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<document id="AC4316ACCF5295C99F5FAB12CD792DB0" ID-DOI="10.1093/zoolinnean/zlad078" ID-ISSN="0024-4082" ID-Zenodo-Dep="11240120" ID-ZooBank="5D6C7EBF-E568-4100-9364-2DD357003878" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716361325736" checkinUser="plazi" docAuthor="Marceniuk, Alexandre Pires, Oliveira, Claudio &amp; Ferraris Jr, Carl J." docDate="2024" docId="03F32B77FFEAFFAF095EFB07FD814CC5" docLanguage="en" docName="zlad078.pdf" docOrigin="Zoological Journal of the Linnean Society 200 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad078" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Notarius Gill 1863" docType="treatment" docVersion="2" lastPageNumber="440" masterDocId="FFCA530FFFE7FFA10D44FFD6FFAB4A1E" masterDocTitle="A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data" masterLastPageNumber="476" masterPageNumber="426" pageNumber="439" updateTime="1716362088827" updateUser="ExternalLinkService">
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<mods:title id="4C74CCE5057D1A72E7BF099329A4A8B1">A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data</mods:title>
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<mods:namePart id="E6FFDA929BAD33B6C3EEF6B57D865B19">Marceniuk, Alexandre Pires</mods:namePart>
<mods:affiliation id="7C9B6116CBACC0E2FC24BCF6CDEC2D42">Programa de Pós-Graduação em Ciências Biológicas, Universidade Federal da Paraíba, conjunto Presidente Castelo Branco III, João Pessoa, PB 58050 - 000, Brazil &amp; Programa de Pós-Graduação em Ecologia e Conservação, Universidade Estadual da Paraíba, Campina Grande, PB 58429 - 500, Brazil &amp; Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
<mods:nameIdentifier id="1080BDA40EDAE6E6BE969A3BBCEEB195" type="email">a_marceniuk@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="C4AB6B6ACE886BECE02FBBB303EB9958">Oliveira, Claudio</mods:namePart>
<mods:affiliation id="C79EC8B67EBDAFEE5A974C7F688A805F">Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
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<mods:namePart id="40AA85CCDD0BED703B9E58E88F4A0702">Ferraris Jr, Carl J.</mods:namePart>
<mods:affiliation id="1B896D4B3CE3D88D123F4B6A51F4E4C4">545 NE Floral Pl., Portland, OR 97232, USA</mods:affiliation>
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<mods:title id="41BD6255F1597AF3FBCBFAA236BCFAD3">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="FBAF09D7F0B5A7942D7AD50243131F45">2024</mods:date>
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<mods:number id="A5947A8491F5C9446BE086FE44552FEA">2023-08-21</mods:number>
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<treatment id="03F32B77FFEAFFAF095EFB07FD814CC5" LSID="urn:lsid:plazi:treatment:03F32B77FFEAFFAF095EFB07FD814CC5" httpUri="http://treatment.plazi.org/id/03F32B77FFEAFFAF095EFB07FD814CC5" lastPageId="14" lastPageNumber="440" pageId="13" pageNumber="439">
<subSubSection id="C340C9EAFFEAFFAC095EFB07FB474F03" pageId="13" pageNumber="439" type="nomenclature">
<paragraph id="8BE59A61FFEAFFAC095EFB07FB4A4EF5" blockId="13.[1050,1249,1232,1259]" box="[1050,1249,1232,1259]" pageId="13" pageNumber="439">
<heading id="D0AD2D0DFFEAFFAC095EFB07FB4A4EF5" bold="true" box="[1050,1249,1232,1259]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="439" reason="2">
<taxonomicName id="4C5AE1E2FFEAFFAC095EFB07FB4A4EF5" ID-CoL="64Z5" ID-ENA="286639" authority="Gill, 1863" authorityName="Gill" authorityYear="1863" box="[1050,1249,1232,1259]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="439" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFEAFFAC095EFB07FB4A4EF5" bold="true" box="[1050,1249,1232,1259]" pageId="13" pageNumber="439">
<emphasis id="B92E4673FFEAFFAC095EFB07FBD84EF5" bold="true" box="[1050,1139,1233,1259]" italics="true" pageId="13" pageNumber="439">Notarius</emphasis>
Gill, 1863
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE59A61FFEAFFAC0954FAD5FB474F03" blockId="13.[1040,1260,1283,1309]" box="[1040,1260,1283,1309]" pageId="13" pageNumber="439">
<heading id="D0AD2D0DFFEAFFAC0954FAD5FB474F03" allCaps="true" bold="true" box="[1040,1260,1283,1309]" centered="true" fontSize="11" level="1" pageId="13" pageNumber="439" reason="1">
<emphasis id="B92E4673FFEAFFAC0954FAD5FB474F03" bold="true" box="[1040,1260,1283,1309]" pageId="13" pageNumber="439">MP 100, BI 1, ML 93</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340C9EAFFEAFFAF095BFAF9FD814CC5" lastPageId="14" lastPageNumber="440" pageId="13" pageNumber="439" type="description">
<paragraph id="8BE59A61FFEAFFAC095BFAF9FB774F57" blockId="13.[1055,1244,1327,1354]" box="[1055,1244,1327,1354]" pageId="13" pageNumber="439">
<emphasis id="B92E4673FFEAFFAC095BFAF9FB774F57" bold="true" box="[1055,1244,1327,1354]" pageId="13" pageNumber="439">
(
<figureCitation id="136186E4FFEAFFAC096EFAF9FB284F57" box="[1066,1155,1327,1353]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[129,194,1805,1829]" captionStartId-1="3.[825,890,1803,1827]" captionStartId-2="4.[113,178,1813,1837]" captionTargetBox-1="[153,1434,148,1768]" captionTargetBox-2="[151,708,146,1775]" captionTargetId-2="figure-643@4.[149,717,144,1776]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-1="Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-2="Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." figureDoi-1="http://doi.org/10.5281/zenodo.11240122" figureDoi-2="http://doi.org/10.5281/zenodo.11240126" httpUri-1="https://zenodo.org/record/11240122/files/figure.png" httpUri-2="https://zenodo.org/record/11240126/files/figure.png" pageId="13" pageNumber="439">Figs 1–3</figureCitation>
,
<figureCitation id="136186E4FFEAFFAC09CAFAF9FB004F57" box="[1166,1195,1327,1353]" captionStart="Figure 9" captionStartId="12.[113,178,1620,1644]" captionTargetBox="[116,1458,151,1592]" captionTargetId="figure-88@12.[114,1458,144,1593]" captionTargetPageId="12" captionText="Figure 9. Representative skulls, in dorsal view, and distribution of Cathoropsini tribe nov. A, Cathorops hypophthalmus, USNM 293275. B, Notarius grandicassis, MPEG 30647. * marine, ** brackish waters, *** freshwater." figureDoi="http://doi.org/10.5281/zenodo.11240139" httpUri="https://zenodo.org/record/11240139/files/figure.png" pageId="13" pageNumber="439">9B</figureCitation>
,
<figureCitation id="136186E4FFEAFFAC09F2FAF9FB794F57" box="[1206,1234,1327,1353]" captionStart="Figure 11" captionStartId="14.[113,178,648,672]" captionTargetBox="[114,1458,143,619]" captionTargetId="figure-488@14.[114,1458,143,619]" captionTargetPageId="14" captionText="Figure 11. Notarius grandicassis, MNHN A-4608, Atlantic South America, type species of the genus." figureDoi="http://doi.org/10.5281/zenodo.11240143" httpUri="https://zenodo.org/record/11240143/files/figure.png" pageId="13" pageNumber="439">11</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BE59A61FFEAFFAC0E7DFAAFFA934F8F" blockId="13.[825,1336,1400,1425]" box="[825,1336,1400,1425]" pageId="13" pageNumber="439">
<emphasis id="B92E4673FFEAFFAC0E7DFAAFFC1E4F8E" box="[825,949,1401,1424]" italics="true" pageId="13" pageNumber="439">
<typeStatus id="54E124C3FFEAFFAC0E7DFAAFFCC24F8E" box="[825,873,1401,1424]" pageId="13" pageNumber="439">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C5AE1E2FFEAFFAC0EFEFAAFFA9F4F8F" authority="Valenciennes, 1840" authorityName="Valenciennes" authorityYear="1840" box="[954,1332,1400,1425]" family="Ariidae" genus="Arius" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="439" phylum="Chordata" rank="species" species="grandicassis">
<emphasis id="B92E4673FFEAFFAC0EFEFAAFFBCC4F8E" box="[954,1127,1400,1424]" italics="true" pageId="13" pageNumber="439">Arius grandicassis</emphasis>
Valenciennes, 1840
</taxonomicName>
.
</paragraph>
<paragraph id="8BE59A61FFEAFFAC090BFA69FB074FC6" blockId="13.[825,1474,1471,1783]" box="[1103,1196,1471,1496]" pageId="13" pageNumber="439">
<emphasis id="B92E4673FFEAFFAC090BFA69FB074FC6" box="[1103,1196,1471,1496]" italics="true" pageId="13" pageNumber="439">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE59A61FFEAFFAC0E7DFA32FAA84CE9" blockId="13.[825,1474,1471,1783]" pageId="13" pageNumber="439">Epiphyseal bar transversely elongate and longitudinally narrow (29, 0&gt; 1); medial groove of cranium absent 30 (0&gt; 1); vomer anterior margin very pronounced and acute (50, 0&gt; 1); parasphenoid very wide in ventral view (66, 0&gt; 1); posterior process of exoccipital absent (90, 1&gt; 0); exoccipital bony crest parallel to vertebral column and mesially folded (92, 1&gt; 2); sesamoid bone I irregularly shaped (145, 1&gt; 2); accessory crest connecting transverse and median crests associated with neural spine of fourth vertebra present (204, 0&gt; 1).</paragraph>
<paragraph id="8BE59A61FFEAFFAC0E7DF8CEFAD24DD3" blockId="13.[825,1475,1816,1998]" pageId="13" pageNumber="439">
<emphasis id="B92E4673FFEAFFAC0E7DF8CEFB854D2E" box="[825,1070,1816,1840]" italics="true" pageId="13" pageNumber="439">Ambiguous optimization:</emphasis>
Lateral horn of lateral ethmoid acute (11, 1&gt; 0); interior margin of otolith straight or slightly irregular (71, 0&gt; 1); anterior opening of aortic canal at base of subvertebral process and anteroventrally oriented (
<emphasis id="B92E4673FFEAFFAC0985F8A1FA894D90" bold="true" box="[1217,1314,1910,1935]" pageId="13" pageNumber="439">77,?&gt; 1</emphasis>
); ventral tip of subvertebral process acute (80,?&gt; 2); adipose-fin base long (222, 3&gt; 1); gas bladder lateral diverticula present (243, 0&gt; 1).
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FF9CFEF54A41" blockId="14.[113,350,74,96]" box="[113,350,74,96]" pageId="14" pageNumber="440">
<emphasis id="B92E4673FFE9FFAF0D35FF9CFF1C4A7E" bold="true" box="[113,183,74,96]" pageId="14" pageNumber="440">440 •</emphasis>
Marceniuk
<emphasis id="B92E4673FFE9FFAF0C70FF9DFEF54A41" box="[308,350,74,96]" italics="true" pageId="14" pageNumber="440">et al.</emphasis>
</paragraph>
<caption id="DF25CAE9FFE9FFAF0D35FD5EFBB448BF" ID-DOI="http://doi.org/10.5281/zenodo.11240143" ID-Zenodo-Dep="11240143" box="[113,1055,648,673]" httpUri="https://zenodo.org/record/11240143/files/figure.png" pageId="14" pageNumber="440" startId="14.[113,178,648,672]" targetBox="[114,1458,143,619]" targetPageId="14" targetType="figure">
<paragraph id="8BE59A61FFE9FFAF0D35FD5EFBB448BF" blockId="14.[113,1055,648,673]" box="[113,1055,648,673]" pageId="14" pageNumber="440">
<emphasis id="B92E4673FFE9FFAF0D35FD5EFF7D48BE" bold="true" box="[113,214,648,672]" pageId="14" pageNumber="440">Figure 11.</emphasis>
<taxonomicName id="4C5AE1E2FFE9FFAF0D9EFD5FFE5A48BE" authority=", MNHN" authorityName="MNHN" box="[218,497,648,672]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="grandicassis">
<emphasis id="B92E4673FFE9FFAF0D9EFD5FFE3148BE" box="[218,410,648,672]" italics="true" pageId="14" pageNumber="440">Notarius grandicassis</emphasis>
, MNHN
</taxonomicName>
A-4608, Atlantic South America, type species of the genus.
</paragraph>
</caption>
<paragraph id="8BE59A61FFE9FFAF0C29FD06FE5548F4" blockId="14.[113,673,720,1346]" box="[365,510,720,746]" pageId="14" pageNumber="440">
<emphasis id="B92E4673FFE9FFAF0C29FD06FE5548F4" box="[365,510,720,746]" italics="true" pageId="14" pageNumber="440">Included species</emphasis>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FD21FD884930" blockId="14.[113,673,720,1346]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FD21FE944910" box="[113,319,758,782]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="armbrusteri">
<emphasis id="B92E4673FFE9FFAF0D35FD21FE944910" box="[113,319,758,782]" italics="true" pageId="14" pageNumber="440">Notarius armbrusteri</emphasis>
</taxonomicName>
Betancur-R. &amp; Acero P., 2006
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FCC0FF5A4933" box="[113,241,789,813]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="biffi">
<emphasis id="B92E4673FFE9FFAF0D35FCC0FF5A4933" box="[113,241,789,813]" italics="true" pageId="14" pageNumber="440">Notarius biffi</emphasis>
</taxonomicName>
Betancur-R. &amp; Acero P., 2004
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FCE0FE024953" blockId="14.[113,673,720,1346]" box="[113,425,821,845]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FCE0FE024953" authority="(Miles, 1945)" baseAuthorityName="Miles" baseAuthorityYear="1945" box="[113,425,821,845]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="bonillai">
<emphasis id="B92E4673FFE9FFAF0D35FCE0FEBE4953" box="[113,277,821,845]" italics="true" pageId="14" pageNumber="440">Notarius bonillai</emphasis>
(Miles, 1945)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FC83FD8A4992" blockId="14.[113,673,720,1346]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FC83FEA34972" box="[113,264,852,876]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="cookei">
<emphasis id="B92E4673FFE9FFAF0D35FC83FEA34972" box="[113,264,852,876]" italics="true" pageId="14" pageNumber="440">Notarius cookei</emphasis>
</taxonomicName>
(Acero P. &amp; Betancur-R., 2002)
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FCA2FD8A4992" authority="(Valenciennes, 1840)" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[113,545,883,908]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="grandicassis">
<emphasis id="B92E4673FFE9FFAF0D35FCA2FE944995" box="[113,319,883,907]" italics="true" pageId="14" pageNumber="440">Notarius grandicassis</emphasis>
(Valenciennes, 1840)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FC42FD9D49B5" blockId="14.[113,673,720,1346]" box="[113,566,915,939]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FC42FE8749B5" baseAuthorityName="Jordan &amp; Gilbert" baseAuthorityYear="1883" box="[113,300,915,939]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="insculptus">
<emphasis id="B92E4673FFE9FFAF0D35FC42FE8749B5" box="[113,300,915,939]" italics="true" pageId="14" pageNumber="440">Notarius insculptus</emphasis>
</taxonomicName>
(
<collectingCountry id="F34DDAF1FFE9FFAF0C04FC45FE2F49B5" box="[320,388,915,939]" name="Jordan" pageId="14" pageNumber="440">Jordan</collectingCountry>
&amp; Gilbert, 1883)
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FC65FE5949D4" blockId="14.[113,673,720,1346]" box="[113,498,946,970]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FC65FE5949D4" authority="(Steindachner, 1876)" baseAuthorityName="Steindachner" baseAuthorityYear="1876" box="[113,498,946,970]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="kessleri">
<emphasis id="B92E4673FFE9FFAF0D35FC65FEBA49D4" box="[113,273,946,970]" italics="true" pageId="14" pageNumber="440">Notarius kessleri</emphasis>
(Steindachner, 1876)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FC04FE9E49F7" blockId="14.[113,673,720,1346]" box="[113,309,977,1001]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FC04FE9E49F7" baseAuthorityName="Steindachner" baseAuthorityYear="1876" box="[113,309,977,1001]" family="Ariidae" genus="Notarius" isUncertain="true" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="kessleri">
<emphasis id="B92E4673FFE9FFAF0D35FC04FF6C49F7" box="[113,199,978,1001]" italics="true" pageId="14" pageNumber="440">Notarius</emphasis>
aff.
<emphasis id="B92E4673FFE9FFAF0DB4FC07FE9E49F7" box="[240,309,977,1001]" italics="true" pageId="14" pageNumber="440">kessleri</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FC24FDAC4E36" blockId="14.[113,673,720,1346]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FC24FD354E17" authority="(Eigenmann &amp; Eigenmann, 1888)" baseAuthorityName="Eigenmann &amp; Eigenmann" baseAuthorityYear="1888" box="[113,670,1009,1033]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="lentiginosus">
<emphasis id="B92E4673FFE9FFAF0D35FC24FE974E17" box="[113,316,1009,1033]" italics="true" pageId="14" pageNumber="440">Notarius lentiginosus</emphasis>
(Eigenmann &amp; Eigenmann, 1888)
</taxonomicName>
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FBC7FDAC4E36" authority="(Valenciennes, 1840)" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[113,519,1040,1064]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="luniscutis">
<emphasis id="B92E4673FFE9FFAF0D35FBC7FE8E4E36" box="[113,293,1040,1064]" italics="true" pageId="14" pageNumber="440">Notarius luniscutis</emphasis>
(Valenciennes, 1840)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FBE6FD814E79" blockId="14.[113,673,720,1346]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FBE6FEF14E59" box="[113,346,1072,1095]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="neogranatensis">
<emphasis id="B92E4673FFE9FFAF0D35FBE6FEF14E59" box="[113,346,1072,1095]" italics="true" pageId="14" pageNumber="440">Notarius neogranatensis</emphasis>
</taxonomicName>
(Acero P. &amp; Betancur-R., 2002)
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FB99FEBB4E79" baseAuthorityName="Jordan &amp; Gilbert" baseAuthorityYear="1883" box="[113,272,1103,1127]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="osculus">
<emphasis id="B92E4673FFE9FFAF0D35FB99FEBB4E79" box="[113,272,1103,1127]" italics="true" pageId="14" pageNumber="440">Notarius osculus</emphasis>
</taxonomicName>
(
<collectingCountry id="F34DDAF1FFE9FFAF0C60FB99FEC34E79" box="[292,360,1103,1127]" name="Jordan" pageId="14" pageNumber="440">Jordan</collectingCountry>
&amp; Gilbert, 1883) *
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FBB9FD9A4E98" blockId="14.[113,673,720,1346]" box="[113,561,1134,1158]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FBB9FD894E98" authority="(Valenciennes, 1840)" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[113,546,1134,1158]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="parmocassis">
<emphasis id="B92E4673FFE9FFAF0D35FBB9FEEA4E98" box="[113,321,1135,1158]" italics="true" pageId="14" pageNumber="440">Notarius parmocassis</emphasis>
(Valenciennes, 1840)
</taxonomicName>
*
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FB58FDBF4EB8" blockId="14.[113,673,720,1346]" box="[113,532,1165,1190]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FB58FDBF4EB8" authority="(Valenciennes, 1840)" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[113,532,1165,1190]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="phrygiatus">
<emphasis id="B92E4673FFE9FFAF0D35FB58FE984EBB" box="[113,307,1165,1189]" italics="true" pageId="14" pageNumber="440">Notarius phrygiatus</emphasis>
(Valenciennes, 1840)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FB7BFDAE4EDB" blockId="14.[113,673,720,1346]" box="[113,517,1197,1221]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FB7BFDAE4EDB" authority="(Steindachner, 1876)" baseAuthorityName="Steindachner" baseAuthorityYear="1876" box="[113,517,1197,1221]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="planiceps">
<emphasis id="B92E4673FFE9FFAF0D35FB7BFE8E4EDB" box="[113,293,1197,1221]" italics="true" pageId="14" pageNumber="440">Notarius planiceps</emphasis>
(Steindachner, 1876)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FB1BFEE24EFA" blockId="14.[113,673,720,1346]" box="[113,329,1228,1252]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FB1BFEE24EFA" baseAuthorityName="Steindachner" baseAuthorityYear="1876" box="[113,329,1228,1252]" family="Ariidae" genus="Notarius" isUncertain="true" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="planiceps">
<emphasis id="B92E4673FFE9FFAF0D35FB1BFF6C4EFA" box="[113,199,1229,1252]" italics="true" pageId="14" pageNumber="440">Notarius</emphasis>
aff.
<emphasis id="B92E4673FFE9FFAF0DB4FB1AFEE24EFA" box="[240,329,1228,1252]" italics="true" pageId="14" pageNumber="440">planiceps</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FB3AFD894F1A" blockId="14.[113,673,720,1346]" box="[113,546,1259,1284]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FB3AFD894F1A" authority="(Valenciennes, 1840)" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[113,546,1259,1284]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="quadriscutis">
<emphasis id="B92E4673FFE9FFAF0D35FB3AFEEA4F1D" box="[113,321,1259,1283]" italics="true" pageId="14" pageNumber="440">Notarius quadriscutis</emphasis>
(Valenciennes, 1840)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FADDFDA14F3D" blockId="14.[113,673,720,1346]" box="[113,522,1291,1315]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FADDFDA14F3D" authority="(Valenciennes, 1840)" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[113,522,1291,1315]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="rugispinis">
<emphasis id="B92E4673FFE9FFAF0D35FADDFE824F3C" box="[113,297,1291,1314]" italics="true" pageId="14" pageNumber="440">Notarius rugispinis</emphasis>
(Valenciennes, 1840)
</taxonomicName>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FAFDFE034F5C" blockId="14.[113,673,720,1346]" box="[113,424,1322,1346]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0D35FAFDFE0F4F5C" authority="(Gill, 1863)" baseAuthorityName="Gill" baseAuthorityYear="1863" box="[113,420,1322,1346]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="species" species="troschelii">
<emphasis id="B92E4673FFE9FFAF0D35FAFDFE8A4F5C" box="[113,289,1322,1346]" italics="true" pageId="14" pageNumber="440">Notarius troschelii</emphasis>
(Gill, 1863)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FA89FDCA4F88" blockId="14.[113,762,1375,1430]" pageId="14" pageNumber="440">
<emphasis id="B92E4673FFE9FFAF0D35FA89FEDB4F69" box="[113,368,1375,1399]" italics="true" pageId="14" pageNumber="440">Habitat and distribution:</emphasis>
Brackish and marine waters, eastern and western Central and South America (
<figureCitation id="136186E4FFE9FFAF0F5CFAA8FDFB4F88" box="[536,592,1406,1430]" captionStart="Figure 9" captionStartId="12.[113,178,1620,1644]" captionTargetBox="[116,1458,151,1592]" captionTargetId="figure-88@12.[114,1458,144,1593]" captionTargetPageId="12" captionText="Figure 9. Representative skulls, in dorsal view, and distribution of Cathoropsini tribe nov. A, Cathorops hypophthalmus, USNM 293275. B, Notarius grandicassis, MPEG 30647. * marine, ** brackish waters, *** freshwater." figureDoi="http://doi.org/10.5281/zenodo.11240139" httpUri="https://zenodo.org/record/11240139/files/figure.png" pageId="14" pageNumber="440">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0CC8FA78FE4B4FD6" blockId="14.[396,480,1454,1480]" box="[396,480,1454,1480]" pageId="14" pageNumber="440">
<emphasis id="B92E4673FFE9FFAF0CC8FA78FE4B4FD6" box="[396,480,1454,1480]" italics="true" pageId="14" pageNumber="440">Remarks</emphasis>
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0D35FA03FDEF4CB6" blockId="14.[113,763,1493,1705]" pageId="14" pageNumber="440">
The total-evidence analysis corroborates the monophyly, relationships, and species composition of
<taxonomicName id="4C5AE1E2FFE9FFAF0F72FA23FD274C12" authorityName="Gill" authorityYear="1863" box="[566,652,1525,1548]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFE9FFAF0F72FA23FD274C12" box="[566,652,1525,1548]" italics="true" pageId="14" pageNumber="440">Notarius</emphasis>
</taxonomicName>
as previously uncovered by molecular evidence (Betancur-R.
<emphasis id="B92E4673FFE9FFAF0F80F9C2FD504C35" box="[708,763,1555,1579]" italics="true" pageId="14" pageNumber="440">et al.</emphasis>
2007; Betancur-R. 2009). The generic names
<taxonomicName id="4C5AE1E2FFE9FFAF0F1CF9E5FD604C55" authorityName="Marceniuk &amp; Menezes" authorityYear="2007" box="[600,715,1587,1611]" family="Ariidae" genus="Amphiarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFE9FFAF0F1CF9E5FD604C55" box="[600,715,1587,1611]" italics="true" pageId="14" pageNumber="440">Amphiarius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE1E2FFE9FFAF0D35F985FF6A4C74" authorityName="D.S.Jordan &amp; Evermann" authorityYear="1898" box="[113,193,1619,1642]" family="Ariidae" genus="Aspistor" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFE9FFAF0D35F985FF6A4C74" box="[113,193,1619,1642]" italics="true" pageId="14" pageNumber="440">Aspistor</emphasis>
</taxonomicName>
, recognized as valid in previous morphological analyses (
<bibRefCitation id="EFCBE790FFE9FFAF0DF5F9A7FE494C94" author="Marceniuk AP &amp; Menezes NA" box="[177,482,1649,1674]" pageId="14" pageNumber="440" pagination="1 - 126" refId="ref30826" refString="Marceniuk AP, Menezes NA. Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera. Zootaxa 2007; 1416: 1 - 126." type="journal article" year="2007">Marceniuk and Menezes 2007</bibRefCitation>
,
<bibRefCitation id="EFCBE790FFE9FFAF0CA9F9A7FD6D4C94" author="Marceniuk AP &amp; Menezes NA &amp; Britto MR" box="[493,710,1649,1674]" pageId="14" pageNumber="440" pagination="534 - 669" refId="ref30859" refString="Marceniuk AP, Menezes NA, Britto MR. Phylogenetic analysis of the family Ariidae (Ostariophysi: Siluriformes), with a hypothesis on the monophyly and relationships of the genera. Zoological Journal of the Linnean Society 2012; 165: 534 - 669. https: // doi. org / 10.1111 / j. 1096 - 3642.2012.00822. x" type="journal article" year="2012">
Marceniuk
<emphasis id="B92E4673FFE9FFAF0F24F9A4FD254C97" box="[608,654,1649,1673]" italics="true" pageId="14" pageNumber="440">et al.</emphasis>
2012
</bibRefCitation>
), are treated herein as junior synonyms of
<taxonomicName id="4C5AE1E2FFE9FFAF0CACF944FD954CB7" authorityName="Gill" authorityYear="1863" box="[488,574,1682,1705]" family="Ariidae" genus="Notarius" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="genus">
<emphasis id="B92E4673FFE9FFAF0CACF944FD954CB7" box="[488,574,1682,1705]" italics="true" pageId="14" pageNumber="440">Notarius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE59A61FFE9FFAF0C05F917FD814CC5" blockId="14.[321,554,1729,1755]" box="[321,554,1729,1755]" pageId="14" pageNumber="440">
<emphasis id="B92E4673FFE9FFAF0C05F917FD814CC5" bold="true" box="[321,554,1729,1755]" pageId="14" pageNumber="440">
<taxonomicName id="4C5AE1E2FFE9FFAF0C05F917FE694CC5" authority="Marceniuk &amp; Oliveira &amp; Ferraris Jr, 2024" authorityName="Marceniuk &amp; Oliveira &amp; Ferraris Jr" authorityYear="2024" box="[321,450,1729,1755]" family="Ariidae" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="440" phylum="Chordata" rank="tribe" status="trib. nov." tribe="Genidentini">Genidentini</taxonomicName>
<taxonomicNameLabel id="A21DFB08FFE9FFAF0C83F917FD814CC5" box="[455,554,1729,1755]" pageId="14" pageNumber="440" rank="tribe">tribe nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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