treatments-xml/data/03/F3/37/03F3372EFF93FFB6C387CDF3FE6F38B7.xml
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<document id="E5FB4CC59F917380D8B439906319E58F" ID-CLB-Dataset="32316" ID-DOI="10.11646/zootaxa.4311.3.2" ID-GBIF-Dataset="38da213d-7ad2-4afe-bc41-a9087c9e6555" ID-ISSN="1175-5326" ID-Zenodo-Dep="847854" ID-ZooBank="F6Bf844D-60Fe-4D1F-8Fca-Dfbd533F0770" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503550462087" checkinUser="plazi" docAuthor="Achterberg, Cornelis Van" docDate="2017" docId="03F3372EFF93FFB6C387CDF3FE6F38B7" docLanguage="en" docName="zootaxa.4311.3.2.pdf" docOrigin="Zootaxa 4311 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yezoceryx coronatus Achterberg 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="354" masterDocId="FFCA4F56FF94FFBFC310CC2FFF9B3A1F" masterDocTitle="First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species" masterLastPageNumber="372" masterPageNumber="345" pageNumber="352" updateTime="1698467142797" updateUser="ExternalLinkService">
<mods:mods id="FF79E300CDD1E1F4220C8D8CD816BAED" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D70FED561A7717816EB5616AC65DCCDC">First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species</mods:title>
</mods:titleInfo>
<mods:name id="D10B8B3401C05FFEBC7388BE51FE6C93" type="personal">
<mods:role id="4E6A6FD9F2DD07EA37CA36457B75ADC5">
<mods:roleTerm id="9C19A13B89C369E1BBBD1B0D3726DD50">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0C5584FCDA58C7CE231BE5CE8C5CF9A8">Achterberg, Cornelis Van</mods:namePart>
</mods:name>
<mods:typeOfResource id="1F1929FDCBBCECBAE44F33C15E63F6DC">text</mods:typeOfResource>
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<mods:titleInfo id="16FD55638CA015806D0D1788977C2907">
<mods:title id="B3059968E0F5889CA9AF0463C7874B1A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B6CE4A4768E3B7331FF1DB7B03B5BF27">
<mods:date id="62341F99DAC9F566B82722A6B7661C2A">2017</mods:date>
<mods:detail id="C3A83A8860B7EA338A984D80683AE849" type="pubDate">
<mods:number id="C70C6B8E05DDA7613E98DC058B7512E9">2017-08-24</mods:number>
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<mods:detail id="F445CE423D7677EFA68DF31547FF7D04" type="volume">
<mods:number id="F9F543B9A1E9C9AFCA67B6022DD163E7">4311</mods:number>
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<mods:number id="4A04A89CA9BC9E16EC1F5C7F60AD1D55">3</mods:number>
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<mods:classification id="F8637C46FBDB778E2104F3712044C11E">journal article</mods:classification>
<mods:identifier id="EFB5F78A83D89DF759BFC9E090FDEE38" type="CLB-Dataset">32316</mods:identifier>
<mods:identifier id="D379C0FD93233BE98C327FE36799EDE0" type="DOI">10.11646/zootaxa.4311.3.2</mods:identifier>
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<mods:identifier id="620A504B4864A9D679AA3839CFC46BDA" type="Zenodo-Dep">847854</mods:identifier>
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</mods:mods>
<treatment id="03F3372EFF93FFB6C387CDF3FE6F38B7" ID-DOI="http://doi.org/10.5281/zenodo.6051713" ID-GBIF-Taxon="132923417" ID-Zenodo-Dep="6051713" LSID="urn:lsid:plazi:treatment:03F3372EFF93FFB6C387CDF3FE6F38B7" httpUri="http://treatment.plazi.org/id/03F3372EFF93FFB6C387CDF3FE6F38B7" lastPageId="9" lastPageNumber="354" pageId="7" pageNumber="352">
<subSubSection id="C340D5B3FF93FFB8C387CDF3FE9C3808" pageId="7" pageNumber="352" type="nomenclature">
<paragraph id="8BE58638FF93FFB8C387CDF3FE693BE9" blockId="7.[151,498,476,535]" box="[151,498,476,502]" pageId="7" pageNumber="352">
<heading id="D0AD3154FF93FFB8C387CDF3FE693BE9" bold="true" box="[151,498,476,502]" fontSize="11" level="1" pageId="7" pageNumber="352" reason="1">
<emphasis id="B92E5A2AFF93FFB8C387CDF3FE693BE9" bold="true" box="[151,498,476,502]" pageId="7" pageNumber="352">
<taxonomicName id="4C5AFDBBFF93FFB8C387CDF3FE153BE9" authority="Achterberg, 2017" authorityName="Achterberg" authorityYear="2017" box="[151,398,476,502]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="coronatus" status="sp. nov.">
<emphasis id="B92E5A2AFF93FFB8C387CDF3FE153BE9" bold="true" box="[151,398,476,502]" italics="true" pageId="7" pageNumber="352">Yezoceryx coronatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21DE751FF93FFB8C285CDF3FE693BE9" box="[405,498,476,502]" pageId="7" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE58638FF93FFB8C387CDD1FE9C3808" blockId="7.[151,498,476,535]" box="[151,263,510,535]" pageId="7" pageNumber="352">
(
<figureCitation id="13619ABDFF93FFB8C38ECDD1FF643808" box="[158,255,510,535]" captionStart="FIGURE 4" captionStartId="8.[151,250,1744,1766]" captionTargetBox="[160,1420,454,1714]" captionTargetId="figure@8.[160,1420,454,1714]" captionTargetPageId="8" captionText="FIGURE 4. Yezoceryx coronatus sp. nov.: a. lateral view; b. dorsal view; c. face; d. lateral view of mesosoma; e. lateral view of first metasomal tergite; f. lateral view of hypopygium." httpUri="https://zenodo.org/record/847862/files/figure.png" pageId="7" pageNumber="352">Figure 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF93FFB8C387CE69FC9D38B8" pageId="7" pageNumber="352" type="diagnosis">
<paragraph id="8BE58638FF93FFB8C387CE69FC9D38B8" blockId="7.[151,1437,582,2011]" pageId="7" pageNumber="352">
<emphasis id="B92E5A2AFF93FFB8C387CE69FE8A3840" bold="true" box="[151,273,582,607]" pageId="7" pageNumber="352">Diagnosis.</emphasis>
Yellow with black marks; mesoscutum with black mark rather crown-shaped; face densely granulate, granulations radiating from centre; first tergite 1.6× as long as posterior width; first sternite projection blunt, hooked anteriorly; ovipositor 1.8× as long as hind tibia.
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF93FFB8C3D7CE9DFC393928" pageId="7" pageNumber="352" type="materials_examined">
<paragraph id="8BE58638FF93FFB8C3D7CE9DFC393928" blockId="7.[151,1437,582,2011]" pageId="7" pageNumber="352">
<materialsCitation id="3B328C65FF93FFB8C3D7CE9DFD4E38F0" ID-GBIF-Occurrence="1586985600" collectingDate="2007-04-09" collectingDateMax="2007-04-26" collectingDateMin="2007-04-09" collectionCode="RMNH" collectorName="Cat Tien NP &amp; Mai &amp; T. M. Nguyen" location="Crocodile" pageId="7" pageNumber="352" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai" typeStatus="holotype">
<emphasis id="B92E5A2AFF93FFB8C3D7CE9DFE2F38D4" bold="true" box="[199,436,690,715]" pageId="7" pageNumber="352">Material examined.</emphasis>
<typeStatus id="54E1389AFF93FFB8C2AECE9DFDB038D4" box="[446,555,690,715]" pageId="7" pageNumber="352" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D5C4DB1FF93FFB8C129CE9CFDD538D5" box="[569,590,691,714]" pageId="7" pageNumber="352" type="female"></specimenCount>
(
<collectionCode id="ED4B1EFDFF93FFB8C171CE9CFD2038D4" box="[609,699,691,715]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="352">RMNH</collectionCode>
),
<collectingRegion id="499E48DAFF93FFB8C1C1CE9CFCDE38D4" box="[721,837,690,715]" country="Vietnam" name="Dong Nai" pageId="7" pageNumber="352">Dong Nai</collectingRegion>
,
<collectorName id="26AFE3EEFF93FFB8C045CE9CFC7C38D5" box="[853,999,690,715]" pageId="7" pageNumber="352">Cat Tien NP</collectorName>
,
<location id="8E85D0E3FF93FFB8C0E6CE9DFBFF38D4" LSID="urn:lsid:plazi:treatment:03F3372EFF93FFB6C387CDF3FE6F38B7:8E85D0E3FF93FFB8C0E6CE9DFBFF38D4" box="[1014,1124,690,715]" name="Crocodile" pageId="7" pageNumber="352" stateProvince="Dong Nai">Crocodile</location>
trail,
<date id="FFE4A0F8FF93FFB8C7BECE9DFAD938D4" box="[1198,1346,690,715]" pageId="7" pageNumber="352" value="2007-04-09" valueMax="2007-04-26" valueMin="2007-04-09">
<collectingDate id="EFA05910FF93FFB8C7BECE9DFAD938D4" box="[1198,1346,690,715]" pageId="7" pageNumber="352" value="2007-04-09" valueMax="2007-04-26" valueMin="2007-04-09">
926.iv.
<quantity id="4CA22BDDFF93FFB8C616CE9DFAD938D4" box="[1286,1346,690,715]" metricMagnitude="6" metricUnit="m" metricValue="2.0071" pageId="7" pageNumber="352" unit="m" value="2007100.0">2007</quantity>
</collectingDate>
</date>
, 100 m a.s.l.,
<collectingMethod id="521BFE2FFF93FFB8C3C9CEF9FEF338F1" box="[217,360,726,751]" pageId="7" pageNumber="352">Malaise trap</collectingMethod>
, P. Q.
<collectorName id="26AFE3EEFF93FFB8C2A2CEF9FE4438F0" box="[434,479,726,751]" pageId="7" pageNumber="352">Mai</collectorName>
&amp;
<collectorName id="26AFE3EEFF93FFB8C114CEF8FD3A38F0" box="[516,673,727,751]" pageId="7" pageNumber="352">T. M. Nguyen</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B328C65FF93FFB8C1F1CEF8FBF6390C" ID-GBIF-Occurrence="1586985597" collectingDate="2000-02-10" collectingDateMax="2000-03-15" collectingDateMin="2000-02-10" collectionCode="IEBR, RMNH" collectorName="Cuc Phuong NP &amp; P. Q. Mai" location="Ninh Binh" pageId="7" pageNumber="352" specimenCount="2" specimenCount-female="2" stateProvince="Ninh Binh" typeStatus="paratype">
<typeStatus id="54E1389AFF93FFB8C1F1CEF8FCC938F0" box="[737,850,727,751]" pageId="7" pageNumber="352" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5C4DB1FF93FFB8C04ECEF8FC1B38F0" box="[862,896,727,751]" pageId="7" pageNumber="352" type="female">1♀</specimenCount>
(
<collectionCode id="ED4B1EFDFF93FFB8C080CEF8FC4E38F0" box="[912,981,727,751]" pageId="7" pageNumber="352">IEBR</collectionCode>
), similar data as
<typeStatus id="54E1389AFF93FFB8C783CEF9FB6338F0" box="[1171,1272,726,751]" pageId="7" pageNumber="352" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D5C4DB1FF93FFB8C614CEF8FABC38F0" box="[1284,1319,727,751]" pageId="7" pageNumber="352" type="female">1♀</specimenCount>
(
<collectionCode id="ED4B1EFDFF93FFB8C626CEF8FA0B38F0" box="[1334,1424,727,751]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="352">RMNH</collectionCode>
),
<collectingRegion id="499E48DAFF93FFB8C387CED5FE8A390C" box="[151,273,762,787]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="352">Ninh Binh</collectingRegion>
,
<collectorName id="26AFE3EEFF93FFB8C20ECED4FE4C390D" box="[286,471,762,787]" pageId="7" pageNumber="352">Cuc Phuong NP</collectorName>
, near centre,
<date id="FFE4A0F8FF93FFB8C161CED5FCB6390C" box="[625,813,762,787]" pageId="7" pageNumber="352" value="2000-02-10" valueMax="2000-03-15" valueMin="2000-02-10">
<collectingDate id="EFA05910FF93FFB8C161CED5FCB6390C" box="[625,813,762,787]" pageId="7" pageNumber="352" value="2000-02-10" valueMax="2000-03-15" valueMin="2000-02-10">
10.ii15.iii.
<quantity id="4CA22BDDFF93FFB8C1FECED5FCB6390C" box="[750,813,762,787]" metricMagnitude="6" metricUnit="m" metricValue="2.0002249999999995" pageId="7" pageNumber="352" unit="m" value="2000225.0">2000</quantity>
</collectingDate>
</date>
, 225 m a.s.l,
<collectorName id="26AFE3EEFF93FFB8C0DACED4FBA3390C" box="[970,1080,762,787]" pageId="7" pageNumber="352">P. Q. Mai</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B328C65FF93FFB8C76BCED4FC393928" ID-GBIF-Occurrence="1586985606" collectingDate="2000-05-15" collectingDateMax="2000-05-27" collectingDateMin="2000-05-15" collectionCode="RMNH" collectorName="Cuc Phuong NP &amp; P. Q. Mai" location="Ninh Binh" pageId="7" pageNumber="352" specimenCount="1" specimenCount-female="1" stateProvince="Ninh Binh" typeStatus="paratype">
<specimenCount id="9D5C4DB1FF93FFB8C76BCED4FB05390C" box="[1147,1182,763,787]" pageId="7" pageNumber="352" type="female">1♀</specimenCount>
(
<collectionCode id="ED4B1EFDFF93FFB8C7BFCED4FA92390C" box="[1199,1289,763,787]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="352">RMNH</collectionCode>
),
<collectingRegion id="499E48DAFF93FFB8C60ECED5FA03390C" box="[1310,1432,762,787]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="352">Ninh Binh</collectingRegion>
,
<collectorName id="26AFE3EEFF93FFB8C387CF30FED03929" box="[151,331,798,823]" pageId="7" pageNumber="352">Cuc Phuong NP</collectorName>
, near centre,
<date id="FFE4A0F8FF93FFB8C2F0CF31FDE03928" box="[480,635,798,823]" pageId="7" pageNumber="352" value="2000-05-15" valueMax="2000-05-27" valueMin="2000-05-15">
<collectingDate id="EFA05910FF93FFB8C2F0CF31FDE03928" box="[480,635,798,823]" pageId="7" pageNumber="352" value="2000-05-15" valueMax="2000-05-27" valueMin="2000-05-15">
1527.v.
<quantity id="4CA22BDDFF93FFB8C12DCF31FDE03928" box="[573,635,798,823]" metricMagnitude="6" metricUnit="m" metricValue="2.0002249999999995" pageId="7" pageNumber="352" unit="m" value="2000225.0">2000</quantity>
</collectingDate>
</date>
, 225 m a.s.l,
<collectorName id="26AFE3EEFF93FFB8C01FCF30FCE23928" box="[783,889,798,823]" pageId="7" pageNumber="352">P. Q. Mai</collectorName>
leg
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF93FFB6C3D7CF6DFDCE3822" lastPageId="9" lastPageNumber="354" pageId="7" pageNumber="352" type="description">
<paragraph id="8BE58638FF93FFB8C3D7CF6DFC8A3E48" blockId="7.[151,1437,582,2011]" pageId="7" pageNumber="352">
<emphasis id="B92E5A2AFF93FFB8C3D7CF6DFDD13944" bold="true" box="[199,586,834,859]" pageId="7" pageNumber="352">
Description of
<typeStatus id="54E1389AFF93FFB8C291CF6DFE733944" box="[385,488,834,859]" pageId="7" pageNumber="352" type="holotype">holotype</typeStatus>
female.
</emphasis>
Head. Antenna with 25 flagellomeres, first flagellomere 1.2× length of second; frons concave and polished medially, punctate laterally, carina between antennal sockets extending to elevated part of face; face with dense, radiating granulation, 0.65× as high as wide, medially swollen; clypeus about 0.5× as high as wide, sculptured as face, subapical margin with median and lateral tubercles; malar space equal to basal width of mandible; mandible with transverse striation proximally, lower tooth of mandible much longer than upper, inner margins of eyes divergent ventrally; diameter of lateral ocellus 0.8× ocellar-ocular distance; vertex with sparse, fine punctures; occipital carina evenly curved, ventrally meeting hypostomal carina about 0.6× basal mandible width from base of mandible.
</paragraph>
<paragraph id="8BE58638FF93FFB8C3D7C84CFE9C3C51" blockId="7.[151,1437,582,2011]" pageId="7" pageNumber="352">
Mesosoma. Epomia equal to mandible basal width; pronotum with strong striations across median groove, posterior corner densely punctate; mesoscutum 0.85× as long as wide at anterior level of tegulae, with dense, moderate-sized punctures, pubescent, convergence area of notauli with two long median carinae and strong irregular wrinkles laterally; scuto-scutellar groove with one median and two oblique lateral carinae; scutellum convex, densely punctate, pubescent, lateral carina present anteriorly; postscutellum polished, impunctate; mesopleuron with moderate-sized punctures, pubescent, except concave, smooth area medially, epicnemial carina complete, sharp, posterior transverse carina of mesosternum present medially; metapleuron densely punctate, pubescent; propodeum with dense, small punctures anteriorly, area basalis wider than long, anterior transverse carina weakly angulate medially, area dentipara nearly defined, pleural area polished, impunctate, pleural parts of anterior and posterior transverse carinae present; fore basitarsus 2.3× second tarsomere; mid basitarsus 2.7× second tarsomere; hind leg with femur 2.7× as long as wide, 0.7× length of tibia, tarsus 0.7× tibia length, basitarsus 2.6× second tarsomere, fifth tarsomere 2.4× longer than third. Fore wing length
<quantity id="4CA22BDDFF93FFB8C0FDC9C0FBDD3C19" box="[1005,1094,1519,1543]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="7" pageNumber="352" unit="mm" value="7.8">7.8 mm</quantity>
, vein
<emphasis id="B92E5A2AFF93FFB8C79BC9C0FB4F3C19" box="[1163,1236,1519,1542]" italics="true" pageId="7" pageNumber="352">Rs&amp;M</emphasis>
slightly distad of
<emphasis id="B92E5A2AFF93FFB8C387CA3CFF523C35" box="[151,201,1555,1578]" italics="true" pageId="7" pageNumber="352">cu-a</emphasis>
, vein 2
<emphasis id="B92E5A2AFF93FFB8C20FCA3CFEC93C35" box="[287,338,1555,1578]" italics="true" pageId="7" pageNumber="352">rs-m</emphasis>
about 5× distance between 2
<emphasis id="B92E5A2AFF93FFB8C1B0CA3CFD4F3C35" box="[672,724,1555,1578]" italics="true" pageId="7" pageNumber="352">rs-m</emphasis>
and 2
<emphasis id="B92E5A2AFF93FFB8C00BCA3CFCC83C35" box="[795,851,1555,1578]" italics="true" pageId="7" pageNumber="352">m-cu</emphasis>
, hind wing with first abscissa of vein
<emphasis id="B92E5A2AFF93FFB8C61ACA3CFAB03C35" box="[1290,1323,1555,1578]" italics="true" pageId="7" pageNumber="352">Cu</emphasis>
1 equal to vein
<emphasis id="B92E5A2AFF93FFB8C3DECA18FE9C3C51" box="[206,263,1591,1614]" italics="true" pageId="7" pageNumber="352">cu-a.</emphasis>
</paragraph>
<paragraph id="8BE58638FF93FFB8C3D7CA74FD633CC0" blockId="7.[151,1437,582,2011]" pageId="7" pageNumber="352">Metasoma. Tergites with small punctures, pubescent; first tergite 1.6× as long as posterior width, punctate medially and apico-laterally, lateral longitudinal carina present not extending to level of spiracle; second tergite 0.4× as long as first tergite, equal to third; first sternite with projection hooked anteriorly; ovipositor 1.8× as long as hind tibia; hypopygium 2.2× as long as second tergite.</paragraph>
<paragraph id="8BE58638FF93FFB7C3D7CAC5FC153ACA" blockId="7.[151,1437,582,2011]" lastBlockId="8.[151,1436,151,392]" lastPageId="8" lastPageNumber="353" pageId="7" pageNumber="352">Colour. Yellow. Antenna black, except scape, pedicel and first two flagellomeres yellow ventrally; frons with paired black spots at concave, impunctate area, vertex with a black band extending from eyes to ocelli, occipital area black; mesoscutum with black mark at anterior edge of median lobe and trilobed black mark posteriorly, shaped rather like a crown, tegula black, apex of scutellum and postscutellum black, mesopleuron with black mark along anterior margin, metapleuron with black spot at anterior margin, propodeum black anteriorly, metasomal tergite 1 black medially and baso-laterally, tergites from second onwards each with black stripe anteriorly, ovipositor sheath light brown, blackish apically, fore wings with distal part and area below prestigma infuscate, mid leg with brown on coxa and femur, hind leg with black marks on coxa, trochanter, trochantellus and femur, tibia reddish with black marks proximally and distally, tarsus black.</paragraph>
<paragraph id="8BE58638FF9CFFB7C3D7CCF0FE1E3AE7" blockId="8.[151,1436,151,392]" box="[199,389,223,248]" pageId="8" pageNumber="353">Male. Unknown.</paragraph>
<paragraph id="8BE58638FF9CFFB7C3D7CD2BFB1B3B97" blockId="8.[151,1436,151,392]" pageId="8" pageNumber="353">
Variation:
<typeStatus id="54E1389AFF9CFFB7C22FCD2AFE373B02" box="[319,428,261,285]" pageId="8" pageNumber="353" type="paratype">Paratypes</typeStatus>
have antennae with 22-23 flagellomeres. Fore wing vein 2
<emphasis id="B92E5A2AFF9CFFB7C744CD2AFB133B03" box="[1108,1160,261,284]" italics="true" pageId="8" pageNumber="353">rs-m</emphasis>
varies from 1.0 to 3.3× distance between 2
<emphasis id="B92E5A2AFF9CFFB7C262CD06FE3D3B5F" box="[370,422,297,320]" italics="true" pageId="8" pageNumber="353">rs-m</emphasis>
and 2
<emphasis id="B92E5A2AFF9CFFB7C2FFCD06FDBC3B5F" box="[495,551,297,320]" italics="true" pageId="8" pageNumber="353">m-cu</emphasis>
. Hind wing first abscissa of vein
<emphasis id="B92E5A2AFF9CFFB7C0A4CD07FC4E3B20" box="[948,981,296,319]" italics="true" pageId="8" pageNumber="353">Cu</emphasis>
1 is slightly longer than vein
<emphasis id="B92E5A2AFF9CFFB7C63DCD06FAC53B5F" box="[1325,1374,297,320]" italics="true" pageId="8" pageNumber="353">cu-a</emphasis>
. The
<typeStatus id="54E1389AFF9CFFB7C387CD62FF6C3B7A" box="[151,247,333,357]" pageId="8" pageNumber="353" type="paratype">paratype</typeStatus>
collected in Cat Tien NP (similar data to the
<typeStatus id="54E1389AFF9CFFB7C004CD63FCE33B7A" box="[788,888,332,357]" pageId="8" pageNumber="353" type="holotype">holotype</typeStatus>
) has fore wing vein
<emphasis id="B92E5A2AFF9CFFB7C77DCD63FB2C3B7C" box="[1133,1207,332,355]" italics="true" pageId="8" pageNumber="353">Rs&amp;M</emphasis>
opposite
<emphasis id="B92E5A2AFF9CFFB7C63ACD62FAC73B7B" box="[1322,1372,333,356]" italics="true" pageId="8" pageNumber="353">cu-a</emphasis>
. One
<typeStatus id="54E1389AFF9CFFB7C387CD5FFF6C3B97" box="[151,247,368,392]" pageId="8" pageNumber="353" type="paratype">paratype</typeStatus>
collected in Cuc Phuong NP with first tergite 1.4× as long as its posterior width.
</paragraph>
<caption id="DF25D6B0FF9CFFB7C387CAFFFD273D1A" httpUri="https://zenodo.org/record/847862/files/figure.png" pageId="8" pageNumber="353" targetBox="[160,1420,454,1714]" targetPageId="8">
<paragraph id="8BE58638FF9CFFB7C387CAFFFD273D1A" blockId="8.[151,1435,1744,1797]" pageId="8" pageNumber="353">
<emphasis id="B92E5A2AFF9CFFB7C387CAFFFE893CFA" bold="true" box="[151,274,1744,1766]" pageId="8" pageNumber="353">FIGURE 4.</emphasis>
<taxonomicName id="4C5AFDBBFF9CFFB7C208CAFDFE7D3CF8" authority="Achterberg, 2017" authorityName="Achterberg" authorityYear="2017" box="[280,486,1746,1767]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="coronatus" status="sp. nov.">
<emphasis id="B92E5A2AFF9CFFB7C208CAFDFE7D3CF8" box="[280,486,1746,1767]" italics="true" pageId="8" pageNumber="353">Yezoceryx coronatus</emphasis>
</taxonomicName>
<emphasis id="B92E5A2AFF9CFFB7C2FCCAFEFDA73CF9" bold="true" box="[492,572,1745,1766]" pageId="8" pageNumber="353">
<taxonomicNameLabel id="A21DE751FF9CFFB7C2FCCAFEFDA73CF9" box="[492,572,1745,1766]" pageId="8" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: a. lateral view; b. dorsal view; c. face; d. lateral view of mesosoma; e. lateral view of first metasomal tergite; f. lateral view of hypopygium.
</paragraph>
</caption>
<paragraph id="8BE58638FF9CFFB6C3D7CB1DFB8A3BD0" blockId="8.[151,1436,1842,2011]" lastBlockId="9.[151,1437,151,680]" lastPageId="9" lastPageNumber="354" pageId="8" pageNumber="353">
Comparison. In general colour pattern, the new species is similar to
<taxonomicName id="4C5AFDBBFF9CFFB7C0FBCB1CFA073D55" authority="(Morley, 1913)" authorityName="Morley" authorityYear="1913" baseAuthorityName="Morley" baseAuthorityYear="1913" box="[1003,1436,1842,1867]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="biumbratus">
<emphasis id="B92E5A2AFF9CFFB7C0FBCB1CFB7E3D55" box="[1003,1253,1842,1866]" italics="true" pageId="8" pageNumber="353">Yezoceryx biumbratus</emphasis>
(Morley, 1913)
</taxonomicName>
(holotype female in BMNH, examined) and
<taxonomicName id="4C5AFDBBFF9CFFB7C189CB78FC5E3D71" authority="(Cameron, 1899)" authorityName="Cameron" authorityYear="1899" baseAuthorityName="Cameron" baseAuthorityYear="1899" box="[665,965,1878,1902]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="B92E5A2AFF9CFFB7C189CB78FD623D71" box="[665,761,1878,1902]" italics="true" pageId="8" pageNumber="353">Y. luteus</emphasis>
(
<bibRefCitation id="EFCBFBC9FF9CFFB7C01BCB79FC253D71" author="Cameron" box="[779,958,1878,1902]" pageId="8" pageNumber="353" refString="Cameron, P. (1899) Hymenoptera Orientalia, or Contributions to a knowledge of the Hymenoptera of the Oriental Zoological Region. Part VIII. The Hymenoptera of the Khasia Hills. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 43, 1 - 220." type="journal article" year="1899">Cameron, 1899</bibRefCitation>
)
</taxonomicName>
(holotype female in BMNH, examined), particularly the former, which is also structurally closest. However, the color of the mesoscutum, mesopleuron and metasomal tergite 1 of
<taxonomicName id="4C5AFDBBFF9CFFB7C28CCBB0FDBD3DA9" authority="Achterberg, 2017" authorityName="Achterberg" authorityYear="2017" box="[412,550,1951,1974]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="coronatus" status="sp. nov.">
<emphasis id="B92E5A2AFF9CFFB7C28CCBB0FDBD3DA9" box="[412,550,1951,1974]" italics="true" pageId="8" pageNumber="353">Y. coronatus</emphasis>
</taxonomicName>
<emphasis id="B92E5A2AFF9CFFB7C13ECBB2FD133DA9" bold="true" box="[558,648,1949,1974]" pageId="8" pageNumber="353">
<taxonomicNameLabel id="A21DE751FF9CFFB7C13ECBB2FD133DA9" box="[558,648,1949,1974]" pageId="8" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs. In
<taxonomicName id="4C5AFDBBFF9CFFB7C016CBB0FC3A3DA9" baseAuthorityName="Morley" baseAuthorityYear="1913" box="[774,929,1950,1974]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="biumbratus">
<emphasis id="B92E5A2AFF9CFFB7C016CBB0FC3A3DA9" box="[774,929,1950,1974]" italics="true" pageId="8" pageNumber="353">Y. biumbratus</emphasis>
</taxonomicName>
the black markings are more restricted, with an entirely yellow metapleuron. In
<taxonomicName id="4C5AFDBBFF9CFFB7C132CBECFD1B3DC5" baseAuthorityName="Cameron" baseAuthorityYear="1899" box="[546,640,1986,2010]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="B92E5A2AFF9CFFB7C132CBECFD1B3DC5" box="[546,640,1986,2010]" italics="true" pageId="8" pageNumber="353">Y. luteus</emphasis>
</taxonomicName>
, the median lobe of the mesoscutum is black anteriorly, amongst other differences. In
<bibRefCitation id="EFCBFBC9FF9DFFB6C22FCCB7FE483AAF" author="Wang" box="[319,467,152,176]" pageId="9" pageNumber="354" refString="Wang, S. - F. (1993) Studies on the genus Yezoceryx Uchida of China (Ichneumonidae: Acaenitinae). Scientific Treatise on Systematic and Evolutionary Zoology, 2, 203 - 218." type="journal article" year="1993">Wang (1993)</bibRefCitation>
, this new species keys to
<taxonomicName id="4C5AFDBBFF9DFFB6C1E2CCB6FBC13AB0" authority="Wang. It" authorityName="Wang. It" authorityYear="1993" box="[754,1114,152,176]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="apicicarinus">
<emphasis id="B92E5A2AFF9DFFB6C1E2CCB6FC683AB0" box="[754,1011,152,176]" italics="true" pageId="9" pageNumber="354">Yezoceryx apicicarinus</emphasis>
Wang. It
</taxonomicName>
differs from the latter by the sculpture of the face and the colour pattern of the mesoscutum and mesopleuron. Two females from Papua New Guinea in BMNH (Bulolo, Mankica,
<date id="FFE4A0F8FF9DFFB6C159CCF0FD533AE7" box="[585,712,223,248]" pageId="9" pageNumber="354" value="1981-12-11">11.xii.1981</date>
) are very similar to the type series from Vietnam, differing in minor details of colour pattern, such as the more extensive black markings on the mesopleuron and hind femur; we refrain from including these specimens in the type series.
<taxonomicName id="4C5AFDBBFF9DFFB6C02DCD06FBAD3B5F" baseAuthorityName="Morley" baseAuthorityYear="1913" box="[829,1078,296,320]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="biumbratus">
<emphasis id="B92E5A2AFF9DFFB6C02DCD06FBAD3B5F" box="[829,1078,296,320]" italics="true" pageId="9" pageNumber="354">Yezoceryx biumbratus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5AFDBBFF9DFFB6C757CD06FB4E3B5F" authority="Achterberg, 2017" authorityName="Achterberg" authorityYear="2017" box="[1095,1237,297,320]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="coronatus" status="sp. nov.">
<emphasis id="B92E5A2AFF9DFFB6C757CD06FB4E3B5F" box="[1095,1237,297,320]" italics="true" pageId="9" pageNumber="354">Y. coronatus</emphasis>
</taxonomicName>
<emphasis id="B92E5A2AFF9DFFB6C7CFCD08FAA63B5F" bold="true" box="[1247,1341,295,320]" pageId="9" pageNumber="354">
<taxonomicNameLabel id="A21DE751FF9DFFB6C7CFCD08FAA63B5F" box="[1247,1341,295,320]" pageId="9" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and the specimens from Papua New Guinea may conceivably be at the opposite ends of a colour spectrum, from palest in India (
<taxonomicName id="4C5AFDBBFF9DFFB6C3F4CD5EFE1A3B97" baseAuthorityName="Morley" baseAuthorityYear="1913" box="[228,385,368,392]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="biumbratus">
<emphasis id="B92E5A2AFF9DFFB6C3F4CD5EFE1A3B97" box="[228,385,368,392]" italics="true" pageId="9" pageNumber="354">Y. biumbratus</emphasis>
</taxonomicName>
) through to darkest in Papua New Guinea. All these specimens share an apically widened antennal flagellum relative to the slenderer proximal segments. However, we treat these as separate species for now, given the known disjunct range and distinct differences in colour patterns.
</paragraph>
<paragraph id="8BE58638FF9DFFB6C3D7CDF2FDCE3822" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="354">
We also take the opportunity to note that
<taxonomicName id="4C5AFDBBFF9DFFB6C184CDF2FCD63BEB" baseAuthorityName="Cameron" baseAuthorityYear="1899" box="[660,845,476,500]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="B92E5A2AFF9DFFB6C184CDF2FCD63BEB" box="[660,845,476,500]" italics="true" pageId="9" pageNumber="354">Yezoceryx luteus</emphasis>
</taxonomicName>
(original combination
<taxonomicName id="4C5AFDBBFF9DFFB6C75ECDF2FF493807" authority="Cameron, 1899" authorityName="Cameron" authorityYear="1899" class="Insecta" genus="Macrogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="B92E5A2AFF9DFFB6C75ECDF2FAB33BEB" box="[1102,1320,476,500]" italics="true" pageId="9" pageNumber="354">Macrogaster luteus</emphasis>
<bibRefCitation id="EFCBFBC9FF9DFFB6C63ECDF3FF493807" author="Cameron" pageId="9" pageNumber="354" refString="Cameron, P. (1899) Hymenoptera Orientalia, or Contributions to a knowledge of the Hymenoptera of the Oriental Zoological Region. Part VIII. The Hymenoptera of the Khasia Hills. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 43, 1 - 220." type="journal article" year="1899">Cameron, 1899</bibRefCitation>
</taxonomicName>
, holotype male in BMNH, examined) is a senior synonym of
<taxonomicName id="4C5AFDBBFF9DFFB6C087CE2FFAB33807" authority="Morley, 1913" authorityName="Morley" authorityYear="1913" box="[919,1320,511,536]" class="Insecta" family="Ichneumonidae" genus="Acaenitus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="xanthorius">
<emphasis id="B92E5A2AFF9DFFB6C087CE2FFB1D3807" box="[919,1158,512,536]" italics="true" pageId="9" pageNumber="354">Acaenitus xanthorius</emphasis>
Morley, 1913
</taxonomicName>
(holotype female in BMNH, examined):
<emphasis id="B92E5A2AFF9DFFB6C2FECE0BFDCE3822" bold="true" box="[494,597,548,573]" pageId="9" pageNumber="354">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF9DFFB6C3D7CE68FB833840" box="[199,1048,583,608]" pageId="9" pageNumber="354" type="etymology">
<paragraph id="8BE58638FF9DFFB6C3D7CE68FB833840" blockId="9.[151,1437,151,680]" box="[199,1048,583,608]" pageId="9" pageNumber="354">
<emphasis id="B92E5A2AFF9DFFB6C3D7CE68FED5387F" bold="true" box="[199,334,583,608]" pageId="9" pageNumber="354">Etymology.</emphasis>
Named after the crown-shaped black mark on the mesoscutum.
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF9DFFB6C3D7CE43FE6F38B7" pageId="9" pageNumber="354" type="distribution">
<paragraph id="8BE58638FF9DFFB6C3D7CE43FE6F38B7" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="354">
<emphasis id="B92E5A2AFF9DFFB6C3D7CE43FEFB389A" bold="true" box="[199,352,620,645]" pageId="9" pageNumber="354">Distribution.</emphasis>
Currently known only from Cuc Phuong NP,
<collectingRegion id="499E48DAFF9DFFB6C062CE43FBCC389A" box="[882,1111,620,645]" country="Vietnam" name="Ninh Binh" pageId="9" pageNumber="354">Ninh Binh Province</collectingRegion>
and Cat Tien NP,
<collectingRegion id="499E48DAFF9DFFB6C63CCE42FF6538B7" country="Vietnam" name="Dong Nai" pageId="9" pageNumber="354">Dong Nai Province</collectingRegion>
,
<collectingCountry id="F34DC6A8FF9DFFB6C218CEA0FEFC38B7" box="[264,359,655,680]" name="Vietnam" pageId="9" pageNumber="354">Vietnam</collectingCountry>
(
<figureCitation id="13619ABDFF9DFFB6C266CEA0FE7D38B7" box="[374,486,655,680]" captionStart="FIGURE 19" captionStartId="25.[151,250,1940,1962]" captionTargetBox="[241,1347,195,1919]" captionTargetId="figure@25.[241,1350,195,1923]" captionTargetPageId="25" captionText="FIGURE 19. Map of sampling sites of Yezoceryx spp in Vietnam: 1. Cao Bang Province; 2. Bac Kan Province; 3. Vinh Phuc Province, 4. Son La Province; 5. Hoa Binh Province; 6. Ninh Binh Province; 7. Nghe An Province; 8. Thua Thien-Hue Province; 9. Kon Tum Province; 10. Dak Lak Province; 11. Dong Nai Province." httpUri="https://zenodo.org/record/847892/files/figure.png" pageId="9" pageNumber="354">Figure 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>