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335 lines
45 KiB
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<mods:title id="9C22A973C6A5B4E56C7E302FFE513ED5">On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands</mods:title>
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<taxonomicName id="4C5ADDCAEF6E5E44FC2F3B56FB35F86B" ID-CoL="7PFQH" authority="Albers, 1860" authorityName="Albers" authorityYear="1860" box="[1009,1255,1917,1941]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="116" phylum="Mollusca" rank="genus">
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Albers, 1860
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<emphasis id="B92E7A5BEF6E5E44FCF33B96FC51F82C" box="[813,899,1981,2002]" italics="true" pageId="1" pageNumber="116">Rhagada</emphasis>
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Albers, 1860
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[in Martens & Albers]: 108–109 (
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species
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<emphasis id="B92E7A5BEF6E5E44FC8C3BF2FC1CF810" box="[850,974,2009,2030]" italics="true" pageId="1" pageNumber="116">Helix reinga</emphasis>
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Pfeiffer, 1846
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by original designation);
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<treatmentCitation id="0AFB8058EF6E5E44FABD3BF2FC3DF7F5" author="Pilsbry HA" page="185" pageId="1" pageNumber="116" year="1890">
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<bibRefCitation id="EFCBDBB8EF6E5E44FABD3BF2FC3DF7F5" author="Pilsbry HA" pageId="1" pageNumber="116" pagination="1 - 69" refId="ref6302" refString="Pilsbry HA (1890) Helicidae, Vol. IV. Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Second Series: Pulmonata. Conchological Section, Academy of Natural Sciences, Philadelphia. Pp. 1 - 324, pls. 1 - 69." type="book chapter" year="1890">Pilsbry, 1890: 185–191</bibRefCitation>
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</treatmentCitation>
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,
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<treatmentCitation id="0AFB8058EF6E5E44FC233BDDFB48F7F5" author="Pilsbry HA" box="[1021,1178,2038,2059]" page="135" pageId="1" pageNumber="116" year="1894">
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<bibRefCitation id="EFCBDBB8EF6E5E44FC233BDDFB48F7F5" author="Pilsbry HA" box="[1021,1178,2038,2059]" pageId="1" pageNumber="116" pagination="1 - 71" refId="ref6354" refString="Pilsbry HA (1894) Helicidae, Vol. VII. Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Second Series: Pulmonata. Conchological Section, Academy of Natural Sciences, Philadelphia. Pp. 1 - 366, pls. 1 - 71." type="book chapter" year="1894">1894: 135–136</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AFB8058EF6E5E44FB763BDDFA75F7F5" author="Martens E von" box="[1192,1447,2038,2059]" page="235" pageId="1" pageNumber="116" year="1891">
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<bibRefCitation id="EFCBDBB8EF6E5E44FB763BDDFA75F7F5" author="Martens E von" box="[1192,1447,2038,2059]" pageId="1" pageNumber="116" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens, 1891: 235–236</bibRefCitation>
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;
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<treatmentCitation id="0AFB8058EF6E5E44FC8C3438FBDCF7D6" author="Preston HB" box="[850,1038,2067,2088]" page="120" pageId="1" pageNumber="116" year="1908">
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<bibRefCitation id="EFCBDBB8EF6E5E44FC8C3438FBDCF7D6" author="Preston HB" box="[850,1038,2067,2088]" pageId="1" pageNumber="116" pagination="120" refId="ref6406" refString="Preston HB (1908) Description of a new species of Rhagada from Western Australia. Proceedings of the Malacological Society of London, 8: 120." type="journal article" year="1908">Preston, 1908: 120</bibRefCitation>
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,
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<treatmentCitation id="0AFB8058EF6E5E44FBC53438FBA6F7D6" author="Preston HB" box="[1051,1140,2067,2088]" page="13" pageId="1" pageNumber="116" year="1914">
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<bibRefCitation id="EFCBDBB8EF6E5E44FBC53438FBA6F7D6" author="Preston HB" box="[1051,1140,2067,2088]" pageId="1" pageNumber="116" pagination="13 - 18" refId="ref6434" refString="Preston HB (1914) Description of new species of land and marine shells from the Montebello Island, Western Australia. Proceedings of the Malacological Society of London, 11: 13 - 18." type="journal article" year="1914">1914: 13</bibRefCitation>
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<treatmentCitation id="0AFB8058EF6E5E44FB5C3438FA80F7D6" author="Hedley C" box="[1154,1362,2067,2088]" page="69" pageId="1" pageNumber="116" year="1916">
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<bibRefCitation id="EFCBDBB8EF6E5E44FB5C3438FA80F7D6" author="Hedley C" box="[1154,1362,2067,2088]" pageId="1" pageNumber="116" pagination="1 - 77" refId="ref5828" refString="Hedley C (1916) A preliminary index of the Mollusca of Western Australia. Journal of the Royal Society of Western Australia, 1: 1 - 77." type="journal article" year="1916">Hedley, 1916: 69–70</bibRefCitation>
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<bibRefCitation id="EFCBDBB8EF6C5E46FEFB3C94FDC2FF2A" author="Iredale T" box="[293,528,191,212]" pageId="3" pageNumber="118" pagination="83 - 124" refId="ref5858" refString="Iredale T (1938) A basic list of the land Mollusca of Australia. Part III. Australian Zoologist, 9: 83 - 124." type="journal article" year="1938">Iredale, 1938: 112–113</bibRefCitation>
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<treatmentCitation id="0AFB8058EF6C5E46FDC33C94FD4CFF2A" author="Iredale T" box="[541,670,191,212]" page="58" pageId="3" pageNumber="118" year="1939">
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<bibRefCitation id="EFCBDBB8EF6C5E46FDC33C94FD4CFF2A" author="Iredale T" box="[541,670,191,212]" pageId="3" pageNumber="118" pagination="1 - 88" refId="ref5885" refString="Iredale T (1939) A review of the land Mollusca of Western Australia. Records of the Western Australian Museum and Art Gallery, 2: 1 - 88." type="journal article" year="1939">1939: 58–63</bibRefCitation>
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<bibRefCitation id="EFCBDBB8EF6C5E46FD7F3D18FEA2FE9A" author="Johnson MS & Hamilton ZR & Murphy CE & MacLeay CA & Roberts B & Kendrick PG" pageId="3" pageNumber="118" pagination="341 - 355" refId="ref6000" refString="Johnson MS, Hamilton ZR, Murphy CE, MacLeay CA, Roberts B & Kendrick PG (2004) Evolutionary genetics of island and mainland species of Rhagada (Gastropoda: Pulmonata) in the Pilbara Region, Western Australia. Australian Journal of Zoology, 52: 341 - 355." type="journal article" year="2004">Johnson et al., 2004: 341–355</bibRefCitation>
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<treatmentCitation id="0AFB8058EF6C5E46FEA33D64FE2EFE9A" author="Johnson MS & Hamilton ZR & Fitzpatrick J" box="[381,508,335,356]" page="45" pageId="3" pageNumber="118" year="2006">
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<bibRefCitation id="EFCBDBB8EF6C5E46FEA33D64FE2EFE9A" author="Johnson MS & Hamilton ZR & Fitzpatrick J" box="[381,508,335,356]" pageId="3" pageNumber="118" pagination="45 - 50" refId="ref5964" refString="Johnson MS, Hamilton ZR & Fitzpatrick J (2006) Genetic diversity of Rhagada land snails on Barrow Island. Journal of the Royal Society of Western Australia, 89: 45 - 50." type="journal article" year="2006">2006: 45–50</bibRefCitation>
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<treatmentCitation id="0AFB8058EF6C5E46FDD73D64FD3EFE9A" author="Maassen WJM" box="[521,748,335,356]" page="65" pageId="3" pageNumber="118" year="2009">
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FDD73D64FD3EFE9A" author="Maassen WJM" box="[521,748,335,356]" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009: 65–69</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AFB8058EF6C5E46FF493D47FE51FE7F" author="Kohler F" box="[151,387,364,385]" page="270" pageId="3" pageNumber="118" year="2011">
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FF493D47FE51FE7F" author="Kohler F" box="[151,387,364,385]" pageId="3" pageNumber="118" pagination="203 - 406" refId="ref6101" refString="Kohler F (2011) The camaenid species of the Kimberley Islands, Western Australia (Stylommatophora: Helicoidea). Malacologia, 54: 203 - 406." type="journal article" year="2011">Köhler, 2011: 270–287</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AFB8058EF6C5E46FE4E3D47FD79FE7F" author="Stankowski S" box="[400,683,364,385]" page="756" pageId="3" pageNumber="118" year="2011">
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FE4E3D47FD79FE7F" author="Stankowski S" box="[400,683,364,385]" pageId="3" pageNumber="118" pagination="756 - 769" refId="ref6847" refString="Stankowski S (2011) Extreme, continuous variation in an island snail: local diversification and association of shell form with the current environment. Biological Journal of the Linnean Society, 104: 756 - 769." type="journal article" year="2011">Stankowski, 2011: 756–769</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AFB8058EF6C5E46FD663D47FED4FE60" author="Stankowski S" page="2726" pageId="3" pageNumber="118" year="2013">
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FD663D47FED4FE60" author="Stankowski S" pageId="3" pageNumber="118" pagination="2726 - 2741" refId="ref6886" refString="Stankowski S (2013) Ecological speciation in an island snail: evidence for the parallel evolution of a novel ecotype and maintenance by ecologically dependent postzygotic isolation. Molecular Ecology, 22: 2726 - 2741." type="journal article" year="2013">2013: 2726–2741</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AFB8058EF6C5E46FED33DA2FDFAFE60" author="Johnson MS & Hamilton ZR & Teale ROY & Kendrick PG" box="[269,552,393,414]" page="316" pageId="3" pageNumber="118" year="2012">
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FED33DA2FDFAFE60" author="Johnson MS & Hamilton ZR & Teale ROY & Kendrick PG" box="[269,552,393,414]" pageId="3" pageNumber="118" pagination="316 - 327" refId="ref6053" refString="Johnson MS, Hamilton ZR, Teale ROY & Kendrick PG (2012) Endemic evolutionary radiation of Rhagada land snails (Pulmonata: Camaenidae) in a continental archipelago in northern Western Australia. Biological Journal of the Linnean Society, 106: 316 - 327." type="journal article" year="2012">Johnson et al., 2012: 316–327</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Köhler & Criscione, 2013: 1971–1982.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<caption id="DF25F6C1EF6D5E47FF563BFDFA11F7BF" ID-DOI="http://doi.org/10.5281/zenodo.4502138" ID-Zenodo-Dep="4502138" httpUri="https://zenodo.org/record/4502138/files/figure.png" pageId="2" pageNumber="117" startId="2.[136,171,2006,2027]" targetBox="[143,1467,293,1913]" targetPageId="2">
|
||
<paragraph id="8BE5A649EF6D5E47FF563BFDFA11F7BF" blockId="2.[136,1475,2006,2113]" pageId="2" pageNumber="117">
|
||
Fig. 2. Penial anatomy, with penial chamber opened showing internal anatomy. A,
|
||
<taxonomicName id="4C5ADDCAEF6D5E47FC603BFDFB12F815" ID-CoL="4RYCR" authority=", Solor" authorityName="Solor" box="[958,1216,2006,2027]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="117" phylum="Mollusca" rank="species" species="solorensis">
|
||
<emphasis id="B92E7A5BEF6D5E47FC603BFDFBAFF815" box="[958,1149,2006,2027]" italics="true" pageId="2" pageNumber="117">Rhagada solorensis</emphasis>
|
||
, Solor
|
||
</taxonomicName>
|
||
Island, AM C.471154; B,
|
||
<taxonomicName id="4C5ADDCAEF6D5E47FF563BD9FE98F7F9" ID-CoL="4RYC9" authorityName="Falconieri" authorityYear="1995" box="[136,330,2034,2055]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="117" phylum="Mollusca" rank="species" species="marghitae">
|
||
<emphasis id="B92E7A5BEF6D5E47FF563BD9FE98F7F9" box="[136,330,2034,2055]" italics="true" pageId="2" pageNumber="117">Rhagada marghitae</emphasis>
|
||
</taxonomicName>
|
||
, Nusa Penida AM C.327851; C,
|
||
<taxonomicName id="4C5ADDCAEF6D5E47FD463BD9FC7CF7F9" ID-CoL="7HZXW" authority="Köhler, 2014" authorityName="Köhler" authorityYear="2014" box="[664,942,2034,2055]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="117" phylum="Mollusca" rank="subSpecies" species="setzeri" status="subsp. nov." subSpecies="atauroensis">
|
||
<emphasis id="B92E7A5BEF6D5E47FD463BD9FC7CF7F9" box="[664,942,2034,2055]" italics="true" pageId="2" pageNumber="117">Rhagada setzeri atauroensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DC720EF6D5E47FC693BD9FBC0F7F9" box="[951,1042,2034,2055]" pageId="2" pageNumber="117" rank="subSpecies">n. subsp.</taxonomicNameLabel>
|
||
, Atauro Island, holotype AM C.478282; D,
|
||
<taxonomicName id="4C5ADDCAEF6D5E47FF563424FEF5F7DA" ID-CoL="6X63X" authority="Köhler, 2014" authorityName="Köhler" authorityYear="2014" box="[136,295,2063,2084]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="117" phylum="Mollusca" rank="species" species="abbasi" status="sp. nov.">
|
||
<emphasis id="B92E7A5BEF6D5E47FF563424FEF5F7DA" box="[136,295,2063,2084]" italics="true" pageId="2" pageNumber="117">Rhagada abbasi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DC720EF6D5E47FEEE3424FEBAF7DA" box="[304,360,2063,2084]" pageId="2" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
|
||
, Lembata Island, holotype AM C.478280. Abbreviations: ep = epiphallus; fl = epiphallic flagellum; gp = genital pore; lwp = longitudinal wall pilaster; pv = penial verge; rm = penial retractor muscle; va = vagina; vd = vas deferens. Scale bars = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE5A649EF6C5E46FFAF3DE8FDB7FE0A" blockId="3.[113,753,191,565]" pageId="3" pageNumber="118">
|
||
<treatmentCitationGroup id="AB4A8167EF6C5E46FFAF3DE8FDB7FE0A" pageId="3" pageNumber="118">
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FFAF3DE8FDC5FE26" ID-CoL="7f815de1-aae9-4a64-86e4-b5431af91063" authority="Iredale, 1938: 114" authorityName="Iredale" authorityPageNumber="114" authorityYear="1938" box="[113,535,451,472]" class="Gastropoda" family="Camaenidae" genus="Bellrhagada" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="genus">
|
||
<treatmentCitation id="0AFB8058EF6C5E46FFAF3DE8FE4AFE26" author="Iredale T" box="[113,408,451,472]" page="114" pageId="3" pageNumber="118" year="1938">
|
||
<emphasis id="B92E7A5BEF6C5E46FFAF3DE8FF35FE26" box="[113,231,451,472]" italics="true" pageId="3" pageNumber="118">Bellrhagada</emphasis>
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FF323DE8FE4AFE26" author="Iredale T" box="[236,408,451,472]" pageId="3" pageNumber="118" pagination="83 - 124" refId="ref5858" refString="Iredale T (1938) A basic list of the land Mollusca of Australia. Part III. Australian Zoologist, 9: 83 - 124." type="journal article" year="1938">Iredale, 1938: 114</bibRefCitation>
|
||
</treatmentCitation>
|
||
,
|
||
<treatmentCitation id="0AFB8058EF6C5E46FE7F3DE8FDC5FE26" author="Iredale T" box="[417,535,451,472]" page="71" pageId="3" pageNumber="118" year="1939">
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FE7F3DE8FDC5FE26" author="Iredale T" box="[417,535,451,472]" pageId="3" pageNumber="118" pagination="1 - 88" refId="ref5885" refString="Iredale T (1939) A review of the land Mollusca of Western Australia. Records of the Western Australian Museum and Art Gallery, 2: 1 - 88." type="journal article" year="1939">1939: 71–72</bibRefCitation>
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54E118EBEF6C5E46FDFC3DE8FD9EFE26" box="[546,588,451,472]" pageId="3" pageNumber="118">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FD443DE8FEBBFE0A" ID-CoL="4RYCH" authority="Preston, 1914" authorityName="Preston" authorityYear="1914" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="plicata">
|
||
<emphasis id="B92E7A5BEF6C5E46FD443DE8FF08FE0A" italics="true" pageId="3" pageNumber="118">Rhagada plicata</emphasis>
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FF3C3DF4FEBBFE0A" author="Preston HB" box="[226,361,479,500]" pageId="3" pageNumber="118" pagination="13 - 18" refId="ref6434" refString="Preston HB (1914) Description of new species of land and marine shells from the Montebello Island, Western Australia. Proceedings of the Malacological Society of London, 11: 13 - 18." type="journal article" year="1914">Preston, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
, by original designation).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE5A649EF6C5E46FFAF3DD4FE34FDCB" blockId="3.[113,753,191,565]" pageId="3" pageNumber="118">
|
||
<treatmentCitationGroup id="AB4A8167EF6C5E46FFAF3DD4FE34FDCB" pageId="3" pageNumber="118">
|
||
<treatmentCitation id="0AFB8058EF6C5E46FFAF3DD4FE57FDEA" author="Iredale T" box="[113,389,511,532]" page="62" pageId="3" pageNumber="118" year="1939">
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FFAF3DD4FE57FDEA" ID-CoL="d43dd8ac-2d30-4247-9e3e-cbf50af70b56" authority="Iredale, 1939: 62" authorityName="Iredale" authorityPageNumber="62" authorityYear="1939" box="[113,389,511,532]" class="Gastropoda" family="Camaenidae" genus="Tumegada" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B92E7A5BEF6C5E46FFAF3DD4FF07FDEA" box="[113,213,511,532]" italics="true" pageId="3" pageNumber="118">Tumegada</emphasis>
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FF003DD4FE57FDEA" author="Iredale T" box="[222,389,511,532]" pageId="3" pageNumber="118" pagination="1 - 88" refId="ref5885" refString="Iredale T (1939) A review of the land Mollusca of Western Australia. Records of the Western Australian Museum and Art Gallery, 2: 1 - 88." type="journal article" year="1939">Iredale, 1939: 62</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(
|
||
<typeStatus id="54E118EBEF6C5E46FE4A3DD4FE6DFDEA" box="[404,447,511,532]" pageId="3" pageNumber="118">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FDC93DD5FF1DFDCB" ID-CoL="7RL49" authority="Cox, 1876" authorityName="Cox" authorityYear="1876" class="Gastropoda" family="Camaenidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="convicta">
|
||
<emphasis id="B92E7A5BEF6C5E46FDC93DD5FD61FDEB" box="[535,691,510,533]" italics="true" pageId="3" pageNumber="118">Helix convicta</emphasis>
|
||
Cox, 1876
|
||
</taxonomicName>
|
||
, by original designation).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BE5A649EF6C5E46FFAF3E75FDCAFA4B" blockId="3.[113,753,606,1461]" pageId="3" pageNumber="118">
|
||
<emphasis id="B92E7A5BEF6C5E46FFAF3E75FE90FD8B" bold="true" box="[113,322,606,629]" pageId="3" pageNumber="118">Taxonomic history.</emphasis>
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FE983E75FE0CFD8B" author="Solem A" box="[326,478,606,629]" pageId="3" pageNumber="118" pagination="707 - 981" refId="ref6573" refString="Solem A (1985 a) Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). V. Remaining Kimberley genera and addenda to the Kimberley. Records of the Western Australian Museum, Supplement 20: 707 - 981." type="journal article" year="1985">Solem (1985a)</bibRefCitation>
|
||
synonymised
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FDB13E75FD22FD8B" ID-CoL="7f815de1-aae9-4a64-86e4-b5431af91063" authorityName="Iredale" authorityYear="1938" box="[623,752,606,629]" class="Gastropoda" family="Camaenidae" genus="Bellrhagada" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B92E7A5BEF6C5E46FDB13E75FD22FD8B" box="[623,752,606,629]" italics="true" pageId="3" pageNumber="118">Bellrhagada</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FF7B3E55FECBFD6B" ID-CoL="d43dd8ac-2d30-4247-9e3e-cbf50af70b56" authorityName="Iredale" authorityYear="1939" box="[165,281,638,661]" class="Gastropoda" family="Camaenidae" genus="Tumegada" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B92E7A5BEF6C5E46FF7B3E55FECBFD6B" box="[165,281,638,661]" italics="true" pageId="3" pageNumber="118">Tumegada</emphasis>
|
||
</taxonomicName>
|
||
. There are about 35 species in
|
||
<collectingRegion id="499E68ABEF6C5E46FD4B3E55FF07FD4B" country="Australia" name="Western Australia" pageId="3" pageNumber="118">Western Australia</collectingRegion>
|
||
, which are comparatively well-known. Six nominal species have so far been described from the Lesser Sunda Islands.
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FF193EF5FEB2FD0B" author="Rensch B" box="[199,352,734,757]" pageId="3" pageNumber="118" pagination="361 - 396" refId="ref6470" refString="Rensch B (1931) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. I. Zoologische Jahrbucher. Abteilung fur Systematik Okologie und Geographie der Tiere, 61: 361 - 396." type="journal article" year="1931">Rensch (1931)</bibRefCitation>
|
||
grouped the species known at the time into two polytypic species (so-called “Rassenkreise”) based on shell features. He considered
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FE0A3F35FD22FCCB" ID-CoL="4RYBN" authority="(Martens, 1877)" baseAuthorityName="Martens" baseAuthorityYear="1877" box="[468,752,798,821]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="colona">
|
||
<emphasis id="B92E7A5BEF6C5E46FE0A3F35FDE9FCCB" box="[468,571,798,821]" italics="true" pageId="3" pageNumber="118">R. colona</emphasis>
|
||
(Martens, 1877)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FF433F15FDCFFCAB" ID-CoL="4RYCS" authority="(Schepman, 1892)" baseAuthorityName="Schepman" baseAuthorityYear="1892" box="[157,541,830,853]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="supracostulata">
|
||
<emphasis id="B92E7A5BEF6C5E46FF433F15FE85FCAB" box="[157,343,830,853]" italics="true" pageId="3" pageNumber="118">R. supracostulata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FEBD3F15FDC7FCAB" author="Schepman MM" box="[355,533,830,853]" pageId="3" pageNumber="118" pagination="145 - 160" refId="ref6535" refString="Schepman MM (1892) Land and freshwater shells collected by Dr. H. Ten Kate in Soemba, Timor, and other East-Indian Islands. Notes from the Leyden Museum 14: 145 - 160." type="journal article" year="1892">Schepman, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
as subspecies of the Australian “
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FF283F75FDCFFC8B" authority="(Benson, 1853)" baseAuthorityName="Benson" baseAuthorityYear="1853" box="[246,541,862,885]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="plectilis">
|
||
<emphasis id="B92E7A5BEF6C5E46FF283F75FEB8FC8B" box="[246,362,862,885]" italics="true" pageId="3" pageNumber="118">R. plectilis</emphasis>
|
||
(Benson, 1853)
|
||
</taxonomicName>
|
||
” (=
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FD8C3F75FF1AFC6B" ID-CoL="4K3QM" baseAuthorityName="Benson" baseAuthorityYear="1853" class="Gastropoda" family="Camaenidae" genus="Plectorhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="plectilis">
|
||
<emphasis id="B92E7A5BEF6C5E46FD8C3F75FF1AFC6B" italics="true" pageId="3" pageNumber="118">Plectorhagada plectilis</emphasis>
|
||
</taxonomicName>
|
||
) for having a small shell with strong axial ribs. However,
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FF3C3FB5FE80FC4B" ID-CoL="4K3QM" baseAuthorityName="Benson" baseAuthorityYear="1853" box="[226,338,926,949]" class="Gastropoda" family="Camaenidae" genus="Plectorhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="plectilis">
|
||
<emphasis id="B92E7A5BEF6C5E46FF3C3FB5FE80FC4B" box="[226,338,926,949]" italics="true" pageId="3" pageNumber="118">P. plectilis</emphasis>
|
||
</taxonomicName>
|
||
has since been shown to be part of a different genus (
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FEC73F95FE70FC2B" author="Solem A" box="[281,418,958,981]" pageId="3" pageNumber="118" pagination="1461 - 1906" refId="ref6710" refString="Solem A (1997) Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). VII. Taxa from Dampierland through the Nullabor. Records of the Western Australian Museum, Supplement 50: 1461 - 1906." type="journal article" year="1997">Solem, 1997</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FE6C3F95FD99FC2B" author="Rensch B" box="[434,587,958,981]" pageId="3" pageNumber="118" pagination="361 - 396" refId="ref6470" refString="Rensch B (1931) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. I. Zoologische Jahrbucher. Abteilung fur Systematik Okologie und Geographie der Tiere, 61: 361 - 396." type="journal article" year="1931">Rensch (1931)</bibRefCitation>
|
||
also recognised a second polytypic species,
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FE773FF5FD22FC0B" ID-CoL="4RYCR" authority="(Martens, 1863)" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[425,752,990,1013]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
|
||
<emphasis id="B92E7A5BEF6C5E46FE773FF5FDE4FC0B" box="[425,566,990,1013]" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
|
||
(Martens, 1863)
|
||
</taxonomicName>
|
||
with its subspecies
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FEE13FD5FD3EFBEB" ID-CoL="4RYBY" authority="(Martens, 1891)" baseAuthorityName="Martens" baseAuthorityYear="1891" box="[319,748,1022,1045]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="subSpecies" species="solorensis" subSpecies="floresiana">
|
||
<emphasis id="B92E7A5BEF6C5E46FEE13FD5FDE5FBEB" box="[319,567,1022,1045]" italics="true" pageId="3" pageNumber="118">R. solorensis floresiana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FD9B3FD5FD31FBEB" author="Martens E von" box="[581,739,1022,1045]" pageId="3" pageNumber="118" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, as a distinct species for its larger shell and faint axial ribs. Subsequently,
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FED13815FE6EFBAB" author="Maassen WJM" box="[271,444,1086,1109]" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
|
||
reinstated
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FDE93815FD72FBAB" ID-CoL="4RYBN" baseAuthorityName="Martens" baseAuthorityYear="1877" box="[567,672,1086,1109]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="colona">
|
||
<emphasis id="B92E7A5BEF6C5E46FDE93815FD72FBAB" box="[567,672,1086,1109]" italics="true" pageId="3" pageNumber="118">R. colona</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FD073815FEC2FB8B" ID-CoL="4RYCS" baseAuthorityName="Schepman" baseAuthorityYear="1892" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="supracostulata">
|
||
<emphasis id="B92E7A5BEF6C5E46FD073815FEC2FB8B" italics="true" pageId="3" pageNumber="118">R. supracostulata</emphasis>
|
||
</taxonomicName>
|
||
as full species but maintained the subspecies arrangement for
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FEF73855FE67FB6B" ID-CoL="4RYCR" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[297,437,1150,1173]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="solorensis">
|
||
<emphasis id="B92E7A5BEF6C5E46FEF73855FE67FB6B" box="[297,437,1150,1173]" italics="true" pageId="3" pageNumber="118">R. solorensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B92E7A5BEF6C5E46FE313855FD46FB6B" box="[495,660,1150,1173]" italics="true" pageId="3" pageNumber="118">
|
||
R. s.
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FDF53855FD46FB6B" ID-CoL="4RYBY" baseAuthorityName="Martens" baseAuthorityYear="1891" box="[555,660,1150,1173]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="floresiana">floresiana</taxonomicName>
|
||
</emphasis>
|
||
without explanation. In addition, he described a new species from Alor island,
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FF213895FDC8FB2B" ID-CoL="4RYCN" authority="Maassen, 2009" authorityName="Maassen" authorityYear="2009" box="[255,538,1214,1237]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="setzeri">
|
||
<emphasis id="B92E7A5BEF6C5E46FF213895FEB5FB2B" box="[255,359,1214,1237]" italics="true" pageId="3" pageNumber="118">R. setzeri</emphasis>
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FEAC3895FDC8FB2B" author="Maassen WJM" box="[370,538,1214,1237]" pageId="3" pageNumber="118" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
. Earlier,
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FD5C3895FF69FB0B" author="Falconieri A" pageId="3" pageNumber="118" pagination="81 - 84" refId="ref5764" refString="Falconieri A (1995) Rhagada marghitae species nova. World Shells, 14: 81 - 84." type="journal article" year="1995">Falconieri (1995)</bibRefCitation>
|
||
also described a new species from Nusa Penida,
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FFAF38D5FED7FAEB" ID-CoL="4RYC9" authorityName="Falconieri" authorityYear="1995" box="[113,261,1278,1301]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="marghitae">
|
||
<emphasis id="B92E7A5BEF6C5E46FFAF38D5FED7FAEB" box="[113,261,1278,1301]" italics="true" pageId="3" pageNumber="118">R. marghitae</emphasis>
|
||
</taxonomicName>
|
||
. However, this description was apparently overlooked subsequently. Except for the description of
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FD073935FF65FAAB" ID-CoL="4RYCN" authorityName="Maassen" authorityYear="2009" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="species" species="setzeri">
|
||
<emphasis id="B92E7A5BEF6C5E46FD073935FF65FAAB" italics="true" pageId="3" pageNumber="118">R. setzeri</emphasis>
|
||
</taxonomicName>
|
||
, all taxonomic decisions were based on shell features only. Characters considered particularly relevant for the taxonomy of
|
||
<taxonomicName id="4C5ADDCAEF6C5E46FEDE3955FEB2FA6B" ID-CoL="7PFQH" authorityName="Albers" authorityYear="1860" box="[256,352,1406,1429]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="118" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B92E7A5BEF6C5E46FEDE3955FEB2FA6B" box="[256,352,1406,1429]" italics="true" pageId="3" pageNumber="118">Rhagada</emphasis>
|
||
</taxonomicName>
|
||
species were shell size, development of axial ribs, and closure of umbilicus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340F5C2EF6C5E46FFAF39F5FDAFF7EB" pageId="3" pageNumber="118" type="diagnosis">
|
||
<paragraph id="8BE5A649EF6C5E46FFAF39F5FDAFF7EB" blockId="3.[113,753,1502,2069]" pageId="3" pageNumber="118">
|
||
<emphasis id="B92E7A5BEF6C5E46FFAF39F5FF21FA0B" bold="true" box="[113,243,1502,1525]" pageId="3" pageNumber="118">Diagnosis.</emphasis>
|
||
Shell small to medium-sized (D =
|
||
<quantity id="4CA20BACEF6C5E46FD6839F5FF72F9EB" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.5" metricValueMin="0.9" pageId="3" pageNumber="118" unit="mm" value="17.0" valueMax="25.0" valueMin="9.0">9–25 mm</quantity>
|
||
), subglobose with moderately to strongly elevated, exceptionally also low spire; umbilicus frequently concealed by greatly expanded callus, occasionally forming a chink or narrowly open; protoconch smooth to weakly sculptured by radial ridgelets or ribs; teleoconch with faint to well-developed sculpture of axial ridgelets or ribs, mostly on upper sector of whorls; whorls separated by shallow suture; periphery wellrounded to weakly angulated. Shell colour generally variable, background whitish or brownish horn, with well-developed, red subsutural, peripheral and frequently additional, thinner spiral bands. Eversible head wart between ommatophores. Penis without penial sheath, with well-developed, long and coiled epiphallus with flagellum of variable length; penial retractor muscle attached at epiphallus; epiphallus opens to lumen of penial chamber through large, cone-shaped verge with characteristic sperm groove. Bursa copulatrix short to long, simple to kinked (based on
|
||
<bibRefCitation id="EFCBDBB8EF6C5E46FE3E3BD5FDA3F7EB" author="Kohler F" box="[480,625,2046,2069]" pageId="3" pageNumber="118" pagination="203 - 406" refId="ref6101" refString="Kohler F (2011) The camaenid species of the Kimberley Islands, Western Australia (Stylommatophora: Helicoidea). Malacologia, 54: 203 - 406." type="journal article" year="2011">Köhler, 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |