193 lines
23 KiB
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193 lines
23 KiB
XML
<document id="341E70C7D1BE4C12037DF8CF51E04B2B" ID-CLB-Dataset="5029" ID-DOI="10.1649/0010-065X-72.2.219" ID-GBIF-Dataset="a4c1b33e-e026-4c37-8632-04b8402ff337" ID-ISSN="1938-4394" ID-Zenodo-Dep="4788883" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1621946431230" checkinUser="carolina" docAuthor="Parker, Joseph & Owens, Brittany" docDate="2018" docId="03E92E0EFFDD1F66FD25FBE4FC9AFED2" docLanguage="en" docName="TheColeopBull.72.2.219-229.pdf" docOrigin="The Coleopterists Bulletin 72 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-72.2.219" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Batriscydmaenus PARKER AND OWENS 2018" docType="treatment" docVersion="5" lastPageNumber="225" masterDocId="FFD05676FFD81F60FFE0FFADFFA3FFF4" masterDocTitle="Batriscydmaenus Parker and Owens, New Genus, and Convergent Evolution of a “ Reductive ” Ecomorph in Socially Symbiotic Pselaphinae (Coleoptera: Staphylinidae)" masterLastPageNumber="229" masterPageNumber="219" pageNumber="224" updateTime="1698989665874" updateUser="ExternalLinkService">
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<mods:title id="DC10F5109006287BDF073291B00D5A00">Batriscydmaenus Parker and Owens, New Genus, and Convergent Evolution of a “ Reductive ” Ecomorph in Socially Symbiotic Pselaphinae (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="AA16CCC1D60419959F65C96A94068C2A">Parker, Joseph</mods:namePart>
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<mods:namePart id="A4953BFF24D65616CAD4AD6E445F2F8E">Owens, Brittany</mods:namePart>
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<mods:title id="922673AB5FFCD6D4B3A921FE383DDDCB">The Coleopterists Bulletin</mods:title>
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<mods:part id="47F9FCB48036B58AB5B8660815F272FB">
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<mods:date id="B8DD97C6CA99AF7A4954CB0A44DE206B">2018</mods:date>
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<mods:number id="58343BB1CD006A9EF0C74DBB132290DC">2018-06-20</mods:number>
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<mods:number id="46AE032FC155BECBD3E173FF92D8AE5A">72</mods:number>
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<treatment id="03E92E0EFFDD1F66FD25FBE4FC9AFED2" ID-GBIF-Taxon="184009671" LSID="urn:lsid:zoobank.org:act:3AB8EB86-87CD-472E-9B22-E9E0A36ECAC1" httpUri="http://treatment.plazi.org/id/03E92E0EFFDD1F66FD25FBE4FC9AFED2" lastPageId="6" lastPageNumber="225" pageId="5" pageNumber="224">
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<paragraph id="8BFF9F18FFDD1F66FD25FBE4FEBDFED2" blockId="5.[685,1173,194,1730]" lastBlockId="6.[173,660,194,1091]" lastPageId="6" lastPageNumber="225" pageId="5" pageNumber="224">
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<subSubSection id="C35ACC93FFDD1F65FD25FBE4FBC9FBAA" box="[709,1130,1097,1118]" pageId="5" pageNumber="224" type="nomenclature">
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<heading id="D0B72874FFDD1F65FD25FBE4FBC9FBAA" box="[709,1130,1097,1118]" pageId="5" pageNumber="224">
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<emphasis id="B934430AFFDD1F65FD25FBE4FBC9FBAA" bold="true" box="[709,1130,1097,1118]" pageId="5" pageNumber="224">
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Phylogenetic Position of
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<taxonomicName id="4C40E49BFFDD1F65FC5AFBE4FBC7FBAA" ID-CoL="8XJY9" authorityName="PARKER AND OWENS" authorityYear="2018" box="[954,1124,1097,1118]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDD1F65FC5AFBE4FBC7FBAA" bold="true" box="[954,1124,1097,1118]" italics="true" pageId="5" pageNumber="224">Batriscydmaenus</emphasis>
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</taxonomicName>
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.
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</emphasis>
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</heading>
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</subSubSection>
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<subSubSection id="C35ACC93FFDD1F66FB8FFBE4FEBDFED2" lastPageId="6" lastPageNumber="225" pageId="5" pageNumber="224" type="discussion">
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Our initial impression of
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<taxonomicName id="4C40E49BFFDD1F65FC90FBC9FBB0FB8D" authorityName="PARKER AND OWENS" authorityYear="2018" box="[880,1043,1124,1145]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDD1F65FC90FBC9FBB0FB8D" box="[880,1043,1124,1145]" italics="true" pageId="5" pageNumber="224">Batriscydmaenus</emphasis>
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</taxonomicName>
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was of a new and highly unusual genus, almost scydmaenine-like in form, but with shortened elytra and threesegmented tarsi that suggested a placement in
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<taxonomicName id="4C40E49BFFDD1F65FD4DFB63FC82FB17" baseAuthorityName="Chandler" baseAuthorityYear="2001" box="[685,801,1230,1251]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="224" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
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. The new taxon appeared to be highly atypical for a pselaphine, however, with several remarkable character states that are otherwise invariant within the subfamily. The overtly simplified morphology, with profound losses of foveae, sulci, and other characters used routinely for tribal placement, meant that unambiguous external characters to link the new genus to a higher taxonomic group were not forthcoming. One character—the apically notched antennal scape—suggested possible membership in the supertribe
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<taxonomicName id="4C40E49BFFDD1F65FBEAFA75FBCAFA19" authorityName="Reitter" authorityYear="1882" box="[1034,1129,1496,1517]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="224" phylum="Arthropoda" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
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, but the approximation of this character state elsewhere in
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<taxonomicName id="4C40E49BFFDD1F65FD2EF9A0FC5AF9D6" authority="(Kurbatov 2007)" baseAuthorityName="Kurbatov" baseAuthorityYear="2007" box="[718,1017,1549,1570]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="224" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">
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Pselaphinae (
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<bibRefCitation id="EFD1E2E9FFDD1F65FCB6F9A0FC51F9D6" author="Kurbatov, S. A." box="[854,1010,1549,1570]" pageId="5" pageNumber="224" pagination="275 - 289" refId="ref7228" refString="Kurbatov, S. A. 2007. Revision of the genus Intestinarius gen. n. from Southeast Asia, with notes on a probable autapomorphy of Batrisitae (Coleoptera: Staphylinidae: Pselaphinae). Russian Entomological Journal 16: 275 - 289." type="journal article" year="2007">Kurbatov 2007</bibRefCitation>
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)
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</taxonomicName>
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prevented unequivocal placement. Batrisites possess a modified metaventrite margin that is curved at the midline to form a preapical fovea (
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<bibRefCitation id="EFD1E2E9FFDD1F65FC23F9F0FBF8F986" author="Chandler, D. S." box="[963,1115,1629,1650]" pageId="5" pageNumber="224" refId="ref6632" refString="Chandler, D. S. 2001. Biology, Morphology and Systematics of the Ant-Like Litter Beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology, International, Associated Publishers, Gainesville, FL." type="book" year="2001">Chandler 2001</bibRefCitation>
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), and we noticed that
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<taxonomicName id="4C40E49BFFDD1F65FCBDF9D5FBA7F979" authorityName="PARKER AND OWENS" authorityYear="2018" box="[861,1028,1656,1677]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="224" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDD1F65FCBDF9D5FBA7F979" box="[861,1028,1656,1677]" italics="true" pageId="5" pageNumber="224">Batriscydmaenus</emphasis>
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</taxonomicName>
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has this feature. Furthermore, upon dissection we discovered the genus possesses four setae on the labrum—a proposed autapomorphy of
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<taxonomicName id="4C40E49BFFDE1F66FE26FF6FFF4AFF05" authority="(Kurbatov 2007)" authorityName="Reitter" authorityYear="1882" baseAuthorityName="Kurbatov" baseAuthorityYear="2007" class="Insecta" family="Staphylinidae" genus="Batrisitae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="genus" superTribe="Batrisitae">
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Batrisitae (
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<bibRefCitation id="EFD1E2E9FFDE1F66FDD4FF6FFF40FF05" author="Kurbatov, S. A." pageId="6" pageNumber="225" pagination="275 - 289" refId="ref7228" refString="Kurbatov, S. A. 2007. Revision of the genus Intestinarius gen. n. from Southeast Asia, with notes on a probable autapomorphy of Batrisitae (Coleoptera: Staphylinidae: Pselaphinae). Russian Entomological Journal 16: 275 - 289." type="journal article" year="2007">Kurbatov 2007</bibRefCitation>
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)
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</taxonomicName>
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. Still, the radical departure from the typical pselaphine (and batrisine) habitus was perplexing.
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</subSubSection>
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</paragraph>
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<paragraph id="8BFF9F18FFDE1F66FF25FE81FC9AFED2" blockId="6.[173,660,194,1091]" lastBlockId="6.[707,1193,194,294]" pageId="6" pageNumber="225">
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To circumvent the challenge posed by the morphology of the new genus to a definitive tribal placement, we assessed the phylogenetic position of
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<taxonomicName id="4C40E49BFFDE1F66FF4DFED1FEF3FE65" authorityName="PARKER AND OWENS" authorityYear="2018" box="[173,336,380,401]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDE1F66FF4DFED1FEF3FE65" box="[173,336,380,401]" italics="true" pageId="6" pageNumber="225">Batriscydmaenus</emphasis>
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</taxonomicName>
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molecularly. We recovered fragments of nuclear
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<emphasis id="B934430AFFDE1F66FEB5FE3BFE17FE5F" box="[341,436,406,427]" italics="true" pageId="6" pageNumber="225">28s rRNA</emphasis>
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and mitochondrial
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<emphasis id="B934430AFFDE1F66FD92FE3BFF40FE32" italics="true" pageId="6" pageNumber="225">16s rRNA</emphasis>
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and COI from one of the
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<typeStatus id="54FB21BAFFDE1F66FE36FE1CFD97FE32" box="[470,564,433,454]" pageId="6" pageNumber="225" type="paratype">paratypes</typeStatus>
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. Bayesian analysis revealed that
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<taxonomicName id="4C40E49BFFDE1F66FE74FE66FD94FE14" authorityName="PARKER AND OWENS" authorityYear="2018" box="[404,567,459,480]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDE1F66FE74FE66FD94FE14" box="[404,567,459,480]" italics="true" pageId="6" pageNumber="225">Batriscydmaenus</emphasis>
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</taxonomicName>
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emerges within a maximally supported
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<taxonomicName id="4C40E49BFFDE1F66FE17FE4BFDF2FE0F" authorityName="Reitter" authorityYear="1882" box="[503,593,486,507]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
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clade (
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<figureCitation id="137B839DFFDE1F66FF54FDADFF4EFDE1" box="[180,237,512,533]" captionStart="Fig" captionStartId="6.[197,227,1663,1682]" captionTargetBox="[177,655,1152,1644]" captionTargetId="figure-638@6.[175,657,1150,1646]" captionTargetPageId="6" captionText="Fig. 4. Bayesian consensus tree showing phylogenetic position of Batriscydmaenus within Batrisitae. Numbers above branches are posterior probabilities." figureDoi="http://doi.org/10.5281/zenodo.4788961" httpUri="https://zenodo.org/record/4788961/files/figure.png" pageId="6" pageNumber="225">Fig. 4</figureCitation>
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). These results establish with confidence that the new taxon is indeed a morphologically derived member of
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<taxonomicName id="4C40E49BFFDE1F66FECCFD98FE31FDBE" authorityName="Reitter" authorityYear="1882" box="[300,402,565,586]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="224" phylum="Arthropoda" rank="superTribe" superTribe="Batrisitae">Batrisitae</taxonomicName>
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. Our tree includes nine members of
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<taxonomicName id="4C40E49BFFDE1F66FEC1FDFDFED1FD91" authorityName="Reitter" authorityYear="1882" box="[289,370,592,613]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
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as well as
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<taxonomicName id="4C40E49BFFDE1F66FE36FDFDFDDCFD91" authority="Brendel" authorityName="Brendel" authorityYear="1893" box="[470,639,592,613]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDE1F66FE36FDFDFD8EFD91" box="[470,557,592,613]" italics="true" pageId="6" pageNumber="225">Arianops</emphasis>
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Brendel
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</taxonomicName>
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, a Nearctic member of the troglobitic batrisite tribe
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<taxonomicName id="4C40E49BFFDE1F66FF4DFD28FE80FD6E" box="[173,291,645,666]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Amauropini">Amauropini</taxonomicName>
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. The position of
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<taxonomicName id="4C40E49BFFDE1F66FE3CFD28FD90FD6E" authorityName="Brendel" authorityYear="1893" box="[476,563,645,666]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDE1F66FE3CFD28FD90FD6E" box="[476,563,645,666]" italics="true" pageId="6" pageNumber="225">Arianops</emphasis>
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</taxonomicName>
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within a clade otherwise composed of
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<taxonomicName id="4C40E49BFFDE1F66FE35FD0DFD85FD41" authorityName="Reitter" authorityYear="1882" box="[469,550,672,693]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
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genera indicates that
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<taxonomicName id="4C40E49BFFDE1F66FEC9FD17FE38FD3B" box="[297,411,698,719]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Amauropini">Amauropini</taxonomicName>
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should be synonymized with
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<taxonomicName id="4C40E49BFFDE1F66FF06FD78FE99FD1E" authorityName="Reitter" authorityYear="1882" box="[230,314,725,746]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
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, as suggested previously (
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<bibRefCitation id="EFD1E2E9FFDE1F66FDB4FD78FF4FFCF1" author="Parker, J." pageId="6" pageNumber="225" pagination="541 - 566" refId="ref7637" refString="Parker, J. 2016 b. Emergence of a superradiation: Pselaphine rove beetles in mid-Cretaceous amber from Myanmar and their evolutionary implications. Systematic Entomology 41: 541 - 566." type="journal article" year="2016">Parker 2016b</bibRefCitation>
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). We refrain from doing so until a greater range of amauropine genera can be sampled, but nevertheless conclude that
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<taxonomicName id="4C40E49BFFDE1F66FE24FC88FDC4FCCE" authorityName="PARKER AND OWENS" authorityYear="2018" box="[452,615,805,826]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDE1F66FE24FC88FDC4FCCE" box="[452,615,805,826]" italics="true" pageId="6" pageNumber="225">Batriscydmaenus</emphasis>
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</taxonomicName>
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belongs within
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<taxonomicName id="4C40E49BFFDE1F66FEDCFC92FE2EFCA0" authorityName="Reitter" authorityYear="1882" box="[316,397,831,852]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
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rather than
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<taxonomicName id="4C40E49BFFDE1F66FDFAFC92FD33FCA0" box="[538,656,831,852]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Amauropini">Amauropini</taxonomicName>
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. Notably, batrisines have undergone explosive genus-level diversification in the African and, in particular, the East Asian tropics, but up to now only six genera have been recorded from the Neotropics:
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<taxonomicName id="4C40E49BFFDE1F66FF4DFC69FEC9FC2D" authority="LeConte" authorityName="LeConte" box="[173,362,964,985]" class="Insecta" family="Staphylinidae" genus="Arthmius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
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<emphasis id="B934430AFFDE1F66FF4DFC69FEA7FC2D" box="[173,260,964,985]" italics="true" pageId="6" pageNumber="225">Arthmius</emphasis>
|
|
LeConte
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C40E49BFFDE1F66FE61FC69FD94FC2D" authority="Sharp" authorityName="Sharp" authorityYear="1887" box="[385,567,964,985]" class="Insecta" family="Staphylinidae" genus="Batoctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FE61FC69FE49FC2D" box="[385,490,964,985]" italics="true" pageId="6" pageNumber="225">Batoctenus</emphasis>
|
|
Sharp
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C40E49BFFDE1F66FDAEFC69FF55FC00" authority="Raffray" authorityName="Raffray" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Iteticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FDAEFC69FD30FC2D" box="[590,659,964,985]" italics="true" pageId="6" pageNumber="225">Iteticus</emphasis>
|
|
Raffray
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C40E49BFFDE1F66FEE3FC72FE14FC00" authority="Reitter" authorityName="Reitter" authorityYear="1883" box="[259,439,991,1012]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FEE3FC72FEC8FC00" box="[259,363,991,1012]" italics="true" pageId="6" pageNumber="225">Oxarthrius</emphasis>
|
|
Reitter
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C40E49BFFDE1F66FE24FC72FDC0FC00" authority="Reitter" authorityName="Reitter" authorityYear="1882" box="[452,611,991,1012]" class="Insecta" family="Staphylinidae" genus="Syrbatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FE24FC72FDB4FC00" box="[452,535,991,1012]" italics="true" pageId="6" pageNumber="225">Syrbatus</emphasis>
|
|
Reitter
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="4C40E49BFFDE1F66FF4DFC54FE45FBFA" authority="Raffray" authorityName="Raffray" authorityYear="1898" box="[173,486,1017,1038]" class="Insecta" family="Staphylinidae" genus="Syrmocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FF4DFC54FEBEFBFA" box="[173,285,1017,1038]" italics="true" pageId="6" pageNumber="225">Syrmocerus</emphasis>
|
|
Raffray (
|
|
<bibRefCitation id="EFD1E2E9FFDE1F66FE98FC54FE7DFBFA" author="Park, O." box="[376,478,1017,1038]" pageId="6" pageNumber="225" refId="ref7521" refString="Park, O. 1942. A Study in Neotropical Pselaphidae. Northwestern University, Evanston, IL." type="book" year="1942">Park 1942</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
.
|
|
<taxonomicName id="4C40E49BFFDE1F66FE10FC54FD30FBFA" authorityName="PARKER AND OWENS" authorityYear="2018" box="[496,659,1017,1038]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FE10FC54FD30FBFA" box="[496,659,1017,1038]" italics="true" pageId="6" pageNumber="225">Batriscydmaenus</emphasis>
|
|
</taxonomicName>
|
|
thus represents the seventh Neotropical batrisine genus. In our tree,
|
|
<taxonomicName id="4C40E49BFFDE1F66FE85FB83FDABFBB7" authorityName="PARKER AND OWENS" authorityYear="2018" box="[357,520,1070,1091]" class="Insecta" family="Staphylinidae" genus="Batriscydmaenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FE85FB83FDABFBB7" box="[357,520,1070,1091]" italics="true" pageId="6" pageNumber="225">Batriscydmaenus</emphasis>
|
|
</taxonomicName>
|
|
is resolved as sister to the Neotropical genus
|
|
<taxonomicName id="4C40E49BFFDE1F66FBEFFF6FFBD4FF23" authorityName="Reitter" authorityYear="1883" box="[1039,1143,194,215]" class="Insecta" family="Staphylinidae" genus="Oxarthrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B934430AFFDE1F66FBEFFF6FFBD4FF23" box="[1039,1143,194,215]" italics="true" pageId="6" pageNumber="225">Oxarthrius</emphasis>
|
|
</taxonomicName>
|
|
, but limited gene and taxon sampling precludes a more exact placement within
|
|
<taxonomicName id="4C40E49BFFDE1F66FC12FF5AFBEAFEF8" authorityName="Reitter" authorityYear="1882" box="[1010,1097,247,268]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="225" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini</taxonomicName>
|
|
with any confidence.
|
|
</paragraph>
|
|
</treatment>
|
|
</document> |