702 lines
97 KiB
XML
702 lines
97 KiB
XML
<document id="05BA29C4FA4AB062AFB067E329CCF7D4" ID-DOI="10.11646/zootaxa.5405.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10603574" ID-ZooBank="9DA83136-F19A-4A22-8320-EE3A09487013" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706774019317" checkinUser="plazi" docAuthor="Vrbica, Maja, Vesović, Nikola, Rađa, Tonći & Ćurčić, Srećko" docDate="2024" docId="772187F5FFE4FF94FF7DFA93FC923724" docLanguage="en" docName="zootaxa.5405.3.5.pdf" docOrigin="Zootaxa 5405 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5405.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speoplanes biocovensis subsp. bosnicus Vrbica, Vesovic, Rada & Curcic 2024, ssp. n." docType="treatment" docVersion="2" lastPageNumber="430" masterDocId="8B18FF8DFFE7FF9CFFEAFFC6FF813641" masterDocTitle="A new subspecies of the subterranean beetle Speoplanes biocovensis Müller, 1934 (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina: the first occurrence of the genus outside Croatia" masterLastPageNumber="432" masterPageNumber="422" pageNumber="425" updateTime="1706774508358" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="6B522CD661E7D57E3341D091ED925A81" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="68FCF67CEB1ACD56ADDCDA1607A68BEA">
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<mods:title id="4420549CE21E065D6A0F6C16FC07163E">A new subspecies of the subterranean beetle Speoplanes biocovensis Müller, 1934 (Coleoptera: Leiodidae: Cholevinae: Leptodirini) from Bosnia and Herzegovina: the first occurrence of the genus outside Croatia</mods:title>
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<mods:name id="85417FDE1BC882CD2FA9BB24FD571182" type="personal">
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<mods:role id="299D8BB8A2A70C62B9580B4D02E66CA1">
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<mods:roleTerm id="D4AC9295268C6BE83BFDD9D2C69CCFBC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5A2AEA7726456D9719035E041FBE034D">Vrbica, Maja</mods:namePart>
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</mods:name>
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<mods:name id="D3DDA02E289A3A4D4E8423EB68FE68B0" type="personal">
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<mods:role id="F5ED693BFF9A57080243CA9F17018079">
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<mods:roleTerm id="EE13CD9993788B323F94565F7D656A86">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="46F1EBB74513CF1FC23BDD497153DECD">Vesović, Nikola</mods:namePart>
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</mods:name>
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<mods:name id="55B99063A856FC4F59F1BB23D7D483B4" type="personal">
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<mods:roleTerm id="3DEB2CBE0451DA2566BC7B8BBA4AC236">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C2A93EFB301E3531A81BE84B1D72CA63">Rađa, Tonći</mods:namePart>
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</mods:name>
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<mods:name id="7658892E27E655DB21B51ECF93D3DD7B" type="personal">
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<mods:roleTerm id="DD49834478984A6274487F5D13B9D2B4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8C3D8B8EF0B89CC5F573F8C33C694A97">Ćurčić, Srećko</mods:namePart>
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<mods:affiliation id="59BE6C20DC7BF129BE3C453B2212ED23">Corresponding author</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="33A7A5B3ADB4B7E062B20D1D8DD76233">text</mods:typeOfResource>
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<mods:title id="26488BA28DA88AD1FD84A674BE381109">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="24A7B6FCBD0257C7AAC01164D8F475F3">
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<mods:date id="339E3AB03143162F255D7E54FC6108AA">2024</mods:date>
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<mods:detail id="D083588945119A50D36B7A9395344791" type="pubDate">
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<mods:number id="FA845C87CE4C80D1E05FD1B7CDDE38ED">2024-01-31</mods:number>
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<mods:number id="C93CF90FF4B4C060CB12846BCB11E455">5405</mods:number>
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<mods:number id="F491C27AB4BD631DC3D4A341C11C8B74">3</mods:number>
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<mods:start id="02A4CF81516C667D2117D534CCC9CC21">422</mods:start>
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<mods:url id="16E451B70F8BD964A7C402467B5F1F45">http://dx.doi.org/10.11646/zootaxa.5405.3.5</mods:url>
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<mods:classification id="1BA7AC4CDC094717C7FD93ACBD84363D">journal article</mods:classification>
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<mods:identifier id="764C70BD1C7FD82856C3DBCD7C48929A" type="DOI">10.11646/zootaxa.5405.3.5</mods:identifier>
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<mods:identifier id="0BABC5328F4B74F208C8BE11D6B5F737" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A10048FDB277CDEFEA182B3DCC6603CC" type="ZooBank">9DA83136-F19A-4A22-8320-EE3A09487013</mods:identifier>
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<treatment id="772187F5FFE4FF94FF7DFA93FC923724" ID-DOI="http://doi.org/10.5281/zenodo.10603560" ID-Zenodo-Dep="10603560" LSID="urn:lsid:plazi:treatment:772187F5FFE4FF94FF7DFA93FC923724" httpUri="http://treatment.plazi.org/id/772187F5FFE4FF94FF7DFA93FC923724" lastPageId="8" lastPageNumber="430" pageId="3" pageNumber="425">
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<subSubSection id="B7926568FFE4FF9FFF7DFA93FB95332E" box="[151,1044,1364,1395]" pageId="3" pageNumber="425" type="nomenclature">
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<paragraph id="FF3736E3FFE4FF9FFF7DFA93FB95332E" blockId="3.[151,1044,1364,1427]" box="[151,1044,1364,1395]" pageId="3" pageNumber="425">
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<heading id="A47F818FFFE4FF9FFF7DFA93FB95332E" bold="true" box="[151,1044,1364,1395]" fontSize="11" level="1" pageId="3" pageNumber="425" reason="1">
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<emphasis id="CDFCEAF1FFE4FF9FFF7DFA93FB95332E" bold="true" box="[151,1044,1364,1395]" pageId="3" pageNumber="425">
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<taxonomicName id="38884D60FFE4FF9FFF7DFA93FC383332" authority="Vrbica, Vesovic, Rada & Curcic" authorityName="Vrbica, Vesovic, Rada & Curcic" authorityYear="2024" box="[151,953,1364,1395]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="425" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
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<emphasis id="CDFCEAF1FFE4FF9FFF7DFA93FDA23331" bold="true" box="[151,547,1365,1392]" italics="true" pageId="3" pageNumber="425">Speoplanes biocovensis bosnicus</emphasis>
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||
Vrbica, Vesović, Rađa & Ćurčić
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="D6CF578AFFE4FF9FFC2CFA93FB95332E" box="[966,1044,1365,1391]" pageId="3" pageNumber="425" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B7926568FFE4FF9FFF7DFABFFE9333D2" box="[151,274,1401,1427]" pageId="3" pageNumber="425" type="description">
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<paragraph id="FF3736E3FFE4FF9FFF7DFABFFE9333D2" blockId="3.[151,1044,1364,1427]" box="[151,274,1401,1427]" pageId="3" pageNumber="425">
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(
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<figureCitation id="67B32A66FFE4FF9FFF75FABFFE8B33D2" box="[159,266,1401,1427]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="5.[151,250,1019,1043]" captionStartId-1="6.[151,250,1193,1217]" captionStartId-2="6.[151,250,1595,1619]" captionStartId-3="7.[151,250,1577,1601]" captionTargetBox-0="[329,1257,181,995]" captionTargetBox-1="[302,1285,187,1163]" captionTargetBox-2="[496,1091,1292,1558]" captionTargetBox-3="[261,1327,466,1541]" captionTargetId-0="figure-32@5.[329,1257,181,995]" captionTargetId-1="figure-20@6.[296,1291,181,1169]" captionTargetId-2="figure-50@6.[496,1091,1277,1571]" captionTargetId-3="figure-183@7.[248,1340,453,1553]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." captionText-1="FIGURE 4. Morphology of head (left), pronotum (middle) and elytra (right) of Speoplanes biocovensis bosnicus ssp. n. (dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm." captionText-2="FIGURE 5. Speoplanes biocovensis bosnicus ssp. n., mesoventral carina (lateral view). Scale = 0.5 mm." captionText-3="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi-0="http://doi.org/10.5281/zenodo.10603580" figureDoi-1="http://doi.org/10.5281/zenodo.10603582" figureDoi-2="http://doi.org/10.5281/zenodo.10603584" figureDoi-3="http://doi.org/10.5281/zenodo.10603586" httpUri-0="https://zenodo.org/record/10603580/files/figure.png" httpUri-1="https://zenodo.org/record/10603582/files/figure.png" httpUri-2="https://zenodo.org/record/10603584/files/figure.png" httpUri-3="https://zenodo.org/record/10603586/files/figure.png" pageId="3" pageNumber="425">Figs. 3–6</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B7926568FFE4FF9FFF7DFA07FCAB3096" pageId="3" pageNumber="425" type="materials_examined">
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||
<paragraph id="FF3736E3FFE4FF9FFF7DFA07FCAB3096" blockId="3.[151,1437,1473,2040]" pageId="3" pageNumber="425">
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||
<emphasis id="CDFCEAF1FFE4FF9FFF7DFA07FE06339A" bold="true" box="[151,391,1473,1499]" pageId="3" pageNumber="425">Material examined.</emphasis>
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||
<materialsCitation id="4FE03CBEFFE4FF9FFE7FFA07FC163062" collectingDate="2019-04-20" collectorName="Tonci Rada" country="Bosnia and Herzegovina" elevation="66" latitude="43.134308" location="Golubinka na Bubalove Njive Pit" longLatPrecision="1" longitude="17.574278" pageId="3" pageNumber="425" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="20338841FFE4FF9FFE7FFA07FE7C339A" box="[405,509,1473,1499]" pageId="3" pageNumber="425" type="holotype">Holotype</typeStatus>
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male labeled as follows: “
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||
<location id="FA576038FFE4FF9FFCAFFA07FB5B339A" LSID="urn:lsid:plazi:treatment:772187F5FFE4FF94FF7DFA93FC923724:FA576038FFE4FF9FFCAFFA07FB5B339A" box="[837,1242,1473,1500]" country="Bosnia and Herzegovina" latitude="43.134308" longLatPrecision="1" longitude="17.574278" name="Golubinka na Bubalove Njive Pit" pageId="3" pageNumber="425">Golubinka na Bubalove Njive Pit</location>
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||
,
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||
<geoCoordinate id="9ABC5024FFE4FF9FFB06FA07FA19339D" box="[1260,1432,1473,1500]" degrees="43" direction="north" minutes="08" orientation="latitude" pageId="3" pageNumber="425" precision="1" seconds="03.5" value="43.134308">
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||
43
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||
<superScript id="08FD9BABFFE4FF9FFAE2FA07FA8E338E" attach="none" box="[1288,1295,1473,1487]" fontSize="6" pageId="3" pageNumber="425">°</superScript>
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||
08’03.5’’ N
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||
</geoCoordinate>
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||
,
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<geoCoordinate id="9ABC5024FFE4FF9FFF7DFA23FEBF3041" box="[151,318,1509,1536]" degrees="17" direction="east" minutes="34" orientation="longitude" pageId="3" pageNumber="425" precision="1" seconds="27.4" value="17.574278">
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||
17
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<superScript id="08FD9BABFFE4FF9FFF59FA23FF3B33B2" attach="none" box="[179,186,1509,1523]" fontSize="6" pageId="3" pageNumber="425">°</superScript>
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||
34’27.4’’ E
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||
</geoCoordinate>
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||
,
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||
<elevation id="74A5D1D0FFE4FF9FFEBAFA23FE5033BE" box="[336,465,1509,1536]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="425" unit="m" value="66.0">
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||
<quantity id="38709B06FFE4FF9FFEBAFA23FE0E3041" box="[336,399,1509,1536]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="425" unit="m" value="66.0">66 m</quantity>
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a.s.l.
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||
</elevation>
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, Bijača settlement, near the town of Ljubuški, southern
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<collectingCountry id="879F7673FFE4FF9FFB9DFA23FA1933BE" box="[1143,1432,1509,1535]" name="Bosnia and Herzegovina" pageId="3" pageNumber="425">Bosnia and Herzegovina</collectingCountry>
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||
,
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<date id="8B361023FFE4FF9FFF7DF9CFFE923062" box="[151,275,1545,1571]" pageId="3" pageNumber="425" value="2019-04-20">
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||
<collectingDate id="9B72E9CBFFE4FF9FFF7DF9CFFE923062" box="[151,275,1545,1571]" pageId="3" pageNumber="425" value="2019-04-20">20.IV.2019</collectingDate>
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||
</date>
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(hand collection), leg.
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<collectorName id="527D5335FFE4FF9FFDFDF9CFFD1B3062" box="[535,666,1545,1571]" pageId="3" pageNumber="425">Tonći Rađa</collectorName>
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||
” (white label, printed)
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||
</materialsCitation>
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||
/
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||
<materialsCitation id="4FE03CBEFFE4FF9FFC44F9CFFB263006" collectionCode="IZFB" country="Bosnia and Herzegovina" determinerName="Curcic" pageId="3" pageNumber="425" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="20338841FFE4FF9FFC44F9CFFBA23062" box="[942,1059,1545,1571]" pageId="3" pageNumber="425" type="holotype">Holotypus</typeStatus>
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||
<taxonomicName id="38884D60FFE4FF9FFBC0F9CFFE7C300B" authority="Vrbica, Vesovic, Rada & Curcic" authorityName="Vrbica, Vesovic, Rada & Curcic" authorityYear="2024" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="425" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
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||
<emphasis id="CDFCEAF1FFE4FF9FFBC0F9CFFA1D3062" box="[1066,1436,1545,1571]" italics="true" pageId="3" pageNumber="425">Speoplanes biocovensis bosnicus</emphasis>
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||
Vrbica, Vesović, Rađa & Ćurčić
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||
</taxonomicName>
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||
,
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||
<emphasis id="CDFCEAF1FFE4FF9FFDECF9EBFDCE3006" bold="true" box="[518,591,1581,1607]" pageId="3" pageNumber="425">
|
||
<taxonomicNameLabel id="D6CF578AFFE4FF9FFDECF9EBFDCE3006" box="[518,591,1581,1607]" pageId="3" pageNumber="425" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
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||
<determinerName id="0AEBDA27FFE4FF9FFDBFF9E9FD21300B" box="[597,672,1583,1610]" pageId="3" pageNumber="425">Ćurčić</determinerName>
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||
det. 2023 (red label, printed) (
|
||
<collectionCode id="9999AE26FFE4FF9FFC05F9EBFBAE3006" box="[1007,1071,1581,1607]" pageId="3" pageNumber="425">IZFB</collectionCode>
|
||
) (
|
||
<figureCitation id="67B32A66FFE4FF9FFBAFF9EBFB1F3006" box="[1093,1182,1581,1607]" captionStart="FIGURE 3" captionStartId="5.[151,250,1019,1043]" captionTargetBox="[329,1257,181,995]" captionTargetId="figure-32@5.[329,1257,181,995]" captionTargetPageId="5" captionText="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="3" pageNumber="425">Fig. 3A</figureCitation>
|
||
)
|
||
</materialsCitation>
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||
.
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||
<materialsCitation id="4FE03CBEFFE4FF9FFB5AF9E8FD77302A" collectionCode="IZFB" country="Bosnia and Herzegovina" determinerName="Curcic" pageId="3" pageNumber="425" specimenCount="9" specimenCount-female="7" specimenCount-male="2" typeStatus="paratype">
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||
<typeStatus id="20338841FFE4FF9FFB5AF9E8FAA03009" box="[1200,1313,1582,1608]" pageId="3" pageNumber="425" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="E98EFD6AFFE4FF9FFAC1F9E8FA1D3006" box="[1323,1436,1581,1608]" count="2" pageId="3" pageNumber="425" type="male">two males</specimenCount>
|
||
and
|
||
<specimenCount id="E98EFD6AFFE4FF9FFF22F994FEEB302A" box="[200,362,1617,1644]" count="7" pageId="3" pageNumber="425" type="female">seven females</specimenCount>
|
||
, same data as for holotype (
|
||
<collectionCode id="9999AE26FFE4FF9FFD46F997FD6F302A" box="[684,750,1617,1643]" pageId="3" pageNumber="425">IZFB</collectionCode>
|
||
)
|
||
</materialsCitation>
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||
;
|
||
<materialsCitation id="4FE03CBEFFE4FF9FFCEEF994FCA63096" collectingDate="2019-05-12" collectionCode="IZFB" collectorName="Tonci Rada & All" country="Bosnia and Herzegovina" determinerName="Curcic" pageId="3" pageNumber="425" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
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||
<specimenCount id="E98EFD6AFFE4FF9FFCEEF994FCEA302A" box="[772,875,1617,1644]" count="1" pageId="3" pageNumber="425" type="male">one male</specimenCount>
|
||
and
|
||
<specimenCount id="E98EFD6AFFE4FF9FFC4EF997FBBE302A" box="[932,1087,1617,1643]" count="3" pageId="3" pageNumber="425" type="female">three females</specimenCount>
|
||
, same locality as for holotype,
|
||
<date id="8B361023FFE4FF9FFF7DF9B3FE8B30CE" box="[151,266,1653,1679]" pageId="3" pageNumber="425" value="2019-05-12">
|
||
<collectingDate id="9B72E9CBFFE4FF9FFF7DF9B3FE8B30CE" box="[151,266,1653,1679]" pageId="3" pageNumber="425" value="2019-05-12">12.V.2019</collectingDate>
|
||
</date>
|
||
(
|
||
<collectingMethod id="26C94EF4FFE4FF9FFEF3F9B3FEDF30CE" box="[281,350,1653,1679]" pageId="3" pageNumber="425">pitfall</collectingMethod>
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trapping with rotten meat or cheese as bait), leg.
|
||
<collectorName id="527D5335FFE4FF9FFC60F9B3FB8A30CE" box="[906,1035,1653,1679]" pageId="3" pageNumber="425">Tonći Rađa</collectorName>
|
||
(
|
||
<collectionCode id="9999AE26FFE4FF9FFBF0F9B3FBDB30CE" box="[1050,1114,1653,1679]" pageId="3" pageNumber="425">IZFB</collectionCode>
|
||
) (
|
||
<figureCitation id="67B32A66FFE4FF9FFB9BF9B3FB4830CE" box="[1137,1225,1653,1679]" captionStart="FIGURE 3" captionStartId="5.[151,250,1019,1043]" captionTargetBox="[329,1257,181,995]" captionTargetId="figure-32@5.[329,1257,181,995]" captionTargetPageId="5" captionText="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="3" pageNumber="425">Fig. 3B</figureCitation>
|
||
).
|
||
<collectorName id="527D5335FFE4FF9FFB36F9B3FA8130CE" box="[1244,1280,1653,1679]" pageId="3" pageNumber="425">All</collectorName>
|
||
<typeStatus id="20338841FFE4FF9FFAEDF9B0FAF330D1" box="[1287,1394,1654,1680]" pageId="3" pageNumber="425" type="paratype">paratypes</typeStatus>
|
||
are labeled with white, printed locality labels and with red printed labels “
|
||
<typeStatus id="20338841FFE4FF9FFC5AF95FFBA030F2" box="[944,1057,1689,1715]" pageId="3" pageNumber="425" type="paratype">Paratypus</typeStatus>
|
||
<taxonomicName id="38884D60FFE4FF9FFBC3F95FFD80309B" authority="Vrbica, Vesovic, Rada & Curcic" authorityName="Vrbica, Vesovic, Rada & Curcic" authorityYear="2024" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="425" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE4FF9FFBC3F95FFA1D30F2" box="[1065,1436,1689,1715]" italics="true" pageId="3" pageNumber="425">Speoplanes biocovensis bosnicus</emphasis>
|
||
Vrbica, Vesović, Rađa & Ćurčić
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="CDFCEAF1FFE4FF9FFDE6F97BFDD73096" bold="true" box="[524,598,1725,1751]" pageId="3" pageNumber="425">
|
||
<taxonomicNameLabel id="D6CF578AFFE4FF9FFDE6F97BFDD73096" box="[524,598,1725,1751]" pageId="3" pageNumber="425" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<determinerName id="0AEBDA27FFE4FF9FFDB7F979FD29309B" box="[605,680,1727,1754]" pageId="3" pageNumber="425">Ćurčić</determinerName>
|
||
det. 2023”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7926568FFE4FF9AFF2DF927FAC930DD" lastPageId="6" lastPageNumber="428" pageId="3" pageNumber="425" type="description">
|
||
<paragraph id="FF3736E3FFE4FF9FFF2DF927FC02315E" blockId="3.[151,1437,1473,2040]" pageId="3" pageNumber="425">
|
||
<emphasis id="CDFCEAF1FFE4FF9FFF2DF927FED830BA" bold="true" box="[199,345,1761,1787]" pageId="3" pageNumber="425">Description.</emphasis>
|
||
Large-sized leptodirine. TL M
|
||
<quantity id="38709B06FFE4FF9FFD2BF927FCA630BD" box="[705,807,1761,1788]" metricMagnitude="-3" metricUnit="m" metricValue="8.32" pageId="3" pageNumber="425" unit="mm" value="8.32">8.32 mm</quantity>
|
||
(
|
||
<quantity id="38709B06FFE4FF9FFCDCF927FC1F30BD" box="[822,926,1761,1788]" metricMagnitude="-3" metricUnit="m" metricValue="8.39" pageId="3" pageNumber="425" unit="mm" value="8.39">8.39 mm</quantity>
|
||
in males,
|
||
<quantity id="38709B06FFE4FF9FFBFFF927FBFA30BD" box="[1045,1147,1761,1788]" metricMagnitude="-3" metricUnit="m" metricValue="8.29" pageId="3" pageNumber="425" unit="mm" value="8.29">8.29 mm</quantity>
|
||
in females), R
|
||
<quantity id="38709B06FFE4FF9FFAC6F927FF423161" metricMagnitude="-3" metricUnit="m" metricValue="8.315" metricValueMax="8.6" metricValueMin="8.03" pageId="3" pageNumber="425" unit="mm" value="8.315" valueMax="8.6" valueMin="8.03">8.03–8.60 mm</quantity>
|
||
(
|
||
<quantity id="38709B06FFE4FF9FFF3BF8C3FEF73161" box="[209,374,1797,1824]" metricMagnitude="-3" metricUnit="m" metricValue="8.405" metricValueMax="8.6" metricValueMin="8.21" pageId="3" pageNumber="425" unit="mm" value="8.405" valueMax="8.6" valueMin="8.21">8.21–8.60 mm</quantity>
|
||
in males,
|
||
<quantity id="38709B06FFE4FF9FFE03F8C3FD0D3161" box="[489,652,1797,1824]" metricMagnitude="-3" metricUnit="m" metricValue="8.315" metricValueMax="8.6" metricValueMin="8.03" pageId="3" pageNumber="425" unit="mm" value="8.315" valueMax="8.6" valueMin="8.03">8.03–8.60 mm</quantity>
|
||
in females) (
|
||
<tableCitation id="B20A0358FFE4FF9FFCF4F8C3FCF3315E" box="[798,882,1797,1823]" captionStart="TABLE 1" captionStartId="4.[152,244,692,716]" captionText="TABLE 1. Linear measurements and morphometric ratios of Speoplanes biocovensis bosnicus ssp. n., S. biocovensis biocovensis and S. giganteus. The values outside the parentheses are mean values, while the values inside the parentheses are ranges." pageId="3" pageNumber="425">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE4FF9FFF2DF8EFFC3C3102" blockId="3.[151,1437,1473,2040]" box="[199,957,1833,1859]" pageId="3" pageNumber="425">
|
||
Habitus. Body elongate, elliptical, reddish-brown in colour (
|
||
<figureCitation id="67B32A66FFE4FF9FFC81F8EFFC2E3102" box="[875,943,1833,1859]" captionStart="FIGURE 3" captionStartId="5.[151,250,1019,1043]" captionTargetBox="[329,1257,181,995]" captionTargetId="figure-32@5.[329,1257,181,995]" captionTargetPageId="5" captionText="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="3" pageNumber="425">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE4FF9FFF2DF888FAFC31EE" blockId="3.[151,1437,1473,2040]" pageId="3" pageNumber="425">
|
||
Integument. Dull, barely pubescent, smooth on elytra, punctate on head and pronotum, with well-defined microsculpture (
|
||
<figureCitation id="67B32A66FFE4FF9FFEA7F8B7FE1031CA" box="[333,401,1905,1931]" captionStart="FIGURE 4" captionStartId="6.[151,250,1193,1217]" captionTargetBox="[302,1285,187,1163]" captionTargetId="figure-20@6.[296,1291,181,1169]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of head (left), pronotum (middle) and elytra (right) of Speoplanes biocovensis bosnicus ssp. n. (dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10603582" httpUri="https://zenodo.org/record/10603582/files/figure.png" pageId="3" pageNumber="425">Fig. 4</figureCitation>
|
||
). Sparsely distributed yellow erect pubescence of medium length on pronotum, elytra and part of head (frons, vertex and neck). Dense yellow recumbent pubescence only on rest of head (clypeus and genae).
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE4FF98FF2DF87FFBD834C4" blockId="3.[151,1437,1473,2040]" lastBlockId="4.[151,1437,151,645]" lastPageId="4" lastPageNumber="426" pageId="3" pageNumber="425">
|
||
Head. Longer than broad (HL/HW M 1.17, R 1.10–1.22), not narrowed at neck level, without occipital carina (
|
||
<figureCitation id="67B32A66FFE4FF9FFF75F81BFED031B6" box="[159,337,2013,2039]" captionStart="FIGURE 4" captionStartId="6.[151,250,1193,1217]" captionTargetBox="[302,1285,187,1163]" captionTargetId="figure-20@6.[296,1291,181,1169]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of head (left), pronotum (middle) and elytra (right) of Speoplanes biocovensis bosnicus ssp. n. (dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10603582" httpUri="https://zenodo.org/record/10603582/files/figure.png" pageId="3" pageNumber="425">Figs. 4A and D</figureCitation>
|
||
). Labrum slightly emarginate, with several long setae. Penultimate maxillary palpomere widened apically. Ultimate maxillary palpomere short, thin, gradually narrowing apically. Vertex with an impression. Antennae inserted shortly behind middle of head, slender, narrow, longer than body in males and females (AL/TL M 1.08, R 1.03–1.16), longer in males than in females (AL/TL M 1.14, R
|
||
<quantity id="38709B06FFE3FF98FC38FF19FBDC36B8" box="[978,1117,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.8829000000000002" metricValueMax="2.9463999999999997" metricValueMin="2.8194000000000004" pageId="4" pageNumber="426" unit="in" value="1.135" valueMax="1.16" valueMin="1.11">1.11–1.16 in</quantity>
|
||
males; M 1.05, R
|
||
<quantity id="38709B06FFE3FF98FAC6FF19FF2C375C" metricMagnitude="-2" metricUnit="m" metricValue="2.6924" metricValueMax="2.7686" metricValueMin="2.6162" pageId="4" pageNumber="426" unit="in" value="1.06" valueMax="1.09" valueMin="1.03">1.03–1.09 in</quantity>
|
||
females), AL M
|
||
<quantity id="38709B06FFE3FF98FE81FEC5FE4F375C" box="[363,462,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="8.98" pageId="4" pageNumber="426" unit="mm" value="8.98">8.98 mm</quantity>
|
||
(
|
||
<quantity id="38709B06FFE3FF98FE36FEC4FDC0375C" box="[476,577,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="9.54" pageId="4" pageNumber="426" unit="mm" value="9.54">9.54 mm</quantity>
|
||
in males,
|
||
<quantity id="38709B06FFE3FF98FD59FEC5FC96375C" box="[691,791,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="8.71" pageId="4" pageNumber="426" unit="mm" value="8.71">8.71 mm</quantity>
|
||
in females), R
|
||
<quantity id="38709B06FFE3FF98FC2AFEC5FBF0375C" box="[960,1137,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="9.205" metricValueMax="10.03" metricValueMin="8.38" pageId="4" pageNumber="426" unit="mm" value="9.205" valueMax="10.03" valueMin="8.38">8.38–10.03 mm</quantity>
|
||
(
|
||
<quantity id="38709B06FFE3FF98FB95FEC5FAB0375C" box="[1151,1329,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="9.614999999999998" metricValueMax="10.03" metricValueMin="9.2" pageId="4" pageNumber="426" unit="mm" value="9.614999999999998" valueMax="10.03" valueMin="9.2">9.20–10.03 mm</quantity>
|
||
in males,
|
||
<quantity id="38709B06FFE3FF98FF7DFEE1FEB83700" box="[151,313,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="8.88" metricValueMax="9.38" metricValueMin="8.38" pageId="4" pageNumber="426" unit="mm" value="8.88" valueMax="9.38" valueMin="8.38">8.38–9.38 mm</quantity>
|
||
in females) (
|
||
<figureCitation id="67B32A66FFE3FF98FE22FEE1FD8A3700" box="[456,523,295,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1019,1043]" captionTargetBox="[329,1257,181,995]" captionTargetId="figure-32@5.[329,1257,181,995]" captionTargetPageId="5" captionText="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="4" pageNumber="426">Fig. 3</figureCitation>
|
||
). Antennomere I short, narrow proximately, widening towards apex. Antennomere II narrow, elongate, slightly widened distally, significantly longer than antennomere I (A1L/A2L M 0.62, R 0.54– 0.68). Antennomeres II, III, and VIII of similar shape and size, narrow, slightly widened distally. Antennomeres IX and X slightly dilated distally. Antennomere I shortest (M
|
||
<quantity id="38709B06FFE3FF98FCD8FE55FC1837EC" box="[818,921,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="426" unit="mm" value="0.44">0.44 mm</quantity>
|
||
, R
|
||
<quantity id="38709B06FFE3FF98FC28FE54FBEB37EC" box="[962,1130,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="5.2" metricValueMin="4.1" pageId="4" pageNumber="426" unit="mm" value="0.46499999999999997" valueMax="0.52" valueMin="0.41">0.41–0.52 mm</quantity>
|
||
), while antennomere V (M
|
||
<quantity id="38709B06FFE3FF98FF7DFE71FF7F3790" box="[151,254,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="426" unit="mm" value="1.03">1.03 mm</quantity>
|
||
, R
|
||
<quantity id="38709B06FFE3FF98FECCFE70FE4A3790" box="[294,459,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" metricValueMax="1.15" metricValueMin="0.93" pageId="4" pageNumber="426" unit="mm" value="1.04" valueMax="1.15" valueMin="0.93">0.93–1.15 mm</quantity>
|
||
) longest in both sexes. Antennomere VIII relatively short and narrow, longer in males than in females (M
|
||
<quantity id="38709B06FFE3FF98FE91FE1DFE6237B4" box="[379,483,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="426" unit="mm" value="0.81">0.81 mm</quantity>
|
||
, R
|
||
<quantity id="38709B06FFE3FF98FDE7FE1DFD3337B4" box="[525,690,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" metricValueMax="8.8" metricValueMin="7.6" pageId="4" pageNumber="426" unit="mm" value="0.82" valueMax="0.88" valueMin="0.76">0.76–0.88 mm</quantity>
|
||
in males; M
|
||
<quantity id="38709B06FFE3FF98FCA4FE1DFC3737B4" box="[846,950,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="4" pageNumber="426" unit="mm" value="0.69">0.69 mm</quantity>
|
||
, R
|
||
<quantity id="38709B06FFE3FF98FC0AFE1CFB0437B4" box="[992,1157,474,501]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="4" pageNumber="426" unit="mm" value="0.7" valueMax="0.75" valueMin="0.65">0.65–0.75 mm</quantity>
|
||
in females), shorter and narrower than antennomeres VII and IX–XI. Ultimate antennomere slender, widening sub-distally, then narrowing apically, narrower than penultimate or as wide as it (A11W/A10W M 0.89, R 0.76–1.00). Other ratios of length of certain antennomeres: A3L/A2L M 1.04, R 0.97–1.11. A3L/A4L M 0.79, R 0.74–0.87. A6L/A3L M 1.23, R 1.15–1.30. A8L/A3L M 0.98, R 0.92–1.11. A11L/A8L M 1.13, R 0.99–1.22 (
|
||
<tableCitation id="B20A0358FFE3FF98FC1EFDADFBC934C4" box="[1012,1096,619,645]" captionStart="TABLE 1" captionStartId="4.[152,244,692,716]" captionText="TABLE 1. Linear measurements and morphometric ratios of Speoplanes biocovensis bosnicus ssp. n., S. biocovensis biocovensis and S. giganteus. The values outside the parentheses are mean values, while the values inside the parentheses are ranges." pageId="4" pageNumber="426">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ABF7666BFFE3FF98FF72FD72FE8A3554" pageId="4" pageNumber="426" startId="4.[152,244,692,716]" targetType="table">
|
||
<paragraph id="FF3736E3FFE3FF98FF72FD72FE8A3554" blockId="4.[151,1436,692,789]" pageId="4" pageNumber="426">
|
||
<emphasis id="CDFCEAF1FFE3FF98FF72FD72FE93348C" bold="true" box="[152,274,692,717]" pageId="4" pageNumber="426">TABLE 1.</emphasis>
|
||
Linear measurements and morphometric ratios of
|
||
<taxonomicName id="38884D60FFE3FF98FCD7FD72FB20348D" authority="Vrbica & Vesović & Rađa & Ćurčić, 2024" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[829,1185,692,716]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE3FF98FCD7FD72FB20348D" box="[829,1185,692,716]" italics="true" pageId="4" pageNumber="426">Speoplanes biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE3FF98FB41FD73FB72348C" bold="true" box="[1195,1267,693,717]" pageId="4" pageNumber="426">
|
||
<taxonomicNameLabel id="D6CF578AFFE3FF98FB41FD73FB72348C" box="[1195,1267,693,717]" pageId="4" pageNumber="426" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="38884D60FFE3FF98FAE9FD73FE9234B1" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="biocovensis">
|
||
<emphasis id="CDFCEAF1FFE3FF98FAE9FD73FE9234B1" italics="true" pageId="4" pageNumber="426">S. biocovensis biocovensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38884D60FFE3FF98FEACFD1FFE4934B1" baseAuthorityName="Muller" baseAuthorityYear="1911" box="[326,456,729,752]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="CDFCEAF1FFE3FF98FEACFD1FFE4934B1" box="[326,456,729,752]" italics="true" pageId="4" pageNumber="426">S. giganteus</emphasis>
|
||
</taxonomicName>
|
||
. The values outside the parentheses are mean values, while the values inside the parentheses are ranges.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF3736E3FFE3FF98FF49FCE2FAA0305E" pageId="4" pageNumber="426">
|
||
<table id="8D88C443FFE30063FF49FCE2FAC7305E" box="[163,1350,804,1567]" gridcols="4" gridrows="18" pageId="4" pageNumber="426">
|
||
<tr id="41B834A1FFE30063FF49FCE2FAC73524" box="[163,1350,804,869]" gridrow="0" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FCE2FEE83524" box="[163,361,804,869]" gridcol="0" gridrow="0" pageId="4" pageNumber="426">
|
||
<emphasis id="CDFCEAF1FFE3FF98FF49FCE2FE933524" bold="true" pageId="4" pageNumber="426">Species/subspecies Character</emphasis>
|
||
</th>
|
||
<th id="02695DDDFFE30063FE20FCE2FB8E3524" box="[458,1039,804,869]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="4" pageNumber="426">
|
||
<emphasis id="CDFCEAF1FFE3FF98FE20FCE1FD8C3522" bold="true" pageId="4" pageNumber="426">
|
||
<taxonomicName id="38884D60FFE3FF98FE20FCE1FD69357E" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[458,744,807,831]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE3FF98FE20FCE1FD69357E" bold="true" box="[458,744,807,831]" italics="true" pageId="4" pageNumber="426">S. biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicName id="38884D60FFE3FF98FD14FCFFFB8E3510" authority="Vrbica & Vesović & Rađa & Ćurčić, 2024" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[766,1039,825,849]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="ssp. n." subSpecies="biocovensis">
|
||
<emphasis id="CDFCEAF1FFE3FF98FD14FCFFFC8D3510" bold="true" box="[766,780,825,849]" italics="true" pageId="4" pageNumber="426">S</emphasis>
|
||
.
|
||
<emphasis id="CDFCEAF1FFE3FF98FCF3FCFFFB8E3510" bold="true" box="[793,1039,825,849]" italics="true" pageId="4" pageNumber="426">biocovensis biocovensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D6CF578AFFE3FF98FE20FC8DFD8C3522" box="[458,525,843,867]" pageId="4" pageNumber="426" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<th id="02695DDDFFE30063FB92FCE2FAC73524" box="[1144,1350,804,869]" gridcol="3" gridrow="0" pageId="4" pageNumber="426">
|
||
<taxonomicName id="38884D60FFE3FF98FB92FCFFFB793510" authority="TL" authorityName="TL" box="[1144,1272,825,849]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="426" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="CDFCEAF1FFE3FF98FB92FCFFFB793510" bold="true" box="[1144,1272,825,849]" italics="true" pageId="4" pageNumber="426">
|
||
S.
|
||
<emphasis id="CDFCEAF1FFE3FF98FB79FCFFFB793510" bold="true" box="[1171,1272,825,849]" italics="true" pageId="4" pageNumber="426">giganteus</emphasis>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FCBBFAC735D4" box="[163,1350,893,917]" gridrow="1" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FCBBFEE835D4" box="[163,361,893,917]" gridcol="0" gridrow="1" pageId="4" pageNumber="426">TL*</th>
|
||
<td id="02695DDDFFE30063FE20FCBBFD0135D4" box="[458,640,893,917]" gridcol="1" gridrow="1" pageId="4" pageNumber="426">8.32 (8.03–8.60)</td>
|
||
<td id="02695DDDFFE30063FD14FCBBFB8E35D4" box="[766,1039,893,917]" gridcol="2" gridrow="1" pageId="4" pageNumber="426">8.25 (7.22–8.71)</td>
|
||
<td id="02695DDDFFE30063FB92FCBBFAC735D4" box="[1144,1350,893,917]" gridcol="3" gridrow="1" pageId="4" pageNumber="426">7.84 (7.47–8.03)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FC6AFAC73585" box="[163,1350,940,964]" gridrow="2" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FC6AFEE83585" box="[163,361,940,964]" gridcol="0" gridrow="2" pageId="4" pageNumber="426">HL/HW</th>
|
||
<td id="02695DDDFFE30063FE20FC6AFD013585" box="[458,640,940,964]" gridcol="1" gridrow="2" pageId="4" pageNumber="426">1.17 (1.10–1.22)</td>
|
||
<td id="02695DDDFFE30063FD14FC6AFB8E3585" box="[766,1039,940,964]" gridcol="2" gridrow="2" pageId="4" pageNumber="426">1.08 (0.99–1.17)</td>
|
||
<td id="02695DDDFFE30063FB92FC6AFAC73585" box="[1144,1350,940,964]" gridcol="3" gridrow="2" pageId="4" pageNumber="426">1.18 (1.09–1.24)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FC12FAC735AD" box="[163,1350,980,1004]" gridrow="3" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FC12FEE835AD" box="[163,361,980,1004]" gridcol="0" gridrow="3" pageId="4" pageNumber="426">HW/PW</th>
|
||
<td id="02695DDDFFE30063FE20FC12FD0135AD" box="[458,640,980,1004]" gridcol="1" gridrow="3" pageId="4" pageNumber="426">0.69 (0.64–0.73)</td>
|
||
<td id="02695DDDFFE30063FD14FC12FB8E35AD" box="[766,1039,980,1004]" gridcol="2" gridrow="3" pageId="4" pageNumber="426">0.72 (0.70–0.77)</td>
|
||
<td id="02695DDDFFE30063FB92FC12FAC735AD" box="[1144,1350,980,1004]" gridcol="3" gridrow="3" pageId="4" pageNumber="426">0.74 (0.72–0.75)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FC3AFAC73255" box="[163,1350,1020,1044]" gridrow="4" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FC3AFEE83255" box="[163,361,1020,1044]" gridcol="0" gridrow="4" pageId="4" pageNumber="426">AL*</th>
|
||
<td id="02695DDDFFE30063FE20FC3AFD013255" box="[458,640,1020,1044]" gridcol="1" gridrow="4" pageId="4" pageNumber="426">8.98 (8.38–10.03)</td>
|
||
<td id="02695DDDFFE30063FD14FC3AFB8E3255" box="[766,1039,1020,1044]" gridcol="2" gridrow="4" pageId="4" pageNumber="426">9.50 (8.66–10.19)</td>
|
||
<td id="02695DDDFFE30063FB92FC3AFAC73255" box="[1144,1350,1020,1044]" gridcol="3" gridrow="4" pageId="4" pageNumber="426">10.79 (10.42–11.15)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FBE2FAC7327C" box="[163,1350,1060,1085]" gridrow="5" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FBE2FEE8327C" box="[163,361,1060,1085]" gridcol="0" gridrow="5" pageId="4" pageNumber="426">AL/TL</th>
|
||
<td id="02695DDDFFE30063FE20FBE2FD01327C" box="[458,640,1060,1085]" gridcol="1" gridrow="5" pageId="4" pageNumber="426">1.08 (1.03–1.16)</td>
|
||
<td id="02695DDDFFE30063FD14FBE2FB8E327C" box="[766,1039,1060,1085]" gridcol="2" gridrow="5" pageId="4" pageNumber="426">1.13 (1.00–1.20)</td>
|
||
<td id="02695DDDFFE30063FB92FBE2FAC7327C" box="[1144,1350,1060,1085]" gridcol="3" gridrow="5" pageId="4" pageNumber="426">1.31 (1.30–1.32)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FB8BFAC73224" box="[163,1350,1101,1125]" gridrow="6" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FB8BFEE83224" box="[163,361,1101,1125]" gridcol="0" gridrow="6" pageId="4" pageNumber="426">A1L/A2L</th>
|
||
<td id="02695DDDFFE30063FE20FB8BFD013224" box="[458,640,1101,1125]" gridcol="1" gridrow="6" pageId="4" pageNumber="426">0.62 (0.54–0.68)</td>
|
||
<td id="02695DDDFFE30063FD14FB8BFB8E3224" box="[766,1039,1101,1125]" gridcol="2" gridrow="6" pageId="4" pageNumber="426">0.63 (0.60–0.69)</td>
|
||
<td id="02695DDDFFE30063FB92FB8BFAC73224" box="[1144,1350,1101,1125]" gridcol="3" gridrow="6" pageId="4" pageNumber="426">0.61 (0.53–0.67)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FBB3FAC732CC" box="[163,1350,1141,1165]" gridrow="7" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FBB3FEE832CC" box="[163,361,1141,1165]" gridcol="0" gridrow="7" pageId="4" pageNumber="426">A3L/A2L</th>
|
||
<td id="02695DDDFFE30063FE20FBB3FD0132CC" box="[458,640,1141,1165]" gridcol="1" gridrow="7" pageId="4" pageNumber="426">1.04 (0.97–1.11)</td>
|
||
<td id="02695DDDFFE30063FD14FBB3FB8E32CC" box="[766,1039,1141,1165]" gridcol="2" gridrow="7" pageId="4" pageNumber="426">1.09 (1.00–1.20)</td>
|
||
<td id="02695DDDFFE30063FB92FBB3FAC732CC" box="[1144,1350,1141,1165]" gridcol="3" gridrow="7" pageId="4" pageNumber="426">1.17 (1.14–1.20)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FB5BFAC732F4" box="[163,1350,1181,1205]" gridrow="8" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FB5BFEE832F4" box="[163,361,1181,1205]" gridcol="0" gridrow="8" pageId="4" pageNumber="426">A3L/A4L</th>
|
||
<td id="02695DDDFFE30063FE20FB5BFD0132F4" box="[458,640,1181,1205]" gridcol="1" gridrow="8" pageId="4" pageNumber="426">0.79 (0.74–0.87)</td>
|
||
<td id="02695DDDFFE30063FD14FB5BFB8E32F4" box="[766,1039,1181,1205]" gridcol="2" gridrow="8" pageId="4" pageNumber="426">0.78 (0.69–0.81)</td>
|
||
<td id="02695DDDFFE30063FB92FB5BFAC732F4" box="[1144,1350,1181,1205]" gridcol="3" gridrow="8" pageId="4" pageNumber="426">0.82 (0.81–0.83)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FB03FAC7329F" box="[163,1350,1221,1246]" gridrow="9" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FB03FEE8329F" box="[163,361,1221,1246]" gridcol="0" gridrow="9" pageId="4" pageNumber="426">A6L/A3L</th>
|
||
<td id="02695DDDFFE30063FE20FB03FD01329F" box="[458,640,1221,1246]" gridcol="1" gridrow="9" pageId="4" pageNumber="426">1.23 (1.15–1.30)</td>
|
||
<td id="02695DDDFFE30063FD14FB03FB8E329F" box="[766,1039,1221,1246]" gridcol="2" gridrow="9" pageId="4" pageNumber="426">1.27 (1.23–1.29)</td>
|
||
<td id="02695DDDFFE30063FB92FB03FAC7329F" box="[1144,1350,1221,1246]" gridcol="3" gridrow="9" pageId="4" pageNumber="426">1.24 (1.22–1.27)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FB2BFAC73347" box="[163,1350,1261,1286]" gridrow="10" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FB2BFEE83347" box="[163,361,1261,1286]" gridcol="0" gridrow="10" pageId="4" pageNumber="426">A8L/A3L</th>
|
||
<td id="02695DDDFFE30063FE20FB2BFD013347" box="[458,640,1261,1286]" gridcol="1" gridrow="10" pageId="4" pageNumber="426">0.98 (0.92–1.11)</td>
|
||
<td id="02695DDDFFE30063FD14FB2BFB8E3347" box="[766,1039,1261,1286]" gridcol="2" gridrow="10" pageId="4" pageNumber="426">1.03 (0.99–1.05)</td>
|
||
<td id="02695DDDFFE30063FB92FB2BFAC73347" box="[1144,1350,1261,1286]" gridcol="3" gridrow="10" pageId="4" pageNumber="426">1.04 (1.01–1.07)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FAD0FAC7336F" box="[163,1350,1302,1326]" gridrow="11" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FAD0FEE8336F" box="[163,361,1302,1326]" gridcol="0" gridrow="11" pageId="4" pageNumber="426">A11L/A8L</th>
|
||
<td id="02695DDDFFE30063FE20FAD0FD01336F" box="[458,640,1302,1326]" gridcol="1" gridrow="11" pageId="4" pageNumber="426">1.13 (0.99–1.22)</td>
|
||
<td id="02695DDDFFE30063FD14FAD0FB8E336F" box="[766,1039,1302,1326]" gridcol="2" gridrow="11" pageId="4" pageNumber="426">1.02 (0.93–1.19)</td>
|
||
<td id="02695DDDFFE30063FB92FAD0FAC7336F" box="[1144,1350,1302,1326]" gridcol="3" gridrow="11" pageId="4" pageNumber="426">0.91 (0.87–0.96)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FAF8FAC73317" box="[163,1350,1342,1366]" gridrow="12" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FAF8FEE83317" box="[163,361,1342,1366]" gridcol="0" gridrow="12" pageId="4" pageNumber="426">A11W/A10W</th>
|
||
<td id="02695DDDFFE30063FE20FAF8FD013317" box="[458,640,1342,1366]" gridcol="1" gridrow="12" pageId="4" pageNumber="426">0.89 (0.76–1.00)</td>
|
||
<td id="02695DDDFFE30063FD14FAF8FB8E3317" box="[766,1039,1342,1366]" gridcol="2" gridrow="12" pageId="4" pageNumber="426">0.98 (0.93–1.00)</td>
|
||
<td id="02695DDDFFE30063FB92FAF8FAC73317" box="[1144,1350,1342,1366]" gridcol="3" gridrow="12" pageId="4" pageNumber="426">1.05 (0.80–1.30)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FAA0FAC7333E" box="[163,1350,1382,1407]" gridrow="13" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FAA0FEE8333E" box="[163,361,1382,1407]" gridcol="0" gridrow="13" pageId="4" pageNumber="426">PL/PW</th>
|
||
<td id="02695DDDFFE30063FE20FAA0FD01333E" box="[458,640,1382,1407]" gridcol="1" gridrow="13" pageId="4" pageNumber="426">0.99 (0.95–1.03)</td>
|
||
<td id="02695DDDFFE30063FD14FAA0FB8E333E" box="[766,1039,1382,1407]" gridcol="2" gridrow="13" pageId="4" pageNumber="426">1.07 (1.01–1.17)</td>
|
||
<td id="02695DDDFFE30063FB92FAA0FAC7333E" box="[1144,1350,1382,1407]" gridcol="3" gridrow="13" pageId="4" pageNumber="426">1.03 (1.00–1.05)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FA48FAC733E6" box="[163,1350,1422,1447]" gridrow="14" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FA48FEE833E6" box="[163,361,1422,1447]" gridcol="0" gridrow="14" pageId="4" pageNumber="426">PL/HL</th>
|
||
<td id="02695DDDFFE30063FE20FA48FD0133E6" box="[458,640,1422,1447]" gridcol="1" gridrow="14" pageId="4" pageNumber="426">1.24 (1.16–1.29)</td>
|
||
<td id="02695DDDFFE30063FD14FA48FB8E33E6" box="[766,1039,1422,1447]" gridcol="2" gridrow="14" pageId="4" pageNumber="426">1.35 (1.26–1.41)</td>
|
||
<td id="02695DDDFFE30063FB92FA48FAC733E6" box="[1144,1350,1422,1447]" gridcol="3" gridrow="14" pageId="4" pageNumber="426">1.20 (1.09–1.27)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FA71FAC7338E" box="[163,1350,1463,1487]" gridrow="15" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FA71FEE8338E" box="[163,361,1463,1487]" gridcol="0" gridrow="15" pageId="4" pageNumber="426">PB/AM</th>
|
||
<td id="02695DDDFFE30063FE20FA71FD01338E" box="[458,640,1463,1487]" gridcol="1" gridrow="15" pageId="4" pageNumber="426">1.45 (1.34–1.51)</td>
|
||
<td id="02695DDDFFE30063FD14FA71FB8E338E" box="[766,1039,1463,1487]" gridcol="2" gridrow="15" pageId="4" pageNumber="426">1.40 (1.31–1.47)</td>
|
||
<td id="02695DDDFFE30063FB92FA71FAC7338E" box="[1144,1350,1463,1487]" gridcol="3" gridrow="15" pageId="4" pageNumber="426">1.38 (1.35–1.45)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49FA19FAC733B6" box="[163,1350,1503,1527]" gridrow="16" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49FA19FEE833B6" box="[163,361,1503,1527]" gridcol="0" gridrow="16" pageId="4" pageNumber="426">EW/PW</th>
|
||
<td id="02695DDDFFE30063FE20FA19FD0133B6" box="[458,640,1503,1527]" gridcol="1" gridrow="16" pageId="4" pageNumber="426">1.66 (1.54–1.76)</td>
|
||
<td id="02695DDDFFE30063FD14FA19FB8E33B6" box="[766,1039,1503,1527]" gridcol="2" gridrow="16" pageId="4" pageNumber="426">1.66 (1.58–1.72)</td>
|
||
<td id="02695DDDFFE30063FB92FA19FAC733B6" box="[1144,1350,1503,1527]" gridcol="3" gridrow="16" pageId="4" pageNumber="426">1.69 (1.61–1.73)</td>
|
||
</tr>
|
||
<tr id="41B834A1FFE30063FF49F9C1FAC7305E" box="[163,1350,1543,1567]" gridrow="17" pageId="4" pageNumber="426">
|
||
<th id="02695DDDFFE30063FF49F9C1FEE8305E" box="[163,361,1543,1567]" gridcol="0" gridrow="17" pageId="4" pageNumber="426">EL/EW in males</th>
|
||
<td id="02695DDDFFE30063FE20F9C1FD01305E" box="[458,640,1543,1567]" gridcol="1" gridrow="17" pageId="4" pageNumber="426">2.04 (2.00–2.09)</td>
|
||
<td id="02695DDDFFE30063FD14F9C1FB8E305E" box="[766,1039,1543,1567]" gridcol="2" gridrow="17" pageId="4" pageNumber="426">2.16 (2.11–2.20)</td>
|
||
<td id="02695DDDFFE30063FB92F9C1FAC7305E" box="[1144,1350,1543,1567]" gridcol="3" gridrow="17" pageId="4" pageNumber="426">1.97 (1.86–2.13)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE3FF98FF7DF9EBFECC3005" blockId="4.[151,333,1581,1605]" box="[151,333,1581,1605]" pageId="4" pageNumber="426">
|
||
<tableNote id="026E376DFFE3FF98FF7DF9EBFECC3005" box="[151,333,1581,1605]" pageId="4" pageNumber="426" targetBox="[163,1350,804,1567]" targetPageId="4">*—values in mm.</tableNote>
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE3FF98FF2DF9B5FD8E3124" blockId="4.[151,1437,1651,2037]" pageId="4" pageNumber="426">
|
||
Prothorax. Pronotum sub-bell-shaped, about as long as broad, PL/PW M 0.99, R 0.95–1.03 (M 1.00, R 0.99– 1.02, in males; M 0.99, R
|
||
<quantity id="38709B06FFE3FF98FE51F951FDC930F0" box="[443,584,1687,1714]" metricMagnitude="-2" metricUnit="m" metricValue="2.5146" metricValueMax="2.6162" metricValueMin="2.413" pageId="4" pageNumber="426" unit="in" value="0.99" valueMax="1.03" valueMin="0.95">0.95–1.03 in</quantity>
|
||
females). PL/HL M 1.24, R 1.16–1.29. Lateral margins S-shaped. Widest at posterior angles in males and slightly anterior to middle in females, wider than head (HW/PW M 0.69, R 0.64–0.73) (
|
||
<figureCitation id="67B32A66FFE3FF98FF75F919FF6F30B8" box="[159,238,1759,1785]" captionStart="FIGURE 3" captionStartId="5.[151,250,1019,1043]" captionTargetBox="[329,1257,181,995]" captionTargetId="figure-32@5.[329,1257,181,995]" captionTargetPageId="5" captionText="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="4" pageNumber="426">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="67B32A66FFE3FF98FF11F919FEE930B8" box="[251,360,1759,1785]" captionStart="FIGURE 4" captionStartId="6.[151,250,1193,1217]" captionTargetBox="[302,1285,187,1163]" captionTargetId="figure-20@6.[296,1291,181,1169]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of head (left), pronotum (middle) and elytra (right) of Speoplanes biocovensis bosnicus ssp. n. (dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10603582" httpUri="https://zenodo.org/record/10603582/files/figure.png" pageId="4" pageNumber="426">4B and E</figureCitation>
|
||
). Pronotal base almost straight. PB/AM M 1.45, R 1.34–1.51 (M 1.45, R
|
||
<quantity id="38709B06FFE3FF98FB75F919FAAD30B8" box="[1183,1324,1759,1786]" metricMagnitude="-2" metricUnit="m" metricValue="3.7084" metricValueMax="3.8354" metricValueMin="3.5814" pageId="4" pageNumber="426" unit="in" value="1.46" valueMax="1.51" valueMin="1.41">1.41–1.51 in</quantity>
|
||
males; M 1.44, R
|
||
<quantity id="38709B06FFE3FF98FF1AF8C5FEFC315C" box="[240,381,1795,1822]" metricMagnitude="-2" metricUnit="m" metricValue="3.6195" metricValueMax="3.8354" metricValueMin="3.4036" pageId="4" pageNumber="426" unit="in" value="1.425" valueMax="1.51" valueMin="1.34">1.34–1.51 in</quantity>
|
||
females) (
|
||
<tableCitation id="B20A0358FFE3FF98FE18F8C5FDC7315C" box="[498,582,1795,1821]" captionStart="TABLE 1" captionStartId="4.[152,244,692,716]" captionText="TABLE 1. Linear measurements and morphometric ratios of Speoplanes biocovensis bosnicus ssp. n., S. biocovensis biocovensis and S. giganteus. The values outside the parentheses are mean values, while the values inside the parentheses are ranges." pageId="4" pageNumber="426">Table 1</tableCitation>
|
||
). Anterior margin barely convex medially. Anterior pronotal angles distinct, acute, pointed. Posterior pronotal angles slightly acute, almost right, not protruding posteriorly (
|
||
<figureCitation id="67B32A66FFE3FF98FB35F8E1FA113100" box="[1247,1424,1831,1857]" captionStart="FIGURE 4" captionStartId="6.[151,250,1193,1217]" captionTargetBox="[302,1285,187,1163]" captionTargetId="figure-20@6.[296,1291,181,1169]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of head (left), pronotum (middle) and elytra (right) of Speoplanes biocovensis bosnicus ssp. n. (dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10603582" httpUri="https://zenodo.org/record/10603582/files/figure.png" pageId="4" pageNumber="426">Figs. 4B and E</figureCitation>
|
||
). Pronotal disc moderately convex.
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE3FF98FF2DF8A9FF3831B4" blockId="4.[151,1437,1651,2037]" pageId="4" pageNumber="426">
|
||
Mesothorax. Mesoventral carina high anteriorly, low posteriorly (
|
||
<figureCitation id="67B32A66FFE3FF98FC51F8A9FB8331CB" box="[955,1026,1903,1930]" captionStart="FIGURE 5" captionStartId="6.[151,250,1595,1619]" captionTargetBox="[496,1091,1292,1558]" captionTargetId="figure-50@6.[496,1091,1277,1571]" captionTargetPageId="6" captionText="FIGURE 5. Speoplanes biocovensis bosnicus ssp. n., mesoventral carina (lateral view). Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603584" httpUri="https://zenodo.org/record/10603584/files/figure.png" pageId="4" pageNumber="426">Fig. 5</figureCitation>
|
||
). Anterior margin slightly concave, while posterior margin almost straight, with some setae. Tooth prominent, of variable shape (acute/right/obtuseangled), rounded. No mesoventral processus on mesoventrite (
|
||
<figureCitation id="67B32A66FFE3FF98FC8CF871FC2C3193" box="[870,941,1975,2002]" captionStart="FIGURE 5" captionStartId="6.[151,250,1595,1619]" captionTargetBox="[496,1091,1292,1558]" captionTargetId="figure-50@6.[496,1091,1277,1571]" captionTargetPageId="6" captionText="FIGURE 5. Speoplanes biocovensis bosnicus ssp. n., mesoventral carina (lateral view). Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603584" httpUri="https://zenodo.org/record/10603584/files/figure.png" pageId="4" pageNumber="426">Fig. 5</figureCitation>
|
||
). Scutellum large, triangular (
|
||
<figureCitation id="67B32A66FFE3FF98FAEDF871FF2D31B4" captionStart="FIGURE 4" captionStartId="6.[151,250,1193,1217]" captionTargetBox="[302,1285,187,1163]" captionTargetId="figure-20@6.[296,1291,181,1169]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of head (left), pronotum (middle) and elytra (right) of Speoplanes biocovensis bosnicus ssp. n. (dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10603582" httpUri="https://zenodo.org/record/10603582/files/figure.png" pageId="4" pageNumber="426">Figs. 4B and E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ABF7666BFFE2FF99FF7DFC3DFF583276" ID-DOI="http://doi.org/10.5281/zenodo.10603580" ID-Zenodo-Dep="10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="5" pageNumber="427" startId="5.[151,250,1019,1043]" targetBox="[329,1257,181,995]" targetPageId="5" targetType="figure">
|
||
<paragraph id="FF3736E3FFE2FF99FF7DFC3DFF583276" blockId="5.[151,1436,1019,1079]" pageId="5" pageNumber="427">
|
||
<emphasis id="CDFCEAF1FFE2FF99FF7DFC3DFE923252" bold="true" box="[151,275,1019,1043]" pageId="5" pageNumber="427">FIGURE 3.</emphasis>
|
||
<taxonomicName id="38884D60FFE2FF99FEF0FC3DFDEB3252" authority="Vrbica & Vesović & Rađa & Ćurčić, 2024" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[282,618,1019,1043]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="427" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE2FF99FEF0FC3DFDEB3252" box="[282,618,1019,1043]" italics="true" pageId="5" pageNumber="427">Speoplanes biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE2FF99FD9AFC3DFD323252" bold="true" box="[624,691,1019,1043]" pageId="5" pageNumber="427">
|
||
<taxonomicNameLabel id="D6CF578AFFE2FF99FD9AFC3DFD323252" box="[624,691,1019,1043]" pageId="5" pageNumber="427" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF3736E3FFE2FF99FF2DFBA2FD1B323F" blockId="5.[151,1437,1124,2038]" box="[199,666,1124,1151]" pageId="5" pageNumber="427">Metathorax. Metasternum without carina.</paragraph>
|
||
<paragraph id="FF3736E3FFE2FF99FF2DFB4EFE833337" blockId="5.[151,1437,1124,2038]" pageId="5" pageNumber="427">
|
||
Elytra. Elongate, oval, broader in females than in males, EL/EW M 1.93, R 1.78–2.09 (M 2.04, R 2.00–
|
||
<quantity id="38709B06FFE2FF99FAA5FB4EFA1D32E3" box="[1359,1436,1160,1186]" metricMagnitude="-2" metricUnit="m" metricValue="5.308599999999999" pageId="5" pageNumber="427" unit="in" value="2.09">2.09 in</quantity>
|
||
males; M 1.88, R
|
||
<quantity id="38709B06FFE2FF99FE8BFB6DFE6F3284" box="[353,494,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="4.8133" metricValueMax="5.1053999999999995" metricValueMin="4.5212" pageId="5" pageNumber="427" unit="in" value="1.895" valueMax="2.01" valueMin="1.78">1.78–2.01 in</quantity>
|
||
females), distinctly wider than pronotum, especially in females, EW/PW M 1.66, R 1.54–1.76 (M 1.57, R
|
||
<quantity id="38709B06FFE2FF99FE67FB08FD9B32A8" box="[397,538,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="3.9751" metricValueMax="4.0386" metricValueMin="3.9116" pageId="5" pageNumber="427" unit="in" value="1.565" valueMax="1.59" valueMin="1.54">1.54–1.59 in</quantity>
|
||
males; M 1.71, R
|
||
<quantity id="38709B06FFE2FF99FD0CFB09FCF332A8" box="[742,882,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="4.318" metricValueMax="4.4704" metricValueMin="4.1655999999999995" pageId="5" pageNumber="427" unit="in" value="1.7" valueMax="1.76" valueMin="1.64">1.64–1.76 in</quantity>
|
||
females) (
|
||
<tableCitation id="B20A0358FFE2FF99FC0CFB09FBB932A8" box="[998,1080,1231,1257]" captionStart="TABLE 1" captionStartId="4.[152,244,692,716]" captionText="TABLE 1. Linear measurements and morphometric ratios of Speoplanes biocovensis bosnicus ssp. n., S. biocovensis biocovensis and S. giganteus. The values outside the parentheses are mean values, while the values inside the parentheses are ranges." pageId="5" pageNumber="427">Table 1</tableCitation>
|
||
) (
|
||
<figureCitation id="67B32A66FFE2FF99FBB8FB09FB1E32A8" box="[1106,1183,1231,1257]" captionStart="FIGURE 3" captionStartId="5.[151,250,1019,1043]" captionTargetBox="[329,1257,181,995]" captionTargetId="figure-32@5.[329,1257,181,995]" captionTargetPageId="5" captionText="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="5" pageNumber="427">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="67B32A66FFE2FF99FB46FB09FA9532A8" box="[1196,1300,1231,1257]" captionStart="FIGURE 4" captionStartId="6.[151,250,1193,1217]" captionTargetBox="[302,1285,187,1163]" captionTargetId="figure-20@6.[296,1291,181,1169]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of head (left), pronotum (middle) and elytra (right) of Speoplanes biocovensis bosnicus ssp. n. (dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10603582" httpUri="https://zenodo.org/record/10603582/files/figure.png" pageId="5" pageNumber="427">4C and F</figureCitation>
|
||
). Maximum width of elytra slightly in front of middle. Lateral margins arcuate. Shoulders weakly pronounced, obtuse, covered by posterior pronotal angles. Marginal furrows not visible from above. Elytral disc distinctly convex, in lateral view steeply sloping both basally and apically. Parasutural stria absent. Elytral apex slightly scaphoid. Pygidium covered by elytra.
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE2FF99FF2DFA47FDF43045" blockId="5.[151,1437,1124,2038]" pageId="5" pageNumber="427">
|
||
Legs. Elongate and slender (
|
||
<figureCitation id="67B32A66FFE2FF99FDFFFA46FDDC33DB" box="[533,605,1408,1434]" captionStart="FIGURE 3" captionStartId="5.[151,250,1019,1043]" captionTargetBox="[329,1257,181,995]" captionTargetId="figure-32@5.[329,1257,181,995]" captionTargetPageId="5" captionText="FIGURE 3. Speoplanes biocovensis bosnicus ssp. n., habitus (dorsal view). A—holotype male; B—paratype female. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603580" httpUri="https://zenodo.org/record/10603580/files/figure.png" pageId="5" pageNumber="427">Fig. 3</figureCitation>
|
||
). Femora widened basally, narrowed in distal half. Tibiae thin, slightly curved, gradually widening distally. Each anterior tibia has a reduced distal brush-like structure. Female protarsi four-segmented. Male protarsi five-segmented, with first three segments clearly widened and flattened. Tarsal claws elongate, narrow, curved, apically pointed.
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE2FF99FF2DF9C8FDDF3126" blockId="5.[151,1437,1124,2038]" pageId="5" pageNumber="427">
|
||
Male genitalia. Aedeagus large, elongate, straight, sclerotized (
|
||
<figureCitation id="67B32A66FFE2FF99FC61F9C8FBB93069" box="[907,1080,1550,1576]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Figs. 6A and B</figureCitation>
|
||
). Basal bulb small and rounded in dorsal view (
|
||
<figureCitation id="67B32A66FFE2FF99FEAEF9F7FE1C300A" box="[324,413,1585,1611]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6A</figureCitation>
|
||
), while moderately large in lateral view (
|
||
<figureCitation id="67B32A66FFE2FF99FC8DF9F7FC41300A" box="[871,960,1585,1611]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6B</figureCitation>
|
||
). Median lobe in dorsal view straight, subparallel, slightly tapering from middle to apex, rounded anteriorly, slightly bilobed at apex, slightly shorter than parameres (
|
||
<figureCitation id="67B32A66FFE2FF99FEF0F9BEFEF530D3" box="[282,372,1656,1682]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6A</figureCitation>
|
||
). Median lobe curved in lateral view, concave dorsally around middle, with pointed apex, bent downwards sub-apically, slightly bent upwards apically (
|
||
<figureCitation id="67B32A66FFE2FF99FCCBF95AFCFD30F7" box="[801,892,1692,1718]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6B</figureCitation>
|
||
). Parameres slender, narrow, arcuate, each with moderately widened rounded apex in dorsal view (
|
||
<figureCitation id="67B32A66FFE2FF99FD27F979FCA73098" box="[717,806,1727,1753]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6A</figureCitation>
|
||
), while in lateral view they slightly narrow distally (
|
||
<figureCitation id="67B32A66FFE2FF99FA85F979FF3D30BC" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6B</figureCitation>
|
||
). Each paramere with three setae: one short apical, one short sub-apical and one longer sub-apical. Distance between apical seta and longer sub-apical seta greater than that between apical seta and shorter sub-apical seta. Longer sub-apical seta lies slightly below shorter sub-apical seta. Sclerites of inner sac thin and straight in both lateral and dorsal view (
|
||
<figureCitation id="67B32A66FFE2FF99FE4FF88BFDD13126" box="[421,592,1869,1895]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Figs. 6A and B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE2FF99FF2DF8B6FCCE31B7" blockId="5.[151,1437,1124,2038]" pageId="5" pageNumber="427">
|
||
Female genitalia. Gonostyli slender, thin, gradually narrowing distally, straight, apically pointed (
|
||
<figureCitation id="67B32A66FFE2FF99FB11F8B6FAD331CB" box="[1275,1362,1904,1930]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6C</figureCitation>
|
||
). Each gonostylus with one apical seta, three inner setae and one outer seta. Outer seta located at level between two inner setae. Spermatheca short, chitinized, basally slightly curved, almost straight and strongly constricted in middle, where somewhat less sclerotized, apically spherical (
|
||
<figureCitation id="67B32A66FFE2FF99FD0DF81AFCC331B7" box="[743,834,2012,2038]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="5" pageNumber="427">Fig. 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ABF7666BFFE1FF9AFF7DFB6FFCD732A4" ID-DOI="http://doi.org/10.5281/zenodo.10603582" ID-Zenodo-Dep="10603582" httpUri="https://zenodo.org/record/10603582/files/figure.png" pageId="6" pageNumber="428" startId="6.[151,250,1193,1217]" targetBox="[302,1285,187,1163]" targetPageId="6" targetType="figure">
|
||
<paragraph id="FF3736E3FFE1FF9AFF7DFB6FFCD732A4" blockId="6.[151,1436,1193,1254]" pageId="6" pageNumber="428">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFF7DFB6FFE933280" bold="true" box="[151,274,1193,1217]" pageId="6" pageNumber="428">FIGURE 4.</emphasis>
|
||
Morphology of head (left), pronotum (middle) and elytra (right) of
|
||
<taxonomicName id="38884D60FFE1FF9AFC51FB6FFA883280" authority="Vrbica & Vesović & Rađa & Ćurčić, 2024" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[955,1289,1193,1217]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFC51FB6FFA883280" box="[955,1289,1193,1217]" italics="true" pageId="6" pageNumber="428">Speoplanes biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE1FF9AFAE4FB6FFAD13280" bold="true" box="[1294,1360,1193,1217]" pageId="6" pageNumber="428">
|
||
<taxonomicNameLabel id="D6CF578AFFE1FF9AFAE4FB6FFAD13280" box="[1294,1360,1193,1217]" pageId="6" pageNumber="428" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(dorsal view). A–C—holotype male; D–F—paratype female. Scales = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="ABF7666BFFE1FF9AFF7DF9FDFB3E3012" ID-DOI="http://doi.org/10.5281/zenodo.10603584" ID-Zenodo-Dep="10603584" box="[151,1215,1595,1620]" httpUri="https://zenodo.org/record/10603584/files/figure.png" pageId="6" pageNumber="428" startId="6.[151,250,1595,1619]" targetBox="[496,1091,1292,1558]" targetPageId="6" targetType="figure">
|
||
<paragraph id="FF3736E3FFE1FF9AFF7DF9FDFB3E3012" blockId="6.[151,1215,1595,1620]" box="[151,1215,1595,1620]" pageId="6" pageNumber="428">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFF7DF9FDFE923015" bold="true" box="[151,275,1595,1620]" pageId="6" pageNumber="428">FIGURE 5.</emphasis>
|
||
<taxonomicName id="38884D60FFE1FF9AFEF3F9FAFDE83015" authority="Vrbica & Vesović & Rađa & Ćurčić, 2024" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[281,617,1596,1620]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFEF3F9FAFDE83015" box="[281,617,1596,1620]" italics="true" pageId="6" pageNumber="428">Speoplanes biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE1FF9AFD85F9FAFD303015" bold="true" box="[623,689,1596,1620]" pageId="6" pageNumber="428">
|
||
<taxonomicNameLabel id="D6CF578AFFE1FF9AFD85F9FAFD303015" box="[623,689,1596,1620]" pageId="6" pageNumber="428" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, mesoventral carina (lateral view). Scale = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF3736E3FFE1FF9AFF2DF944FAC930DD" blockId="6.[151,1437,1666,2016]" box="[199,1352,1666,1692]" pageId="6" pageNumber="428">
|
||
Female abdominal sternite VIII. Large, transverse, hairy, with short and thin anterior process (
|
||
<figureCitation id="67B32A66FFE1FF9AFB0EF944FABA30DD" box="[1252,1339,1666,1692]" captionStart="FIGURE 6" captionStartId="7.[151,250,1577,1601]" captionTargetBox="[261,1327,466,1541]" captionTargetId="figure-183@7.[248,1340,453,1553]" captionTargetPageId="7" captionText="FIGURE 6. Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of Speoplanes biocovensis bosnicus ssp. n. A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="6" pageNumber="428">Fig. 6E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7926568FFE1FF9BFF2DF963FE17318F" lastPageId="7" lastPageNumber="429" pageId="6" pageNumber="428" type="diagnosis">
|
||
<paragraph id="FF3736E3FFE1FF9AFF2DF963FDB9316D" blockId="6.[151,1437,1666,2016]" pageId="6" pageNumber="428">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFF2DF963FEC330FE" bold="true" box="[199,322,1701,1727]" pageId="6" pageNumber="428">Diagnosis.</emphasis>
|
||
The new subspecies is closely related to two other congeners, of which it is morphologically more similar to
|
||
<taxonomicName id="38884D60FFE1FF9AFEEDF90CFDAF30A5" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[263,558,1738,1764]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="biocovensis">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFEEDF90CFDAF30A5" box="[263,558,1738,1764]" italics="true" pageId="6" pageNumber="428">S. biocovensis biocovensis</emphasis>
|
||
</taxonomicName>
|
||
due to the shape of the sclerites of the inner sac (thin and straight) and the apex of the median lobe (flattened), as well as shorter legs and antennae and smaller AL/TL (
|
||
<tableCitation id="B20A0358FFE1FF9AFBB1F928FB2D3149" box="[1115,1196,1774,1800]" captionStart="TABLE 1" captionStartId="4.[152,244,692,716]" captionText="TABLE 1. Linear measurements and morphometric ratios of Speoplanes biocovensis bosnicus ssp. n., S. biocovensis biocovensis and S. giganteus. The values outside the parentheses are mean values, while the values inside the parentheses are ranges." pageId="6" pageNumber="428">Table 1</tableCitation>
|
||
) (
|
||
<bibRefCitation id="9B194B12FFE1FF9AFB2FF928FAD33149" author="Muller, J." box="[1221,1362,1774,1800]" pageId="6" pageNumber="428" pagination="1 - 4" refId="ref6840" refString="Muller, J. (1911) Neue Hohlenkafer aus dem osterreichischen Karst. Wiener Entomologische Zeitung, 30 (1), 1 - 4." type="journal article" year="1911">Müller 1911</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9B194B12FFE1FF9AFAB6F928FA163149" author="Muller, J." box="[1372,1431,1774,1800]" pageId="6" pageNumber="428" pagination="176 - 181" refId="ref6867" refString="Muller, J. (1934) Diagnosi preliminari di nuovi coleotteri ipogei e cavernicoli. Atti del Museo Civico di Storia Naturale di Trieste, 12, 176 - 181." type="journal article" year="1934">1934</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B194B12FFE1FF9AFF7DF8D4FEAC316D" author="Jeannel, R." box="[151,301,1810,1836]" pageId="6" pageNumber="428" pagination="1 - 436" refId="ref6742" refString="Jeannel, R. (1924) Monographie des Bathysciinae. Biospeologica V. Archives de Zoologie experimentale et generale, 63, 1 - 436." type="journal article" year="1924">Jeannel 1924</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B194B12FFE1FF9AFED3F8D4FDAD316D" author="Mizerakis, V. & Hlavac, P. & Ceplik, D. & Jalzic, B. & Faille, A." box="[313,556,1810,1836]" pageId="6" pageNumber="428" pagination="581 - 594" refId="ref6769" refString="Mizerakis, V., Hlavac, P., Ceplik, D., Jalzic, B. & Faille, A. (2022) New insights into the subterranean genus Speoplanes Muller, 1911 (Coleoptera, Leiodidae, Cholevinae) from the Dinarides. Zootaxa, 5214 (4), 581 - 594. https: // doi. org / 10.11646 / zootaxa. 5214.4.6" type="journal article" year="2022">
|
||
Mizerakis
|
||
<emphasis id="CDFCEAF1FFE1FF9AFE5AF8D5FE60316D" box="[432,481,1810,1836]" italics="true" pageId="6" pageNumber="428">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFE1FF9BFF1DF8F0FB9A37EC" blockId="6.[151,1437,1666,2016]" lastBlockId="7.[151,1437,151,429]" lastPageId="7" lastPageNumber="429" pageId="6" pageNumber="428">
|
||
Specimens of
|
||
<taxonomicName id="38884D60FFE1FF9AFE77F8F0FD2C3111" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[413,685,1846,1872]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFE77F8F0FD2C3111" box="[413,685,1846,1872]" italics="true" pageId="6" pageNumber="428">S. biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE1FF9AFD5DF8F0FC843111" bold="true" box="[695,773,1846,1872]" pageId="6" pageNumber="428">ssp. n.</emphasis>
|
||
differ from
|
||
<taxonomicName id="38884D60FFE1FF9AFC73F8F0FB4B3111" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[921,1226,1846,1872]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="biocovensis">
|
||
<emphasis id="CDFCEAF1FFE1FF9AFC73F8F0FB4B3111" box="[921,1226,1846,1872]" italics="true" pageId="6" pageNumber="428">S. biocovensis biocovensis</emphasis>
|
||
</taxonomicName>
|
||
in the size of the pro- and metatibiae in females (protibiae shorter than head and pronotum together, metatibiae shorter than elytra
|
||
<emphasis id="CDFCEAF1FFE1FF9AFF7DF8B9FF3731D9" box="[151,182,1919,1944]" italics="true" pageId="6" pageNumber="428">vs.</emphasis>
|
||
protibiae as long as head and pronotum together, metatibiae as long as elytra), PL/PW (pronotum on average as long as wide
|
||
<emphasis id="CDFCEAF1FFE1FF9AFEB8F865FEE831FD" box="[338,361,1955,1980]" italics="true" pageId="6" pageNumber="428">vs</emphasis>
|
||
. pronotum on average slightly longer than wide), position of maximum width of pronotum in females (slightly anterior to the middle
|
||
<emphasis id="CDFCEAF1FFE1FF9AFDB3F801FDF631A1" box="[601,631,1991,2016]" italics="true" pageId="6" pageNumber="428">vs.</emphasis>
|
||
at the posterior angles), shape of mesoventral carina (less pronounced, with a more prominent tooth
|
||
<emphasis id="CDFCEAF1FFE0FF9BFE0EFF51FE7A36F1" box="[484,507,151,176]" italics="true" pageId="7" pageNumber="429">vs</emphasis>
|
||
. more pronounced, with a less prominent tooth), and EL/EW in males (elytra less elongate
|
||
<emphasis id="CDFCEAF1FFE0FF9BFF14FF7DFE943695" box="[254,277,187,212]" italics="true" pageId="7" pageNumber="429">vs</emphasis>
|
||
. elytra more elongate). We also found that antennae in
|
||
<taxonomicName id="38884D60FFE0FF9BFC65FF7DFB1B3694" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[911,1178,187,213]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFC65FF7DFB1B3694" box="[911,1178,187,213]" italics="true" pageId="7" pageNumber="429">S. biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE0FF9BFB48FF7DFB6F3694" bold="true" box="[1186,1262,187,213]" pageId="7" pageNumber="429">ssp. n.</emphasis>
|
||
are on average shorter than in the nominotypic subspecies, and that antennomere VIII in the new subspecies is on average slightly shorter than antennomere III (antennomere VIII is on average slightly longer than antennomere III in
|
||
<taxonomicName id="38884D60FFE0FF9BFB14FEC5FE9B3700" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="biocovensis">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFB14FEC5FE9B3700" italics="true" pageId="7" pageNumber="429">S. biocovensis biocovensis</emphasis>
|
||
</taxonomicName>
|
||
), antennomere XI in the new subspecies is on average longer than antennomere VIII (antennomeres XI and VIII are of similar length in
|
||
<taxonomicName id="38884D60FFE0FF9BFDEFFE8DFCAE3724" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[517,815,331,357]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="biocovensis">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFDEFFE8DFCAE3724" box="[517,815,331,357]" italics="true" pageId="7" pageNumber="429">S. biocovensis biocovensis</emphasis>
|
||
</taxonomicName>
|
||
) and the pronotum in the new subspecies is on average about a quarter longer than the head (the pronotum in
|
||
<taxonomicName id="38884D60FFE0FF9BFD1FFEA9FB9F37C8" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[757,1054,367,393]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="biocovensis">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFD1FFEA9FB9F37C8" box="[757,1054,367,393]" italics="true" pageId="7" pageNumber="429">S. biocovensis biocovensis</emphasis>
|
||
</taxonomicName>
|
||
is on average about a third longer than the head) (
|
||
<tableCitation id="B20A0358FFE0FF9BFEAEFE55FE1837EC" box="[324,409,403,429]" captionStart="TABLE 1" captionStartId="4.[152,244,692,716]" captionText="TABLE 1. Linear measurements and morphometric ratios of Speoplanes biocovensis bosnicus ssp. n., S. biocovensis biocovensis and S. giganteus. The values outside the parentheses are mean values, while the values inside the parentheses are ranges." pageId="7" pageNumber="429">Table 1</tableCitation>
|
||
and
|
||
<figureCitation id="67B32A66FFE0FF9BFE25FE55FDA537EC" box="[463,548,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1223,1247]" captionTargetBox="[296,1291,273,1199]" captionTargetId="figure-40@3.[296,1291,273,1199]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of the taxa of the genus Speoplanes (dorsal view). A—S. giganteus; B—S. biocovensis biocovensis. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603578" httpUri="https://zenodo.org/record/10603578/files/figure.png" pageId="7" pageNumber="429">Fig. 2B</figureCitation>
|
||
) (
|
||
<bibRefCitation id="9B194B12FFE0FF9BFDD1FE55FD4837EC" author="Muller, J." box="[571,713,403,429]" pageId="7" pageNumber="429" pagination="1 - 4" refId="ref6840" refString="Muller, J. (1911) Neue Hohlenkafer aus dem osterreichischen Karst. Wiener Entomologische Zeitung, 30 (1), 1 - 4." type="journal article" year="1911">Müller 1911</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9B194B12FFE0FF9BFD3FFE55FC9137EC" author="Muller, J." box="[725,784,403,429]" pageId="7" pageNumber="429" pagination="176 - 181" refId="ref6867" refString="Muller, J. (1934) Diagnosi preliminari di nuovi coleotteri ipogei e cavernicoli. Atti del Museo Civico di Storia Naturale di Trieste, 12, 176 - 181." type="journal article" year="1934">1934</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B194B12FFE0FF9BFCF6FE55FB8E37EC" author="Mizerakis, V. & Hlavac, P. & Ceplik, D. & Jalzic, B. & Faille, A." box="[796,1039,403,429]" pageId="7" pageNumber="429" pagination="581 - 594" refId="ref6769" refString="Mizerakis, V., Hlavac, P., Ceplik, D., Jalzic, B. & Faille, A. (2022) New insights into the subterranean genus Speoplanes Muller, 1911 (Coleoptera, Leiodidae, Cholevinae) from the Dinarides. Zootaxa, 5214 (4), 581 - 594. https: // doi. org / 10.11646 / zootaxa. 5214.4.6" type="journal article" year="2022">
|
||
Mizerakis
|
||
<emphasis id="CDFCEAF1FFE0FF9BFC79FE55FC4537EC" box="[915,964,403,429]" italics="true" pageId="7" pageNumber="429">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ABF7666BFFE0FF9BFF7DF9EFFAC830C8" ID-DOI="http://doi.org/10.5281/zenodo.10603586" ID-Zenodo-Dep="10603586" httpUri="https://zenodo.org/record/10603586/files/figure.png" pageId="7" pageNumber="429" startId="7.[151,250,1577,1601]" targetBox="[261,1327,466,1541]" targetPageId="7" targetType="figure">
|
||
<paragraph id="FF3736E3FFE0FF9BFF7DF9EFFAC830C8" blockId="7.[151,1436,1577,1673]" pageId="7" pageNumber="429">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFF7DF9EFFE943000" bold="true" box="[151,277,1577,1601]" pageId="7" pageNumber="429">FIGURE 6.</emphasis>
|
||
Morphology of certain structures of the holotype male (A and B) and the paratype female (C–E) of
|
||
<taxonomicName id="38884D60FFE0FF9BFAC3F9EFFEF23024" authority="Vrbica & Vesović & Rađa & Ćurčić, 2024" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="biocovensis" status="subsp. nov." subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFAC3F9EFFEF23024" italics="true" pageId="7" pageNumber="429">Speoplanes biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE0FF9BFE95F98BFE493024" bold="true" box="[383,456,1613,1637]" pageId="7" pageNumber="429">
|
||
<taxonomicNameLabel id="D6CF578AFFE0FF9BFE95F98BFE493024" box="[383,456,1613,1637]" pageId="7" pageNumber="429" rank="subSpecies">ssp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A—aedeagus (dorsal view); B—aedeagus (lateral view); C—gonostyli (dorsal view); D— spermatheca (lateral view); E—abdominal sternite VIII (ventral view). Scales = 0.5 mm (A–C and E) and 0.1 mm (D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF3736E3FFE0FF9BFF2DF971FE17318F" blockId="7.[151,1437,1719,1998]" pageId="7" pageNumber="429">
|
||
Specimens of
|
||
<taxonomicName id="38884D60FFE0FF9BFE82F97EFDF03090" authorityName="Vrbica & Vesović & Rađa & Ćurčić" authorityYear="2024" box="[360,625,1719,1746]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="biocovensis" subSpecies="bosnicus">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFE82F97EFDF03090" box="[360,625,1719,1746]" italics="true" pageId="7" pageNumber="429">S. biocovensis bosnicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CDFCEAF1FFE0FF9BFD93F97EFD453093" bold="true" box="[633,708,1720,1746]" pageId="7" pageNumber="429">ssp. n.</emphasis>
|
||
differ from
|
||
<taxonomicName id="38884D60FFE0FF9BFCBAF97EFC5B3090" box="[848,986,1720,1746]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="429" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="CDFCEAF1FFE0FF9BFCBAF97EFC5B3090" box="[848,986,1720,1746]" italics="true" pageId="7" pageNumber="429">S. giganteus</emphasis>
|
||
</taxonomicName>
|
||
by the larger size (TL R
|
||
<quantity id="38709B06FFE0FF9BFB13F971FA1C3093" box="[1273,1437,1719,1746]" metricMagnitude="-3" metricUnit="m" metricValue="8.315" metricValueMax="8.6" metricValueMin="8.03" pageId="7" pageNumber="429" unit="mm" value="8.315" valueMax="8.6" valueMin="8.03">8.03–8.60 mm</quantity>
|
||
<emphasis id="CDFCEAF1FFE0FF9BFF7DF91AFF2F30B4" box="[151,174,1756,1781]" italics="true" pageId="7" pageNumber="429">vs</emphasis>
|
||
.
|
||
<quantity id="38709B06FFE0FF9BFF2AF91AFEE830B7" box="[192,361,1756,1782]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.03" metricValueMin="7.47" pageId="7" pageNumber="429" unit="mm" value="7.75" valueMax="8.03" valueMin="7.47">7.47–8.03 mm</quantity>
|
||
), HL/HW (1.10–1.22
|
||
<emphasis id="CDFCEAF1FFE0FF9BFD82F91AFD0730B4" box="[616,646,1756,1781]" italics="true" pageId="7" pageNumber="429">vs.</emphasis>
|
||
cca. 1.50), shorter antennae (AL R
|
||
<quantity id="38709B06FFE0FF9BFBC1F91AFB5E30B7" box="[1067,1247,1756,1782]" metricMagnitude="-3" metricUnit="m" metricValue="9.205" metricValueMax="10.03" metricValueMin="8.38" pageId="7" pageNumber="429" unit="mm" value="9.205" valueMax="10.03" valueMin="8.38">8.38–10.03 mm</quantity>
|
||
<emphasis id="CDFCEAF1FFE0FF9BFB03F91AFA8130B4" box="[1257,1280,1756,1781]" italics="true" pageId="7" pageNumber="429">vs</emphasis>
|
||
.
|
||
<quantity id="38709B06FFE0FF9BFAFBF91DFF46315B" metricMagnitude="-2" metricUnit="m" metricValue="1.0785" metricValueMax="1.115" metricValueMin="1.042" pageId="7" pageNumber="429" unit="mm" value="10.785" valueMax="11.15" valueMin="10.42">10.42–11.15 mm</quantity>
|
||
), smaller AL/TL (R 1.03–1.16
|
||
<emphasis id="CDFCEAF1FFE0FF9BFDC2F8C6FDBE3158" box="[552,575,1792,1817]" italics="true" pageId="7" pageNumber="429">vs</emphasis>
|
||
. 1.30–1.32), A3L/A2L (0.97–1.11
|
||
<emphasis id="CDFCEAF1FFE0FF9BFC26F8C6FC623158" box="[972,995,1792,1817]" italics="true" pageId="7" pageNumber="429">vs</emphasis>
|
||
. 1.14–1.20), A11L/A8L (0.99–1.22
|
||
<emphasis id="CDFCEAF1FFE0FF9BFA94F8C6FA143158" box="[1406,1429,1792,1817]" italics="true" pageId="7" pageNumber="429">vs</emphasis>
|
||
. 0.87–0.96), size of pro- and metatibiae in females (protibiae shorter than head and pronotum together, metatibiae shorter than elytra
|
||
<emphasis id="CDFCEAF1FFE0FF9BFE98F88EFE103120" box="[370,401,1864,1889]" italics="true" pageId="7" pageNumber="429">vs.</emphasis>
|
||
protibiae longer than head and pronotum together, metatibiae longer than elytra), PL/PW (pronotum as long as wide
|
||
<emphasis id="CDFCEAF1FFE0FF9BFE20F8AAFE6931C4" box="[458,488,1900,1925]" italics="true" pageId="7" pageNumber="429">vs.</emphasis>
|
||
pronotum significantly longer than wide), and the shape of the sclerites of the inner sac (thin and straight
|
||
<emphasis id="CDFCEAF1FFE0FF9BFE7BF856FE3131E8" box="[401,432,1936,1961]" italics="true" pageId="7" pageNumber="429">vs.</emphasis>
|
||
compact and curved) (
|
||
<tableCitation id="B20A0358FFE0FF9BFD53F849FC9131EB" box="[697,784,1935,1962]" captionStart="TABLE 1" captionStartId="4.[152,244,692,716]" captionText="TABLE 1. Linear measurements and morphometric ratios of Speoplanes biocovensis bosnicus ssp. n., S. biocovensis biocovensis and S. giganteus. The values outside the parentheses are mean values, while the values inside the parentheses are ranges." pageId="7" pageNumber="429">Table 1</tableCitation>
|
||
and
|
||
<figureCitation id="67B32A66FFE0FF9BFCA1F849FC2531EB" box="[843,932,1935,1962]" captionStart="FIGURE 2" captionStartId="3.[151,250,1223,1247]" captionTargetBox="[296,1291,273,1199]" captionTargetId="figure-40@3.[296,1291,273,1199]" captionTargetPageId="3" captionText="FIGURE 2. Habitus of the taxa of the genus Speoplanes (dorsal view). A—S. giganteus; B—S. biocovensis biocovensis. Scales = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10603578" httpUri="https://zenodo.org/record/10603578/files/figure.png" pageId="7" pageNumber="429">Fig. 2A</figureCitation>
|
||
) (
|
||
<bibRefCitation id="9B194B12FFE0FF9BFC57F849FBCF31EB" author="Muller, J." box="[957,1102,1935,1962]" pageId="7" pageNumber="429" pagination="1 - 4" refId="ref6840" refString="Muller, J. (1911) Neue Hohlenkafer aus dem osterreichischen Karst. Wiener Entomologische Zeitung, 30 (1), 1 - 4." type="journal article" year="1911">Müller 1911</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B194B12FFE0FF9BFBB6F849FB7531EB" author="Jeannel, R." box="[1116,1268,1935,1962]" pageId="7" pageNumber="429" pagination="1 - 436" refId="ref6742" refString="Jeannel, R. (1924) Monographie des Bathysciinae. Biospeologica V. Archives de Zoologie experimentale et generale, 63, 1 - 436." type="journal article" year="1924">Jeannel 1924</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B194B12FFE0FF9BFAE8F856FA1631EB" author="Pretner, E." box="[1282,1431,1936,1962]" pageId="7" pageNumber="429" pagination="101 - 239" refId="ref7087" refString="Pretner, E. (1973) Koleopteroloska fauna pecina i jama Hrvatske. Krs Jugoslavije, 8 (6), 101 - 239." type="journal article" year="1973">Pretner 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B194B12FFE0FF9BFF7DF875FE0B318F" author="Mizerakis, V. & Hlavac, P. & Ceplik, D. & Jalzic, B. & Faille, A." box="[151,394,1971,1998]" pageId="7" pageNumber="429" pagination="581 - 594" refId="ref6769" refString="Mizerakis, V., Hlavac, P., Ceplik, D., Jalzic, B. & Faille, A. (2022) New insights into the subterranean genus Speoplanes Muller, 1911 (Coleoptera, Leiodidae, Cholevinae) from the Dinarides. Zootaxa, 5214 (4), 581 - 594. https: // doi. org / 10.11646 / zootaxa. 5214.4.6" type="journal article" year="2022">
|
||
Mizerakis
|
||
<emphasis id="CDFCEAF1FFE0FF9BFEE4F872FEBE318C" box="[270,319,1971,1997]" italics="true" pageId="7" pageNumber="429">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7926568FFEFFF94FF2DFF51FC923724" pageId="8" pageNumber="430" type="etymology">
|
||
<paragraph id="FF3736E3FFEFFF94FF2DFF51FF773694" blockId="8.[151,1436,151,359]" pageId="8" pageNumber="430">
|
||
<emphasis id="CDFCEAF1FFEFFF94FF2DFF51FECF36F0" bold="true" box="[199,334,151,177]" pageId="8" pageNumber="430">Etymology.</emphasis>
|
||
After
|
||
<collectingCountry id="879F7673FFEFFF94FE48FF51FD4436F0" box="[418,709,151,177]" name="Bosnia and Herzegovina" pageId="8" pageNumber="430">Bosnia and Herzegovina</collectingCountry>
|
||
, a country where the
|
||
<typeStatus id="20338841FFEFFF94FC3FFF51FB8436F0" box="[981,1029,151,177]" pageId="8" pageNumber="430">type</typeStatus>
|
||
locality of the new subspecies is situated.
|
||
</paragraph>
|
||
<paragraph id="FF3736E3FFEFFF94FF2DFF18FC923724" blockId="8.[151,1436,151,359]" pageId="8" pageNumber="430">
|
||
<emphasis id="CDFCEAF1FFEFFF94FF2DFF18FE7336B9" bold="true" box="[199,498,222,248]" pageId="8" pageNumber="430">Geographic distribution.</emphasis>
|
||
This subspecies inhabits a pit near the town of Ljubuški in southern
|
||
<collectingCountry id="879F7673FFEFFF94FAF6FF19FEA7375C" name="Bosnia and Herzegovina" pageId="8" pageNumber="430">Bosnia and Herzegovina</collectingCountry>
|
||
—the Golubinka na Bubalove Njive Pit. This is the first discovery of the genus
|
||
<taxonomicName id="38884D60FFEFFF94FB21FEC5FACA375C" authorityName="Muller" authorityYear="1911" box="[1227,1355,259,285]" class="Insecta" family="Leiodidae" genus="Speoplanes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="430" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CDFCEAF1FFEFFF94FB21FEC5FACA375C" box="[1227,1355,259,285]" italics="true" pageId="8" pageNumber="430">Speoplanes</emphasis>
|
||
</taxonomicName>
|
||
in this country. Previously, this genus was only known from caves and pits in several mountains in southern
|
||
<collectingCountry id="879F7673FFEFFF94FAA0FEE1FA1D3700" box="[1354,1436,295,321]" name="Croatia" pageId="8" pageNumber="430">Croatia</collectingCountry>
|
||
(Dalmatia) (
|
||
<figureCitation id="67B32A66FFEFFF94FECAFE8DFEE13724" box="[288,352,331,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1198,1222]" captionTargetBox="[151,1436,273,1175]" captionTargetId="figure-78@1.[151,1436,273,1175]" captionTargetPageId="1" captionText="FIGURE 1. The distribution area of the genus Speoplanes. Light blue star—S. biocovensis bosnicus ssp. n.; light blue zones— S. biocovensis biocovensis; red zone—S. giganteus." figureDoi="http://doi.org/10.5281/zenodo.10603576" httpUri="https://zenodo.org/record/10603576/files/figure.png" pageId="8" pageNumber="430">Fig. 1</figureCitation>
|
||
) (
|
||
<bibRefCitation id="9B194B12FFEFFF94FE90FE8DFDEF3724" author="Mizerakis, V. & Hlavac, P. & Ceplik, D. & Jalzic, B. & Faille, A." box="[378,622,331,357]" pageId="8" pageNumber="430" pagination="581 - 594" refId="ref6769" refString="Mizerakis, V., Hlavac, P., Ceplik, D., Jalzic, B. & Faille, A. (2022) New insights into the subterranean genus Speoplanes Muller, 1911 (Coleoptera, Leiodidae, Cholevinae) from the Dinarides. Zootaxa, 5214 (4), 581 - 594. https: // doi. org / 10.11646 / zootaxa. 5214.4.6" type="journal article" year="2022">
|
||
Mizerakis
|
||
<emphasis id="CDFCEAF1FFEFFF94FE19FE8DFDA53724" box="[499,548,331,357]" italics="true" pageId="8" pageNumber="430">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B194B12FFEFFF94FD93FE8AFC863724" author="Ceplik, D." box="[633,775,331,359]" pageId="8" pageNumber="430" pagination="401 - 440" refId="ref6520" refString="Ceplik, D. (2023) Supplementary data and taxonomic changes in the subterranean beetles of the Balkan Peninsula from July 1 st 2016 to September 30 th 2023. Zootaxa, 5383 (4), 401 - 440. https: // doi. org / 10.11646 / zootaxa. 5383.4.1" type="journal article" year="2023">Čeplík 2023</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |