimenCount="1" typeStatus="holotype">
<emphasis id="F28DEABC20104B22FCCCA1BFFC04FA51" bold="true" box="[810,920,1451,1475]" pageId="0" pageNumber="120">
<typeStatus id="1F42880C20104B22FCCCA1BFFC08FA51" box="[810,916,1451,1475]" pageId="0" pageNumber="120">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="A6E8AE6B20104B22FC45A1B8FC4DFA51" box="[931,977,1452,1475]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="0" pageNumber="120">PIN</collectionCode>
, no 3695/141, part and counterpart of the caddisfly fore wing of good preservation.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="88E3652520104B22FCB9A1E0FA9AF9A2" pageId="0" pageNumber="120" type="etymology">
<paragraph id="C04636AE20104B22FCB9A1E0FA9AF9A2" blockId="0.[810,1452,1451,2092]" pageId="0" pageNumber="120">
<emphasis id="F28DEABC20104B22FCB9A1E0FC7CF99E" bold="true" box="[863,992,1524,1548]" pageId="0" pageNumber="120">Etymology.</emphasis>
The new species is named in memory of the late palaeoentomologist Daniil S. Aristov.
</paragraph>
</subSubSection>
<subSubSection id="88E3652520104B22FCB9A228FB99F8E5" pageId="0" pageNumber="120" type="diagnosis">
<paragraph id="C04636AE20104B22FCB9A228FB62F90F" blockId="0.[810,1452,1451,2092]" pageId="0" pageNumber="120">
<emphasis id="F28DEABC20104B22FCB9A228FC48F9C6" bold="true" box="[863,980,1596,1620]" pageId="0" pageNumber="120">Diagnosis.</emphasis>
The fore wing of
<taxonomicName id="07F94D2D20104B22FB79A229FA8DF9C6" authority="SUKATSHEVA &amp; SINITSHENKOVA, 2023" authorityName="SUKATSHEVA &amp; SINITSHENKOVA" authorityYear="2023" box="[1183,1297,1597,1620]" class="Insecta" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="0" pageNumber="120" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
<emphasis id="F28DEABC20104B22FB79A229FA8DF9C6" box="[1183,1297,1597,1620]" italics="true" pageId="0" pageNumber="120">K. aristovi</emphasis>
</taxonomicName>
<emphasis id="F28DEABC20104B22FAFCA228FAEEF9C6" bold="true" box="[1306,1394,1596,1620]" pageId="0" pageNumber="120">
<taxonomicNameLabel id="E9BE57C720104B22FAFCA228FAEEF9C6" box="[1306,1394,1596,1620]" pageId="0" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is of medium size, RS and M branching almost at the same level, the fork on R is absent, RS seven-branched.
</paragraph>
<paragraph id="C04636AE20104B22FCB9A2BDFB99F8E5" blockId="0.[810,1452,1451,2092]" pageId="0" pageNumber="120">
<emphasis id="F28DEABC20104B22FCB9A2BDFBFEF953" bold="true" box="[863,1122,1705,1729]" pageId="0" pageNumber="120">Locality and horizon.</emphasis>
The Kostovaty site, Middle Urals,
<collectingRegion id="023DF84C20104B22FC90A2D9FC79F977" box="[886,997,1741,1765]" country="Russia" name="Udmurtiya" pageId="0" pageNumber="120">Udmurtya</collectingRegion>
, right bank of the Kama River,
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upstream of the Kostovaty village and
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downstream of the Galevo pier. The deposits of Ilyinskaya Member in Lower Urzhum Formation of Urzhum Horizon are dated to the Middle Permian.
</paragraph>
</subSubSection>
<subSubSection id="88E3652520104B23FCB9A397FEC7F871" lastPageId="1" lastPageNumber="121" pageId="0" pageNumber="120" type="description">
<paragraph id="C04636AE20104B23FCB9A397FEC7F871" blockId="0.[810,1452,1451,2092]" lastBlockId="1.[136,778,1414,2092]" lastPageId="1" lastPageNumber="121" pageId="0" pageNumber="120">
<emphasis id="F28DEABC20104B22FCB9A397FC76F809" bold="true" box="[863,1002,1923,1947]" pageId="0" pageNumber="120">Description.</emphasis>
Fore wing rather narrow, length almost three times width, with greatest width at level of SC entering anterior wing margin. Anterior margin straight, apex broadly rounded. SC long, straight, ending in middle of third quarter of wing length. Costal area at wing base twice as wide as at wing apex, crossed by four crossveins. Subcostal area wide in its middle part, R straight, long, with very slight scooplike bend at apex. RS
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simple, with straight crossvein r-rs
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. RS base at beginning of second quarter of wing length, RS seven-branched. RS
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stem 2.6× as long as RS
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stem. F
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three-branched, F
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four-branched. Additional fork on RS
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of medium size, shorter than that on RS
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and significantly longer than that on RS 3. F 1 fork 0,6× as long as F 2 fork. Cells DC and MC open. M with five branches, forks slightly distal to RS fork almost at the middle of wing. M
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, M
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and M
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simple. M 1 bifurcation short, almost equal in length to its trunk. CuA fork of same size as M
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fork. Long arcuate cross vein m
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-cua
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present. Y-shaped vein not visible. CuP simple, short, entering wing trailing margin separately from A
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. A
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long, 0.76× as long as A
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and 2.6× so as A
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. Anal field narrow. Wing membrane with no coloration. Wing length 8.0 mm, width
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.
</paragraph>
</subSubSection>
<caption id="9486662620114B23FF6EA0EBFA87FAA9" ID-DOI="http://doi.org/10.5281/zenodo.7942956" ID-Zenodo-Dep="7942956" httpUri="https://zenodo.org/record/7942956/files/figure.png" pageId="1" pageNumber="121" startId="1.[136,235,1279,1303]" targetBox="[215,1371,189,1256]" targetPageId="1" targetType="figure">
<paragraph id="C04636AE20114B23FF6EA0EBFA87FAA9" blockId="1.[136,1452,1279,1339]" pageId="1" pageNumber="121">
<emphasis id="F28DEABC20114B23FF6EA0EBFE9BFA85" bold="true" box="[136,263,1279,1303]" pageId="1" pageNumber="121">FIGURE 1.</emphasis>
<taxonomicName id="07F94D2D20114B23FEE9A114FE64FA85" authority="SUKATSHEVA &amp; SINITSHENKOVA, 2023" authorityName="SUKATSHEVA &amp; SINITSHENKOVA" authorityYear="2023" box="[271,504,1280,1303]" class="Insecta" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="121" phylum="Arthropoda" rank="species" species="aristovi" status="sp. nov.">
<emphasis id="F28DEABC20114B23FEE9A114FE64FA85" box="[271,504,1280,1303]" italics="true" pageId="1" pageNumber="121">Kamopanorpa aristovi</emphasis>
</taxonomicName>
<emphasis id="F28DEABC20114B23FDE7A0EBFDC8FA85" bold="true" box="[513,596,1279,1303]" pageId="1" pageNumber="121">
<taxonomicNameLabel id="E9BE57C720114B23FDE7A0EBFDC8FA85" box="[513,596,1279,1303]" pageId="1" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype PIN, no 3695/141, locality Kostovaty (Middle Ural, Udmurtya, Russia; Middle Permian, Lower Urzhum Formation, Urzhum Horizon).
<emphasis id="F28DEABC20114B23FCF4A137FCB8FAA9" bold="true" box="[786,804,1315,1339]" pageId="1" pageNumber="121">A</emphasis>
, Photograph.
<emphasis id="F28DEABC20114B23FC55A137FC58FAA9" bold="true" box="[947,964,1315,1339]" pageId="1" pageNumber="121">B</emphasis>
, Fore wing. Scale bars = 2.0 mm.
</paragraph>
</caption>
<subSubSection id="88E3652520114B23FF5BA3E4FC57F90E" pageId="1" pageNumber="121" type="discussion">
<paragraph id="C04636AE20114B23FF5BA3E4FC57F90E" blockId="1.[136,778,1414,2092]" lastBlockId="1.[810,1452,1414,1692]" pageId="1" pageNumber="121">
<emphasis id="F28DEABC20114B23FF5BA3E4FEB0F79A" bold="true" box="[189,300,2032,2056]" pageId="1" pageNumber="121">Remarks.</emphasis>
According to the
<typeStatus id="1F42880C20114B23FE02A3E4FD8EF79A" box="[484,530,2032,2056]" pageId="1" pageNumber="121">type</typeStatus>
of branching of RS and M (almost at the same level), the size of the forewing, the absence of a fork on R the described new species is closest to
<taxonomicName id="07F94D2D20114B23FCADA1BFFB02FA50" authority="Novokshonov, 1993" authorityName="Novokshonov" authorityYear="1993" box="[843,1182,1450,1474]" class="Insecta" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="121" phylum="Arthropoda" rank="species" species="grossa">
<emphasis id="F28DEABC20114B23FCADA1BFFC2AFA50" box="[843,950,1451,1474]" italics="true" pageId="1" pageNumber="121">K. grossa</emphasis>
<bibRefCitation id="A4684B5F20114B23FC24A1BEFB02FA50" author="Novokshonov, V. G." box="[962,1182,1450,1474]" pageId="1" pageNumber="121" pagination="90 - 101" refId="ref2263" refString="Novokshonov, V. G. (1993) Caddis flies (Insecta, Trichoptera, Microptysmatidae). Paleontological Journal, 27 (1 A), 90 - 101." type="journal article" year="1993">Novokshonov, 1993</bibRefCitation>
</taxonomicName>
from the same locality of Kostovaty, differing from it in the much longer stems of R
<subScript id="5C7D34EB20114B23FCBDA217FCEFF983" attach="left" box="[859,883,1539,1553]" fontSize="6" pageId="1" pageNumber="121">1+4</subScript>
and M
<subScript id="5C7D34EB20114B23FC25A217FC7CF983" attach="left" box="[963,992,1539,1553]" fontSize="6" pageId="1" pageNumber="121">1 +3</subScript>
and an open DC. The new species differs from all other species of the genus
<taxonomicName id="07F94D2D20114B23FB79A20CFAA5F9BD" authorityName="Martynov" authorityYear="1928" box="[1183,1337,1560,1583]" class="Insecta" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="F28DEABC20114B23FB79A20CFAA5F9BD" box="[1183,1337,1560,1583]" italics="true" pageId="1" pageNumber="121">Kamopanorpa</emphasis>
</taxonomicName>
, including
<taxonomicName id="07F94D2D20114B23FCCCA228FC04F9C1" authorityName="Novokshonov" authorityYear="1993" box="[810,920,1596,1619]" class="Insecta" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mecoptera" pageId="1" pageNumber="121" phylum="Arthropoda" rank="species" species="grossa">
<emphasis id="F28DEABC20114B23FCCCA228FC04F9C1" box="[810,920,1596,1619]" italics="true" pageId="1" pageNumber="121">K. grossa</emphasis>
</taxonomicName>
, by its seven-branched RS (
<bibRefCitation id="A4684B5F20114B23FB09A228FA3BF9C6" author="Martynov, A. V." box="[1263,1447,1596,1620]" pageId="1" pageNumber="121" pagination="23 - 62" refId="ref2133" refString="Martynov, A. V. (1933) Permian fossil insects from the Arkhangelsk district. Pt. 1. The order Mecoptera. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 2, 23 - 62." type="journal article" year="1933">Martynov, 1933</bibRefCitation>
;
<bibRefCitation id="A4684B5F20114B23FCCCA274FC75F9EA" author="Sukatsheva, I. D." box="[810,1001,1632,1656]" pageId="1" pageNumber="121" pagination="198 - 209" refId="ref2496" refString="Sukatsheva, I. D. (1976) Caddisflies of the suborder Permotrichoptera. Paleontological Journal, 10 (2), 198 - 209." type="journal article" year="1976">Sukatsheva, 1976</bibRefCitation>
:
<bibRefCitation id="A4684B5F20114B23FC14A274FB57F9EA" author="Novokshonov, V. G." box="[1010,1227,1632,1656]" pageId="1" pageNumber="121" pagination="136 - 141" refId="ref2222" refString="Novokshonov, V. G. (1992) Caddisflies of the genus Kamopanorpa (Trichoptera, Microptysmatidae) from the Kungurian of Chekarda (Perm District). Paleontological Journal, 26 (3), 136 - 141." type="journal article" year="1992">Novokshonov, 1992</bibRefCitation>
,
<bibRefCitation id="A4684B5F20114B23FB33A274FA91F9EA" author="Novokshonov, V. G." box="[1237,1293,1632,1656]" pageId="1" pageNumber="121" pagination="90 - 101" refId="ref2263" refString="Novokshonov, V. G. (1993) Caddis flies (Insecta, Trichoptera, Microptysmatidae). Paleontological Journal, 27 (1 A), 90 - 101." type="journal article" year="1993">1993</bibRefCitation>
;
<bibRefCitation id="A4684B5F20114B23FAF1A274FA30F9EA" author="Sukatsheva, I. D. &amp; Aristov, D. S." box="[1303,1452,1632,1656]" pageId="1" pageNumber="121" pagination="374 - 381" refId="ref2687" refString="Sukatsheva, I. D. &amp; Aristov, D. S. (2017) New insects (Insecta: Trichoptera, Eoblattida) from the Lower Permian of Russia. Paleontological Journal, 51 (4), 374 - 381. https: // doi. org / 10.1134 / S 0031030117040116" type="journal article" year="2017">Sukatsheva &amp;</bibRefCitation>
Aristov, 2017).
</paragraph>
</subSubSection>
</treatment>
</document>
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