273 lines
23 KiB
XML
273 lines
23 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3788370" ID-GBIF-Dataset="a757de0f-b7ba-4ef8-9c95-24cfc4226b76" ID-GBIF-Taxon="163747475" ID-ISSN="1313–2970" ID-Zenodo-Dep="3788370" ID-ZooBank="EB891295-3333-4BB1-B7EA-F118C4D6B13A" approvalRequired="26" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="23" approvalRequired_for_treatments="2" checkinTime="1587953814852" checkinUser="plazi" docAuthor="Paulsen, M. J. & Smith, Andrew" docDate="2010" docId="AA0687ED826FFFBBFF63FBCA3A4CFC39" docLanguage="en" docName="ZK_article_2192.pdf" docOrigin="ZooKeys 43 (43)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6ed1fdb7" docStyleName="zookeys.2008.journal_article" docTitle="Chiasognathus beneshi Lacroix 1978" docType="treatment" docVersion="4" lastPageId="11" lastPageNumber="44" masterDocId="563FFF958266FFB0FFEFFFF33E69FFCA" masterDocTitle="Revision of the genus Chiasognathus Stephens of southern South America with the description of a new species (Coleoptera, Lucanidae, Lucaninae, Chiasognathini)" masterLastPageNumber="63" masterPageNumber="33" pageId="9" pageNumber="42" updateTime="1643557281858" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Chiasognathus Stephens of southern South America with the description of a new species (Coleoptera, Lucanidae, Lucaninae, Chiasognathini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Paulsen, M. J.</mods:namePart>
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<mods:affiliation>University of Nebraska State Museum, Lincoln, United States of America</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Smith, Andrew</mods:namePart>
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<mods:affiliation>,, Canada</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>43</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>43</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>63</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.43.397</mods:identifier>
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<mods:identifier type="GBIF-Dataset">a757de0f-b7ba-4ef8-9c95-24cfc4226b76</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576657</mods:identifier>
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<mods:identifier type="ZooBank">EB891295-3333-4BB1-B7EA-F118C4D6B13A</mods:identifier>
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||
</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788370" ID-GBIF-Taxon="163747475" ID-Zenodo-Dep="3788370" LSID="urn:lsid:plazi:treatment:AA0687ED826FFFBBFF63FBCA3A4CFC39" httpUri="http://treatment.plazi.org/id/AA0687ED826FFFBBFF63FBCA3A4CFC39" lastPageId="11" lastPageNumber="44" pageId="9" pageNumber="42">
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<subSubSection pageId="9" pageNumber="42" type="nomenclature">
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<paragraph blockId="9.[140,582,1081,1143]" box="[140,582,1081,1108]" pageId="9" pageNumber="42">
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<heading box="[140,582,1081,1108]" fontSize="11" level="2" pageId="9" pageNumber="42" reason="3">
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<taxonomicName authority="Lacroix, 1978" authorityName="Lacroix" authorityYear="1978" box="[140,582,1081,1108]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="beneshi">
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<emphasis bold="true" box="[140,582,1081,1108]" pageId="9" pageNumber="42">
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<emphasis bold="true" box="[140,409,1081,1108]" italics="true" pageId="9" pageNumber="42">Chiasognathus beneshi</emphasis>
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Lacroix, 1978
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph blockId="9.[140,582,1081,1143]" box="[140,294,1117,1143]" pageId="9" pageNumber="42">Figs 8, 12, 27</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42" type="description">
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<paragraph blockId="9.[140,1108,1187,1319]" pageId="9" pageNumber="42">
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<taxonomicName authority="Lacroix, 1978: 250" authorityName="Lacroix" authorityPageNumber="250" authorityYear="1978" box="[140,611,1187,1214]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="species" species="beneshi">
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<emphasis box="[140,382,1187,1213]" italics="true" pageId="9" pageNumber="42">Chiasognathus beneshi</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608811609" box="[390,611,1187,1214]" collectorName="J. P. Lacroix" country="Chile" location="Province d'Aysen" municipality="Rio Manihuales" pageId="9" pageNumber="42" specimenCount="1" typeStatus="holotype">Lacroix, 1978: 250</materialsCitation>
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</taxonomicName>
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. Type material:
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<typeStatus box="[794,902,1187,1213]" pageId="9" pageNumber="42">Holotype</typeStatus>
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reportedly deposited in the
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<collectorName box="[310,450,1223,1249]" pageId="9" pageNumber="42">J. P. Lacroix</collectorName>
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collection. Lacroix is deceased, and the Lacroix collection is not available for study and is in an unknown state of preservation. Type locality: “
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<collectingCountry box="[196,258,1293,1319]" name="Chile" pageId="9" pageNumber="42">Chili</collectingCountry>
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,
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<location LSID="urn:lsid:plazi:treatment:AA0687ED826FFFBBFF63FBCA3A4CFC39:27706020826FFFB9FEE1FAFE3FBCFAED" box="[270,469,1293,1319]" country="Chile" municipality="Rio Manihuales" name="Province d'Aysen" pageId="9" pageNumber="42">Province d’Aysen</location>
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:
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<collectingMunicipality box="[480,666,1293,1319]" pageId="9" pageNumber="42">Río Manihuales</collectingMunicipality>
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”.
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</paragraph>
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<paragraph blockId="9.[140,1108,1363,1672]" lastBlockId="10.[140,1108,167,1671]" lastPageId="10" lastPageNumber="43" pageId="9" pageNumber="42">
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<emphasis bold="true" box="[140,396,1363,1390]" pageId="9" pageNumber="42">
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<emphasis box="[140,286,1363,1389]" inLineHeading="true" pageId="9" pageNumber="42" reason="1">Description</emphasis>
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.
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<emphasis bold="true" box="[308,387,1363,1390]" italics="true" pageId="9" pageNumber="42">Length</emphasis>
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:
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</emphasis>
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21.7–28.0 mm.
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<emphasis bold="true" box="[615,702,1363,1390]" pageId="9" pageNumber="42">
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<emphasis bold="true" box="[615,694,1363,1390]" italics="true" pageId="9" pageNumber="42">Width</emphasis>
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:
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</emphasis>
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10.0–
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<quantity box="[786,903,1363,1390]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="9" pageNumber="42" unit="mm" value="12.6">12.6 mm</quantity>
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.
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<emphasis bold="true" box="[922,998,1363,1390]" pageId="9" pageNumber="42">
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<emphasis bold="true" box="[922,990,1363,1390]" italics="true" pageId="9" pageNumber="42">Color</emphasis>
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:
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</emphasis>
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Reddish brown with weak green or purple metallic reflections.
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<emphasis bold="true" box="[798,872,1398,1425]" pageId="9" pageNumber="42">
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<emphasis bold="true" box="[798,864,1398,1425]" italics="true" pageId="9" pageNumber="42">Head</emphasis>
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:
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</emphasis>
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Form subquadrate. Surface punctate; punctures fine to coarse, generally setose with short to long setae. Anterior margin of head rounded or concave, not strongly produced beyond anterior angles, lacking median nasus. Anterior angles obtusely rounded in dorsal view. Male mandibles cylindrical, 1.5–2× as long as head, externally almost straight until internally curved apex. Dentate carina beginning with basal tooth on dorsal surface, continuing along internal edge, not becoming internal; dorsal and ventral teeth present either side of apex resulting in scoop-like appearance; apex lacking patch of setae. Base of mandibles lacking ventral tooth. Female mandible externally rounded in basal third, dorsally flattened, with median internal tooth. Galeal brush moderately long, 2× longer than mentum. Antennal scape with sparse area of long setae present in males, or setae absent.
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<emphasis bold="true" box="[594,725,272,299]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[594,718,273,299]" italics="true" pageId="10" pageNumber="43">Pronotum</emphasis>
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:
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</emphasis>
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Posterior and lateral angles dentate, posterior angle more strongly so in males, or angles obsolete. Lateral margins distinct, crenulate. Dorsal surface with transverse ridge, longitudinal median furrow broad, weak depressions indicated either side of disc before middle in most specimens. Surface near margins with distinct, moderately long setae in females, glabrous in males examined.
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<emphasis bold="true" box="[440,523,448,475]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[440,516,448,475]" italics="true" pageId="10" pageNumber="43">Elytra</emphasis>
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:
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</emphasis>
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Surface weakly shiny, appearing wrinkled, densely punctate, setose; setae scale-like, often abraded except on lateral elytral declivity; females lacking distinct, scattered, longer bristle-like setae. Apex rounded, lacking spine. Epipleuron flat.
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<emphasis bold="true" box="[406,466,554,580]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[406,458,554,580]" italics="true" pageId="10" pageNumber="43">Legs</emphasis>
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:
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</emphasis>
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Protibiae short, serrate externally; ventral surface along internal margin with 2–5 small teeth.
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<emphasis bold="true" box="[591,777,588,615]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[591,769,588,615]" italics="true" pageId="10" pageNumber="43">Male genitalia</emphasis>
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:
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</emphasis>
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Flagellum moderately long, subequal in length to basal piece (Fig. 8).
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</paragraph>
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</subSubSection>
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<subSubSection box="[185,1027,659,686]" pageId="10" pageNumber="43" type="distribution">
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<paragraph blockId="10.[140,1108,167,1671]" box="[185,1027,659,686]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[185,344,659,685]" pageId="10" pageNumber="43">
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<emphasis box="[185,339,659,685]" inLineHeading="true" pageId="10" pageNumber="43" reason="1">Distribution</emphasis>
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.
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</emphasis>
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Specimens were examined only fromAisén,
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<collectingCountry box="[851,913,659,685]" name="Chile" pageId="10" pageNumber="43">Chile</collectingCountry>
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(Fig. 27).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="materials_examined">
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<paragraph blockId="10.[140,1108,167,1671]" box="[185,1088,693,722]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[185,274,694,720]" pageId="10" pageNumber="43">
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<collectingCountry box="[185,274,694,720]" name="Chile" pageId="10" pageNumber="43">CHILE</collectingCountry>
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</emphasis>
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(6):
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<emphasis bold="true" box="[330,396,693,720]" pageId="10" pageNumber="43">
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<collectingRegion box="[330,396,693,720]" country="Chile" name="Aysen" pageId="10" pageNumber="43">Aisén</collectingRegion>
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</emphasis>
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(6): Caleta Tortel, Coihaique, Lago Yulton, Río Correntoso.
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</paragraph>
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<paragraph blockId="10.[140,1108,167,1671]" box="[185,757,729,756]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[185,459,729,756]" pageId="10" pageNumber="43">
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<emphasis box="[185,299,730,756]" inLineHeading="true" pageId="10" pageNumber="43" reason="1">Temporal</emphasis>
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distribution.
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</emphasis>
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January (3), February (3).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="43" type="diagnosis">
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<paragraph blockId="10.[140,1108,167,1671]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[185,312,765,791]" pageId="10" pageNumber="43">
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<emphasis box="[185,307,765,791]" inLineHeading="true" pageId="10" pageNumber="43" reason="1">Diagnosis</emphasis>
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.
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</emphasis>
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This species, although externally nondescript (Fig. 12), can be separated from the
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<taxonomicName authorityName="Solier" authorityYear="1851" box="[314,439,800,827]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="latreillei">
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<emphasis box="[314,439,800,827]" italics="true" pageId="10" pageNumber="43">C. latreillei</emphasis>
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</taxonomicName>
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complex by the longer flagellum of the male genitalia and by its geographic distribution.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="44" pageId="10" pageNumber="43" type="discussion">
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<paragraph blockId="10.[140,1108,167,1671]" pageId="10" pageNumber="43">
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<emphasis bold="true" box="[185,298,871,897]" pageId="10" pageNumber="43">
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<emphasis box="[185,293,871,897]" inLineHeading="true" pageId="10" pageNumber="43" reason="1">Remarks</emphasis>
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.
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</emphasis>
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The amateur entomologist J. P. Lacroix described this species. Most taxa named by Lacroix, especially in the
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<taxonomicName authorityName="Burmeister" authorityYear="1847" box="[562,739,906,932]" class="Insecta" family="Lucanidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="tribe" tribe="Chiasognathini">Chiasognathini</taxonomicName>
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, are now considered synonyms, including eight of twelve available Lacroix names in
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<taxonomicName authorityName="Buquet" authorityYear="1838" box="[735,909,941,967]" class="Insecta" family="Curculionidae" genus="Sphaenognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis box="[735,909,941,967]" italics="true" pageId="10" pageNumber="43">Sphaenognathus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation author="Chalumeau F & Brochier B" pageId="10" pageNumber="43" refId="ref14133" refString="Chalumeau F, Brochier B (2007) Chiasognathinae of the Andes. Taita Publishers, Hradec Kralove, 324 pp." type="book" year="2007">Chalumeau and Brochier 2007</bibRefCitation>
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;
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<bibRefCitation author="Paulsen MJ" box="[318,475,976,1002]" pageId="10" pageNumber="43" refId="ref15209" refString="Paulsen MJ (2008) Annotated Checklist of the New World Lucanidae, Version 2. http: // www. museum. unl. edu / research / entomology / Guide / Scarabaeoidea / Lucanidae / Lucanidae-Catalog / LucanidaeC. htm [accessed: December, 2009]" type="url" year="2008">Paulsen 2008</bibRefCitation>
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). The type specimen was not deposited in an institution and is unavailable for study, but we have examined specimens from near the type locality in
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<collectingRegion box="[239,301,1047,1073]" country="Chile" name="Aysen" pageId="10" pageNumber="43">Aisén</collectingRegion>
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that display mandibles identical to those drawn for the
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<typeStatus box="[943,1042,1047,1073]" pageId="10" pageNumber="43">holotype</typeStatus>
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of
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<taxonomicName authorityName="Lacroix" authorityYear="1978" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">
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<emphasis italics="true" pageId="10" pageNumber="43">C. beneshi</emphasis>
|
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</taxonomicName>
|
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. These specimens have a distinctly longer flagellum of the male genitalia than do
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<emphasis box="[177,310,1117,1144]" italics="true" pageId="10" pageNumber="43">
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<taxonomicName authorityName="Solier" authorityYear="1851" box="[177,306,1117,1144]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="latreillei">C. latreillei</taxonomicName>
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.
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</emphasis>
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Because this character appears to be useful in separating other species in the genus, we have refrained from synonymizing
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<taxonomicName authorityName="Lacroix" authorityYear="1978" box="[732,844,1152,1179]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">
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<emphasis box="[732,844,1152,1179]" italics="true" pageId="10" pageNumber="43">C. beneshi</emphasis>
|
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</taxonomicName>
|
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at this time. However, externally there are few strong morphological characters to support the distinction, in part because the
|
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<taxonomicName authorityName="Solier" authorityYear="1851" box="[329,453,1223,1249]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="latreillei">
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<emphasis box="[329,453,1223,1249]" italics="true" pageId="10" pageNumber="43">C. latreillei</emphasis>
|
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</taxonomicName>
|
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complex is so variable and possibly composed of multiple taxa. The median teeth of the male mandibles in
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<taxonomicName authorityName="Solier" authorityYear="1851" box="[695,819,1258,1284]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="latreillei">
|
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<emphasis box="[695,819,1258,1284]" italics="true" pageId="10" pageNumber="43">C. latreillei</emphasis>
|
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</taxonomicName>
|
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generally point distinctly inward, while in the
|
||
<taxonomicName authorityName="Lacroix" authorityYear="1978" box="[375,486,1293,1320]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">
|
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<emphasis box="[375,486,1293,1320]" italics="true" pageId="10" pageNumber="43">C. beneshi</emphasis>
|
||
</taxonomicName>
|
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material studied the teeth remain more or less vertical. The apex of the male mandible in
|
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<taxonomicName authorityName="Lacroix" authorityYear="1978" box="[540,654,1328,1355]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">
|
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<emphasis box="[540,654,1328,1355]" italics="true" pageId="10" pageNumber="43">C. beneshi</emphasis>
|
||
</taxonomicName>
|
||
generally has a strong tooth above and below the apex, giving a scoop-like appearance, and the mandibles overall are more cylindrical and curved internally at the apex. Most specimens of
|
||
<taxonomicName authorityName="Solier" authorityYear="1851" box="[881,1006,1399,1425]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="latreillei">
|
||
<emphasis box="[881,1006,1399,1425]" italics="true" pageId="10" pageNumber="43">C. latreillei</emphasis>
|
||
</taxonomicName>
|
||
have externally straighter mandibles with a simple apex. Unfortunately, a few specimens of the
|
||
<taxonomicName authorityName="Solier" authorityYear="1851" box="[183,308,1469,1496]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="latreillei">
|
||
<emphasis box="[183,308,1469,1496]" italics="true" pageId="10" pageNumber="43">C. latreillei</emphasis>
|
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</taxonomicName>
|
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complex have mandibles similar to those of
|
||
<taxonomicName authorityName="Lacroix" authorityYear="1978" box="[814,926,1469,1496]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">
|
||
<emphasis box="[814,926,1469,1496]" italics="true" pageId="10" pageNumber="43">C. beneshi</emphasis>
|
||
</taxonomicName>
|
||
. The specimens treated as
|
||
<taxonomicName authorityName="Lacroix" authorityYear="1978" box="[257,371,1504,1531]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">
|
||
<emphasis box="[257,371,1504,1531]" italics="true" pageId="10" pageNumber="43">C. beneshi</emphasis>
|
||
</taxonomicName>
|
||
by some authors (
|
||
<bibRefCitation author="Mizunuma T & Nagai S" box="[586,911,1504,1530]" pageId="10" pageNumber="43" refId="ref14725" refString="Mizunuma T, Nagai S (2001) The Lucanid Beetles of the World, 4 th edition. Mushi Sha, Tokyo, 337 pp." type="book" year="2001">Mizunuma and Nagai 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Chalumeau F & Brochier B" pageId="10" pageNumber="43" refId="ref14133" refString="Chalumeau F, Brochier B (2007) Chiasognathinae of the Andes. Taita Publishers, Hradec Kralove, 324 pp." type="book" year="2007">Chalumeau and Brochier 2007</bibRefCitation>
|
||
) represent an undescribed species that we describe below.
|
||
<taxonomicName authorityName="Lacroix" authorityYear="1978" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="43" phylum="Arthropoda" rank="species" species="beneshi">
|
||
<emphasis italics="true" pageId="10" pageNumber="43">Chiasognathus beneshi</emphasis>
|
||
</taxonomicName>
|
||
was previously recorded from
|
||
<collectingRegion box="[569,645,1575,1601]" country="Chile" name="Biobio" pageId="10" pageNumber="43">Biobío</collectingRegion>
|
||
(Las Trancas) but this locality is erroneous due to the misapplication of the name – this species is only known to occur in
|
||
<collectingRegion box="[140,281,1645,1671]" country="Chile" name="Aysen" pageId="10" pageNumber="43">Aisén region</collectingRegion>
|
||
of
|
||
<collectingCountry box="[319,385,1645,1671]" name="Chile" pageId="10" pageNumber="43">Chile</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="11.[140,1061,990,1012]" box="[140,1061,990,1012]" pageId="11" pageNumber="44">
|
||
<emphasis bold="true" box="[140,299,990,1012]" pageId="11" pageNumber="44">Figures Ι2–Ι3.</emphasis>
|
||
Dorsal habitus of
|
||
<taxonomicName box="[471,600,990,1011]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[471,600,990,1011]" italics="true" pageId="11" pageNumber="44">Chiasognathus</emphasis>
|
||
</taxonomicName>
|
||
species, males.
|
||
<emphasis bold="true" box="[744,770,990,1012]" pageId="11" pageNumber="44">Ι2</emphasis>
|
||
<emphasis box="[777,875,990,1011]" italics="true" pageId="11" pageNumber="44">
|
||
<taxonomicName authorityName="Lacroix" authorityYear="1978" box="[777,871,990,1011]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="beneshi">C. beneshi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[921,947,990,1012]" pageId="11" pageNumber="44">Ι3</emphasis>
|
||
<emphasis box="[953,1061,990,1011]" italics="true" pageId="11" pageNumber="44">
|
||
<taxonomicName authorityName="Solier" authorityYear="1851" box="[953,1057,990,1011]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="latreillei">C. latreillei</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |