treatments-xml/data/03/E3/87/03E387DABC2AFF8EFF31FC10FB0FFD5A.xml

<document id="BE132EE8E94E89EF8AA934EC8AA4475B" ID-DOI="10.5281/zenodo.197318" ID-GBIF-Dataset="171ad498-8a10-49d9-8b0c-15f886bb682d" ID-ISSN="1175-5326" ID-Zenodo-Dep="197318" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460266585607" checkinUser="plazi" docAuthor="Rowley, Jodi J. L., Stuart, Bryan L., Thy, Neang &amp; Emmett, David A." docDate="2010" docId="03E387DABC2AFF8EFF31FC10FB0FFD5A" docLanguage="en" docName="zt02567p068.pdf" docOrigin="Zootaxa 2567" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptolalax melicus Rowley, Stuart, Thy &amp; Emmett, 2010, sp. nov." docType="treatment" docVersion="13" lastPageNumber="66" masterDocId="FFDAFFA2BC28FF87FFA6FFB7FFEFFFB8" masterDocTitle="A new species of Leptolalax (Anura: Megophryidae) from northeastern Cambodia" masterLastPageNumber="68" masterPageNumber="57" pageNumber="59" updateTime="1698245923217" updateUser="plazi">
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<mods:title id="7819050BA652DF0086DDBA9083BC48BB">A new species of Leptolalax (Anura: Megophryidae) from northeastern Cambodia</mods:title>
</mods:titleInfo>
<mods:name id="F8EDD041CE7F8885E2B0D76DFFFCAE60" type="personal">
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<mods:namePart id="0938EAA4FE15669435EECA44FF95F802">Rowley, Jodi J. L.</mods:namePart>
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<mods:namePart id="8D98E9086F904F02692F0B02C7D3A64C">Stuart, Bryan L.</mods:namePart>
</mods:name>
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<mods:namePart id="1F4551D4E01E2C62DB3A82A77E4FFE4F">Thy, Neang</mods:namePart>
</mods:name>
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<mods:namePart id="41BA3D483AD6CA9460D9792D00006303">Emmett, David A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4271F3DDB4BCE5E7C494B1BDEBE67037">text</mods:typeOfResource>
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<mods:title id="2C390E937370A6E33F04B01B6C8A215F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D386CFAC5948DD99C97B45E97AE7F235">
<mods:date id="F85A509CD3C35EED49619DD703D7006E">2010</mods:date>
<mods:detail id="886BC4A92D0BF5A5A58A4C56FFFDA6A3" type="volume">
<mods:number id="492489294634E2AB05D93AFABAC4AB22">2567</mods:number>
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<mods:identifier id="2A7A429C5F065D854AE7B6535EDB53B3" type="DOI">10.5281/zenodo.197318</mods:identifier>
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<mods:identifier id="3293E04B4096311BCA9EAE5B61F03F2B" type="Zenodo-Dep">197318</mods:identifier>
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<treatment id="03E387DABC2AFF8EFF31FC10FB0FFD5A" ID-DOI="http://doi.org/10.5281/zenodo.5629033" ID-GBIF-Taxon="119394313" ID-Zenodo-Dep="5629033" LSID="urn:lsid:plazi:treatment:03E387DABC2AFF8EFF31FC10FB0FFD5A" httpUri="http://treatment.plazi.org/id/03E387DABC2AFF8EFF31FC10FB0FFD5A" lastPageId="9" lastPageNumber="66" pageId="2" pageNumber="59">
<subSubSection id="C3506547BC2AFF85FF31FC10FE0FFC79" box="[151,480,935,961]" pageId="2" pageNumber="59" type="nomenclature">
<paragraph id="8BF536CCBC2AFF85FF31FC10FE0FFC79" blockId="2.[151,480,935,961]" box="[151,480,935,961]" pageId="2" pageNumber="59">
<heading id="D0BD81A0BC2AFF85FF31FC10FE0FFC79" bold="true" box="[151,480,935,961]" fontSize="11" level="1" pageId="2" pageNumber="59" reason="1">
<emphasis id="B93EEADEBC2AFF85FF31FC10FE0FFC79" bold="true" box="[151,480,935,961]" pageId="2" pageNumber="59">
<taxonomicName id="4C4A4D4FBC2AFF85FF31FC10FE93FC79" ID-CoL="3TQQC" box="[151,380,935,961]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="melicus" status="sp. nov.">
<emphasis id="B93EEADEBC2AFF85FF31FC10FE93FC79" bold="true" box="[151,380,935,961]" italics="true" pageId="2" pageNumber="59">Leptolalax melicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57A5BC2AFF85FE25FC10FE0FFC79" box="[387,480,935,961]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3506547BC2AFF85FF31FC58FCEBFAE2" pageId="2" pageNumber="59" type="materials_examined">
<paragraph id="8BF536CCBC2AFF85FF31FC58FA78FBEC" blockId="2.[151,1437,1007,2004]" pageId="2" pageNumber="59">
<emphasis id="B93EEADEBC2AFF85FF31FC58FEFDFBB1" bold="true" box="[151,274,1007,1033]" pageId="2" pageNumber="59">
<typeStatus id="54F1886EBC2AFF85FF31FC58FEE1FBB1" box="[151,270,1007,1033]" pageId="2" pageNumber="59" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED5BAE09BC2AFF85FEBCFC47FEB4FBB2" LSID="urn:lsid:biocol.org:col:34777" box="[282,347,1008,1034]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="2" pageNumber="59">MVZ</collectionCode>
258198, an adult male, calling on 
<quantity id="4CB29B29BC2AFF85FD48FC47FCC7FBB2" box="[750,808,1008,1034]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="59" unit="cm" value="2.0">2 cm</quantity>
diameter tree root, 
<quantity id="4CB29B29BC2AFF85FBAAFC47FBA0FBB2" box="[1036,1103,1008,1034]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="59" unit="m" value="0.5">0.5 m</quantity>
from 
<quantity id="4CB29B29BC2AFF85FB30FC47FB2DFBB2" box="[1174,1218,1008,1034]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="59" unit="m" value="2.0">2 m</quantity>
wide, swift, rocky stream in Virachey National Park, Ratanakiri Province, 
<collectingCountry id="F35D765CBC2AFF85FCF6FBA2FC21FB97" box="[848,974,1045,1071]" name="Cambodia" pageId="2" pageNumber="59">Cambodia</collectingCountry>
(
<geoCoordinate id="EE7E500BBC2AFF85FC79FBA2FB6FFB97" box="[991,1152,1045,1071]" direction="north" orientation="latitude" pageId="2" pageNumber="59" precision="1" value="14.19287">14.19287º N</geoCoordinate>
, 
<geoCoordinate id="EE7E500BBC2AFF85FB2BFBA2FAD6FB97" box="[1165,1337,1045,1071]" direction="east" orientation="longitude" pageId="2" pageNumber="59" precision="1" value="106.99612">106.99612º E</geoCoordinate>
, 
<quantity id="4CB29B29BC2AFF85FAE0FBA2FA77FB97" box="[1350,1432,1045,1071]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="59" unit="m" value="650.0">650 m</quantity>
, 
<figureCitation id="13712A49BC2AFF85FF31FB8DFECAFBEC" box="[151,293,1082,1108]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,257,1800,1824]" captionStartId-1="4.[151,255,1872,1896]" captionTargetBox-0="[151,1436,645,1776]" captionTargetBox-1="[151,1436,178,1850]" captionTargetId-0="figure@3.[151,1436,645,1777]" captionTargetId-1="figure@4.[151,1436,176,1851]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Collection sites of Leptolalax melicus sp. nov. from Virachey National Park, Cambodia (black circles), Leptolalax applebyi from Song Thanh Proposed Nature Reserve, Vietnam (dark grey square; Rowley &amp; Cao 2009) and Leptolalax sp. from the Cardamom Mountains, Cambodia (dark grey triangle; Ohler et al. 2002). Areas over 600 m elevation are pale grey. Boundaries of Virachey National Park (hatched area) were obtained from the World Database on Protected Areas 2009." captionText-1="FIGURE 2. Male holotype of Leptolalax melicus sp. nov. (MVZ 258198) in life. (A) Lateral view in the field, in calling posture (nocturnal ventral colouration), (B) dorsal and (C) ventral views (diurnal ventral colouration)." httpUri-0="https://zenodo.org/record/197319/files/figure.png" httpUri-1="https://zenodo.org/record/197320/files/figure.png" pageId="2" pageNumber="59">Figures 1–2</figureCitation>
). Collected by Jodi J. L. Rowley, Bryan L. Stuart and Neang Thy at 19:35 h on 
<date id="FFF4100CBC2AFF85FB6CFB8DFA7CFBEC" box="[1226,1427,1082,1108]" pageId="2" pageNumber="59" value="2007-10-13">13 October 2007</date>
.
</paragraph>
<paragraph id="8BF536CCBC2AFF85FF63FBE8FCEBFAE2" blockId="2.[151,1437,1007,2004]" pageId="2" pageNumber="59">
<emphasis id="B93EEADEBC2AFF85FF63FBE8FEA3FBC1" bold="true" box="[197,332,1119,1145]" pageId="2" pageNumber="59">
<typeStatus id="54F1886EBC2AFF85FF63FBE8FEA8FBC1" box="[197,327,1119,1145]" pageId="2" pageNumber="59" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="ED5BAE09BC2AFF85FEF5FBD7FE7BFBC2" LSID="urn:lsid:biocol.org:col:34777" box="[339,404,1120,1146]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="2" pageNumber="59">MVZ</collectionCode>
258199, an adult male, same data as 
<typeStatus id="54F1886EBC2AFF85FCE2FBD7FC45FBC2" box="[836,938,1120,1146]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
except collected at 19:40 h on 
<date id="FFF4100CBC2AFF85FAB2FBD7FA78FBC2" box="[1300,1431,1120,1146]" pageId="2" pageNumber="59" value="1913-10">13 October</date>
, 2007. 
<collectionCode id="ED5BAE09BC2AFF85FF46FB32FECEFB27" LSID="urn:lsid:biocol.org:col:34777" box="[224,289,1157,1183]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="2" pageNumber="59">MVZ</collectionCode>
258197, adult male, same data as 
<typeStatus id="54F1886EBC2AFF85FD15FB32FCF5FB27" box="[691,794,1157,1183]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
except collected at 19:40 h on 
<date id="FFF4100CBC2AFF85FB21FB32FAA0FB27" box="[1159,1359,1157,1183]" pageId="2" pageNumber="59" value="2007-10-10">10 October 2007</date>
. 
<collectionCode id="ED5BAE09BC2AFF85FAFCFB32FA74FB27" LSID="urn:lsid:biocol.org:col:34777" box="[1370,1435,1157,1183]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="2" pageNumber="59">MVZ</collectionCode>
258074, adult male, calling from boulder, collected by Jodi J. L. Rowley on 
<date id="FFF4100CBC2AFF85FBE6FB1DFB0BFB7C" box="[1088,1252,1194,1220]" pageId="2" pageNumber="59" value="2006-06-22">22 June 2006</date>
from Virachey National Park, Ratanakiri Province, 
<collectingCountry id="F35D765CBC2AFF85FDF3FB67FD3DFB52" box="[597,722,1232,1258]" name="Cambodia" pageId="2" pageNumber="59">Cambodia</collectingCountry>
(ca. 
<geoCoordinate id="EE7E500BBC2AFF85FCB6FB67FC9EFB52" box="[784,881,1232,1258]" direction="north" orientation="latitude" pageId="2" pageNumber="59" precision="5555" value="14.3">14.3º N</geoCoordinate>
, 
<geoCoordinate id="EE7E500BBC2AFF85FCD8FB67FC16FB52" box="[894,1017,1232,1258]" direction="east" orientation="longitude" pageId="2" pageNumber="59" precision="555" value="107.37">107.37º E</geoCoordinate>
, 
<quantity id="4CB29B29BC2AFF85FBA1FB67FBB8FB52" box="[1031,1111,1232,1258]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="59" unit="m" value="600.0">600 m</quantity>
, 
<figureCitation id="13712A49BC2AFF85FBC3FB67FB24FB52" box="[1125,1227,1232,1258]" captionStart="FIGURE 1" captionStartId="3.[151,257,1800,1824]" captionTargetBox="[151,1436,645,1776]" captionTargetId="figure@3.[151,1436,645,1777]" captionTargetPageId="3" captionText="FIGURE 1. Collection sites of Leptolalax melicus sp. nov. from Virachey National Park, Cambodia (black circles), Leptolalax applebyi from Song Thanh Proposed Nature Reserve, Vietnam (dark grey square; Rowley &amp; Cao 2009) and Leptolalax sp. from the Cardamom Mountains, Cambodia (dark grey triangle; Ohler et al. 2002). Areas over 600 m elevation are pale grey. Boundaries of Virachey National Park (hatched area) were obtained from the World Database on Protected Areas 2009." httpUri="https://zenodo.org/record/197319/files/figure.png" pageId="2" pageNumber="59">Figure 1</figureCitation>
). 
<collectionCode id="ED5BAE09BC2AFF85FB41FB67FAC5FB52" LSID="urn:lsid:biocol.org:col:34777" box="[1255,1322,1232,1258]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="2" pageNumber="59">MVZ</collectionCode>
258075– 258077, adult males, all calling on rocks, collected by David A. Emmett on 
<date id="FFF4100CBC2AFF85FB99FB42FB0BFAB7" box="[1087,1252,1269,1295]" pageId="2" pageNumber="59" value="2006-06-22">22 June 2006</date>
from Virachey National Park, Ratanakiri Province, 
<collectingCountry id="F35D765CBC2AFF85FDE7FAADFD56FA8C" box="[577,697,1306,1332]" name="Cambodia" pageId="2" pageNumber="59">Cambodia</collectingCountry>
(ca. 
<geoCoordinate id="EE7E500BBC2AFF85FD52FAADFCBFFA8C" box="[756,848,1306,1332]" direction="north" orientation="latitude" pageId="2" pageNumber="59" precision="5555" value="14.2">14.2º N</geoCoordinate>
, 
<geoCoordinate id="EE7E500BBC2AFF85FCFAFAADFC3CFA8C" box="[860,979,1306,1332]" direction="east" orientation="longitude" pageId="2" pageNumber="59" precision="555" value="107.38">107.38º E</geoCoordinate>
, 
<quantity id="4CB29B29BC2AFF85FC79FAADFBC3FA8C" box="[991,1068,1306,1332]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="59" unit="m" value="850.0">850 m</quantity>
, 
<figureCitation id="13712A49BC2AFF85FB9EFAADFB74FA8C" box="[1080,1179,1306,1332]" captionStart="FIGURE 1" captionStartId="3.[151,257,1800,1824]" captionTargetBox="[151,1436,645,1776]" captionTargetId="figure@3.[151,1436,645,1777]" captionTargetPageId="3" captionText="FIGURE 1. Collection sites of Leptolalax melicus sp. nov. from Virachey National Park, Cambodia (black circles), Leptolalax applebyi from Song Thanh Proposed Nature Reserve, Vietnam (dark grey square; Rowley &amp; Cao 2009) and Leptolalax sp. from the Cardamom Mountains, Cambodia (dark grey triangle; Ohler et al. 2002). Areas over 600 m elevation are pale grey. Boundaries of Virachey National Park (hatched area) were obtained from the World Database on Protected Areas 2009." httpUri="https://zenodo.org/record/197319/files/figure.png" pageId="2" pageNumber="59">Figure 1</figureCitation>
). All specimens were found &lt;
<quantity id="4CB29B29BC2AFF85FF55FAF7FEC0FAE2" box="[243,303,1344,1370]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="59" unit="m" value="10.0">10 m</quantity>
from rocky streams in evergreen forest.
</paragraph>
</subSubSection>
<subSubSection id="C3506547BC2AFF85FF63FAD2FE0BFA1C" pageId="2" pageNumber="59" type="etymology">
<paragraph id="8BF536CCBC2AFF85FF63FAD2FE0BFA1C" blockId="2.[151,1437,1007,2004]" pageId="2" pageNumber="59">
<emphasis id="B93EEADEBC2AFF85FF63FAD2FEB9FAC7" bold="true" box="[197,342,1381,1407]" pageId="2" pageNumber="59">Etymology:</emphasis>
Specific name from 
<taxonomicName id="4C4A4D4FBC2AFF85FDE9FAD2FD46FAC6" box="[591,681,1381,1406]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2AFF85FDE9FAD2FD46FAC6" box="[591,681,1381,1406]" italics="true" pageId="2" pageNumber="59">melicus</emphasis>
</taxonomicName>
L., meaning musical or lyrical, in reference to the complex call structure of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C3506547BC2AFF85FF63FA18FEA3F887" pageId="2" pageNumber="59" type="diagnosis">
<paragraph id="8BF536CCBC2AFF85FF63FA18FEA3F887" blockId="2.[151,1437,1007,2004]" pageId="2" pageNumber="59">
<emphasis id="B93EEADEBC2AFF85FF63FA18FEA8FA71" bold="true" box="[197,327,1455,1481]" pageId="2" pageNumber="59">Diagnosis:</emphasis>
Assigned to the genus 
<taxonomicName id="4C4A4D4FBC2AFF85FDF3FA07FD3DFA71" box="[597,722,1456,1481]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="genus">
<emphasis id="B93EEADEBC2AFF85FDF3FA07FD3DFA71" box="[597,722,1456,1481]" italics="true" pageId="2" pageNumber="59">Leptolalax</emphasis>
</taxonomicName>
on the basis of the following: small size, rounded finger tips, the presence of an elevated inner palmar tubercle not continuous to the thumb, presence of macroglands on body (including supra-axillary, pectoral, and femoral glands), vomerine teeth absent, tubercles on eyelids, anterior tip of snout with vertical white bar (
<bibRefCitation id="EFDB4B3DBC2AFF85FD69F997FC98F982" author="Dubois" box="[719,887,1568,1594]" pageId="2" pageNumber="59" refString="Dubois, A. (1983) Note preliminaire sur le genre Leptolalax Dubois, 1980 (Amphibiens, Anoures), avec diagnose d'une espece novella du Vietnam. Alytes, 2, 147 - 153." type="journal article" year="1983">Dubois 1983</bibRefCitation>
; 
<bibRefCitation id="EFDB4B3DBC2AFF85FC23F997FB92F982" author="Lathrop" box="[901,1149,1568,1594]" pageId="2" pageNumber="59" refString="Lathrop, A., Murphy, R. W., Orlov, N. &amp; Ho, C. T. (1998) Two new species of Leptolalax (Anura: Megophryidae) from northern Vietnam. Amphibia-Reptilia, 19, 253 - 267." type="journal article" year="1998">
Lathrop 
<emphasis id="B93EEADEBC2AFF85FC57F997FBC5F981" box="[1009,1066,1568,1593]" italics="true" pageId="2" pageNumber="59">et al</emphasis>
. 1998
</bibRefCitation>
; 
<bibRefCitation id="EFDB4B3DBC2AFF85FB2AF997FA7FF982" author="Delorme" box="[1164,1424,1568,1594]" pageId="2" pageNumber="59" refString="Delorme, M., Dubois, A., Grosjean, S. &amp; Ohler, A. (2006) Une nouvelle ergotaxinomie des Megophryidae (Amphibia, Anura). Alytes, 24, 6 - 21." type="journal article" year="2006">
Delorme 
<emphasis id="B93EEADEBC2AFF85FAA4F997FAD4F981" box="[1282,1339,1568,1593]" italics="true" pageId="2" pageNumber="59">et al</emphasis>
. 2006
</bibRefCitation>
). 
<taxonomicName id="4C4A4D4FBC2AFF85FF31F9F2FE95F9E6" box="[151,378,1605,1630]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="59" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2AFF85FF31F9F2FE95F9E6" box="[151,378,1605,1630]" italics="true" pageId="2" pageNumber="59">Leptolalax melicus</emphasis>
</taxonomicName>
is distinguished from its congeners by a combination of (1) a white to pale pink ventral surface with diffuse dark brown blotches and distinct white speckling, (2) finger I &lt;II, (3) an absence of webbing and dermal fringes on fingers, (4) slight basal webbing and no dermal fringes on toes, (5) body size (
<quantity id="4CB29B29BC2AFF85FF39F902FEA4F977" box="[159,331,1717,1743]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" metricValueMax="2.27" metricValueMin="1.95" pageId="2" pageNumber="59" unit="mm" value="21.1" valueMax="22.7" valueMin="19.5">19.5–22.7 mm</quantity>
for seven adult males), (6) absence of ventrolateral glandular lines, (7) mostly smooth dorsum with no skin ridges, and (8) unique advertisement call consisting of a single long introductory note containing 8–50 pulses, followed by 3–11 predominantly single-pulse notes, and with an average dominant frequency of 3560–3610 Hz.
</paragraph>
</subSubSection>
<subSubSection id="C3506547BC2AFF8EFF63F8FDFB0FFD5A" lastPageId="9" lastPageNumber="66" pageId="2" pageNumber="59" type="description">
<paragraph id="8BF536CCBC2AFF84FF63F8FDFDDEFDF4" blockId="2.[151,1437,1007,2004]" lastBlockId="3.[151,1437,152,588]" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
<emphasis id="B93EEADEBC2AFF85FF63F8FDFE1DF8DC" bold="true" box="[197,498,1866,1892]" pageId="2" pageNumber="59">
Description of 
<typeStatus id="54F1886EBC2AFF85FEDBF8FDFE02F8DC" box="[381,493,1866,1892]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
:
</emphasis>
Head longer than wide; snout rounded in profile, projecting slightly over lower jaw; nostril closer to tip of snout than eye; canthus rostralis rounded; lores sloping, concave; vertical pupil; eye diameter equal to snout length; tympanum distinct, round, diameter smaller than that of the eye; tympanic annulus elevated relative to skin of temporal region; vomerine teeth absent; pineal ocellus absent; vocal sac openings large, oval; tongue large, broad, with small notch at tip; raised supratympanic ridge running from eye towards axillary gland. Tips of fingers rounded, slightly enlarged; relative finger lengths I &lt;II = IV &lt;III; nuptial pad absent; subarticular tubercles absent; a large, round inner palmar tubercle distinctly separated from small, laterally compressed outer palmar tubercle; no finger webbing or lateral fringes. Tips of toes like fingers; relative toe length I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent, replaced by dermal ridges, distinct on second, third, fourth and fifth toes; small, oval inner metatarsal tubercle pronounced, outer metatarsal tubercle absent; webbing basal, confined to very base of toes; no lateral fringes. Tibia relatively short and stout, width approximately one-third of length; tibiotarsal articulation reaches snout. Skin on dorsum mostly smooth, with fine, scattered tubercles concentrated on eyelids and ventrolateral surfaces; ventral skin smooth; pectoral gland circular, 
<quantity id="4CB29B29BC2BFF84FD67FE5FFCF0FDBA" box="[705,799,488,514]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="60" unit="mm" value="0.8">0.8 mm</quantity>
diameter; femoral gland oval, 
<quantity id="4CB29B29BC2BFF84FB3BFE5FFB14FDBA" box="[1181,1275,488,514]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="60" unit="mm" value="0.7">0.7 mm</quantity>
diameter, on posteroventral surface of thigh, closer to knee than to vent; supra-axillary gland raised, 
<quantity id="4CB29B29BC2BFF84FB61FDBAFACBFD9F" box="[1223,1316,525,551]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="60" unit="mm" value="0.7">0.7 mm</quantity>
diameter. Ventrolateral glandular line absent.
</paragraph>
<caption id="DF356644BC2BFF84FF31F8BFFE6BF827" httpUri="https://zenodo.org/record/197319/files/figure.png" pageId="3" pageNumber="60" targetBox="[151,1436,645,1776]" targetPageId="3">
<paragraph id="8BF536CCBC2BFF84FF31F8BFFE6BF827" blockId="3.[151,1436,1800,1951]" pageId="3" pageNumber="60">
<emphasis id="B93EEADEBC2BFF84FF31F8BFFEF2F898" bold="true" box="[151,285,1800,1824]" pageId="3" pageNumber="60">FIGURE 1.</emphasis>
Collection sites of 
<taxonomicName id="4C4A4D4FBC2BFF84FE51F8BEFD27F898" box="[503,712,1801,1824]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="melicus" status="sp. nov.">
<emphasis id="B93EEADEBC2BFF84FE51F8BEFD27F898" box="[503,712,1801,1824]" italics="true" pageId="3" pageNumber="60">Leptolalax melicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57A5BC2BFF84FD76F8BFFCC6F898" box="[720,809,1800,1824]" pageId="3" pageNumber="60" rank="species">
<emphasis id="B93EEADEBC2BFF84FD76F8BFFCC6F898" bold="true" box="[720,809,1800,1824]" pageId="3" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
from Virachey National Park, Cambodia (black circles), 
<taxonomicName id="4C4A4D4FBC2BFF84FF31F89EFE82F8F8" box="[151,365,1833,1856]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B93EEADEBC2BFF84FF31F89EFE82F8F8" box="[151,365,1833,1856]" italics="true" pageId="3" pageNumber="60">Leptolalax applebyi</emphasis>
</taxonomicName>
from Song Thanh Proposed Nature Reserve, Vietnam (dark grey square; Rowley &amp; Cao 2009) and 
<taxonomicName id="4C4A4D4FBC2BFF84FF31F8FEFEE2F8D8" box="[151,269,1865,1888]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="B93EEADEBC2BFF84FF31F8FEFEE2F8D8" box="[151,269,1865,1888]" italics="true" pageId="3" pageNumber="60">Leptolalax</emphasis>
</taxonomicName>
sp. from the Cardamom Mountains, Cambodia (dark grey triangle; Ohler 
<emphasis id="B93EEADEBC2BFF84FBE0F8FEFB98F8D8" box="[1094,1143,1865,1888]" italics="true" pageId="3" pageNumber="60">et al</emphasis>
. 2002). Areas over 600 m elevation are pale grey. Boundaries of Virachey National Park (hatched area) were obtained from the World Database on Protected Areas 2009.
</paragraph>
</caption>
<caption id="DF356644BC2CFF83FF31F8E7FB3AF83F" httpUri="https://zenodo.org/record/197320/files/figure.png" pageId="4" pageNumber="61" targetBox="[151,1436,178,1850]" targetPageId="4">
<paragraph id="8BF536CCBC2CFF83FF31F8E7FB3AF83F" blockId="4.[151,1436,1872,1927]" pageId="4" pageNumber="61">
<emphasis id="B93EEADEBC2CFF83FF31F8E7FEF6F8D0" bold="true" box="[151,281,1872,1896]" pageId="4" pageNumber="61">FIGURE 2.</emphasis>
Male holotype of 
<taxonomicName id="4C4A4D4FBC2CFF83FE7BF8E6FD48F8D0" box="[477,679,1873,1896]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="61" phylum="Chordata" rank="species" species="melicus" status="sp. nov.">
<emphasis id="B93EEADEBC2CFF83FE7BF8E6FD48F8D0" box="[477,679,1873,1896]" italics="true" pageId="4" pageNumber="61">Leptolalax melicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57A5BC2CFF83FD08F8E7FCECF8D0" box="[686,771,1872,1896]" pageId="4" pageNumber="61" rank="species">
<emphasis id="B93EEADEBC2CFF83FD08F8E7FCECF8D0" bold="true" box="[686,771,1872,1896]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
(MVZ 258198) in life. (A) Lateral view in the field, in calling posture (nocturnal ventral colouration), (B) dorsal and (C) ventral views (diurnal ventral colouration).
</paragraph>
</caption>
<paragraph id="8BF536CCBC2CFF82FF63F801FD67FDF4" blockId="4.[151,1436,1974,2037]" lastBlockId="5.[151,1437,152,1074]" lastPageId="5" lastPageNumber="62" pageId="4" pageNumber="61">
<emphasis id="B93EEADEBC2CFF83FF63F801FDE1F868" bold="true" box="[197,526,1974,2000]" pageId="4" pageNumber="61">
Colour of 
<typeStatus id="54F1886EBC2CFF83FEE1F801FE5BF868" box="[327,436,1974,2000]" pageId="4" pageNumber="61" type="holotype">holotype</typeStatus>
in life:
</emphasis>
Dorsal surface brown; distinct dark brown markings lined with diffuse pale copper colour on dorsum, V-shaped interorbital marking, W-shaped marking between axillae and inverted Vshaped marking above sacrum; dorsal surface of head anterior to eyes pale copper; fine tubercles on dorsum reddish; vertical blackish brown bars on background of pale copper on upper lip; black line along canthus rostralis, through eye, and continuing along supratympanic ridge, encompassing most of tympanum, terminating above axilla; pale copper coloured area under supratympanic ridge, fine reddish line above supratympanic ridge; transverse blackish brown bars on dorsal surface of limbs; large, black blotch on flank and numerous, smaller black spots on sides from groin to axilla; ventral surface of elbow and upper arms without dark bars but with distinct copper colouration; fingers and toes with transverse barring; pale pink ventral surface with diffuse dark blotches and distinct white speckling concentrated on belly, but also scattered on throat, and lateral surfaces of body and upper arms, nearly absent from ventral surface of legs; ventral margin of throat bearing broken row of slightly larger white spots. At night, ventral surface much paler, off-white. Supra-axillary gland copper; femoral and pectoral glands white, lined with dark brown. Iris dark gold with minute, black reticulations.
</paragraph>
<paragraph id="8BF536CCBC2DFF82FF63FDE0FE9DFD04" blockId="5.[151,1437,152,1074]" pageId="5" pageNumber="62">
<emphasis id="B93EEADEBC2DFF82FF63FDE0FD6FFDC9" bold="true" box="[197,640,599,625]" pageId="5" pageNumber="62">
Colour of 
<typeStatus id="54F1886EBC2DFF82FEE0FDE0FE5DFDC9" box="[326,434,599,625]" pageId="5" pageNumber="62" type="holotype">holotype</typeStatus>
in preservative:
</emphasis>
Dorsum dark brown with slightly paler limbs. Ventral surface pale greyish brown. Macroglands white. Patterns on dorsal surface and white speckling on ventral surface less distinct (
<figureCitation id="13712A49BC2DFF82FF5AFD15FE8DFD04" box="[252,354,674,700]" captionStart="FIGURE 3" captionStartId="6.[151,257,1623,1647]" captionTargetBox="[151,1422,207,1601]" captionTargetId="figure@6.[151,1436,197,1602]" captionTargetPageId="6" captionText="FIGURE 3. (A) Dorsal view, (B) ventral view and (C) lateral view of head of preserved holotype (MVZ 258198) of Leptolalax melicus sp. nov. Scale bars = 2 mm." httpUri="https://zenodo.org/record/197321/files/figure.png" pageId="5" pageNumber="62">Figure 3</figureCitation>
).
</paragraph>
<paragraph id="8BF536CCBC2DFF82FF63FD70FC2BFCBF" blockId="5.[151,1437,152,1074]" pageId="5" pageNumber="62">
<emphasis id="B93EEADEBC2DFF82FF63FD70FE68FD59" bold="true" box="[197,391,711,737]" pageId="5" pageNumber="62">Measurements:</emphasis>
<typeStatus id="54F1886EBC2DFF82FE36FD7FFDEDFD5A" box="[400,514,712,738]" pageId="5" pageNumber="62" type="holotype">Holotype</typeStatus>
: SVL 20.1, HDL 7.7, HDW 7.4, SNT 3.0, EYE 2.7, IOD 2.4, TMP 1.2, TEY 0.6, TIB 9.6, ML 4.8, PL 9.1, F1L 1.5, F2L 1.8, F3L 3.5, weight 
<quantity id="4CB29B29BC2DFF82FC36FD5AFC2BFCBF" box="[912,964,749,775]" metricMagnitude="-4" metricUnit="kg" metricValue="7.0" pageId="5" pageNumber="62" unit="g" value="0.7">0.7g</quantity>
</paragraph>
<paragraph id="8BF536CCBC2DFF82FF63FCA5FEE3FB8A" blockId="5.[151,1437,152,1074]" pageId="5" pageNumber="62">
<emphasis id="B93EEADEBC2DFF82FF63FCA5FED2FC94" bold="true" box="[197,317,786,812]" pageId="5" pageNumber="62">Variation</emphasis>
. Size and extent of white speckling on ventral and ventrolateral surfaces may be such that anterior half of belly is mostly white with pinkish brown mottling (MVZ 258075, 258197), rather than mostly off-white or pale pink. Specimens vary in the number and size of black blotches ventrolaterally. MVZ 258197 has large white blotches ventrolaterally, in addition to black blotches. In some specimens, dorsal markings join along midline to form a continuous dark brown marking (MVZ 258077, 258199). A single specimen (MVZ 258199) has small non-pigmented patches of skin on knee and groin. Relative finger lengths may be I &lt;II ≤ IV &lt;III. Measurements of the 
<typeStatus id="54F1886EBC2DFF82FDCBFC45FD4CFBB4" box="[621,675,1010,1036]" pageId="5" pageNumber="62">type</typeStatus>
series are shown in 
<tableCitation id="C6C80377BC2DFF82FC00FC45FBEAFBB4" box="[934,1029,1010,1036]" captionStart="TABLE 1" captionStartId="5.[151,239,1122,1146]" captionTargetBox="[151,1434,1248,1825]" captionTargetPageId="5" captionText="TABLE 1. Measurements (mm) of adult male Leptolalax melicus, sp. nov. Abbreviations defined in text." httpUri="http://table.plazi.org/id/DF356644BC2DFF82FF31FBD5FB15FBC1" pageId="5" pageNumber="62" tableUuid="DF356644BC2DFF82FF31FBD5FB15FBC1">Table 1</tableCitation>
. Females of the new species are unknown.
</paragraph>
<caption id="DF356644BC2DFF82FF31FBD5FB15FBC1" ID-Table-UUID="DF356644BC2DFF82FF31FBD5FB15FBC1" box="[151,1274,1122,1146]" httpUri="http://table.plazi.org/id/DF356644BC2DFF82FF31FBD5FB15FBC1" pageId="5" pageNumber="62" targetBox="[151,1434,1248,1825]" targetIsTable="true" targetPageId="5">
<paragraph id="8BF536CCBC2DFF82FF31FBD5FB15FBC1" blockId="5.[151,1434,1122,1228]" box="[151,1274,1122,1146]" pageId="5" pageNumber="62">
<emphasis id="B93EEADEBC2DFF82FF31FBD5FEEDFBC2" bold="true" box="[151,258,1122,1146]" pageId="5" pageNumber="62">TABLE 1</emphasis>
. Measurements (mm) of adult male 
<taxonomicName id="4C4A4D4FBC2DFF82FD2EFBD5FCBDFBC1" box="[648,850,1122,1145]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="62" phylum="Chordata" rank="species" species="melicus" status="sp. nov.">
<emphasis id="B93EEADEBC2DFF82FD2EFBD5FCBDFBC1" box="[648,850,1122,1145]" italics="true" pageId="5" pageNumber="62">Leptolalax melicus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A20D57A5BC2DFF82FCF9FBD5FC5BFBC2" box="[863,948,1122,1146]" pageId="5" pageNumber="62" rank="species">
<emphasis id="B93EEADEBC2DFF82FCF9FBD5FC5BFBC2" bold="true" box="[863,948,1122,1146]" pageId="5" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
Abbreviations defined in text.
</paragraph>
</caption>
<paragraph id="8BF536CCBC2DFF82FEF2FB22FA75FB14" blockId="5.[151,1434,1122,1228]" box="[340,1434,1173,1196]" pageId="5" pageNumber="62">M V Z MVZ 258075 MVZ 258076 MVZ 258077 MVZ 258197 M V Z MVZ 258199</paragraph>
<paragraph id="8BF536CCBC2DFF82FEF2FB02FB58FB74" blockId="5.[151,1434,1122,1228]" box="[340,1207,1205,1228]" pageId="5" pageNumber="62">258074 258198*</paragraph>
<paragraph id="8BF536CCBC2DFF82FF39FB57FAC6F899" pageId="5" pageNumber="62">
<table id="F94AC46CBC2D0078FF31FB57FA75F899" box="[151,1434,1248,1825]" gridcols="7" gridrows="9" pageId="5" pageNumber="62">
<tr id="357A348EBC2D0078FF31FB57FA75FA99" box="[151,1434,1248,1313]" gridrow="0" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31FB57FF31FA99" box="[151,222,1248,1313]" gridcol="0" gridrow="0" pageId="5" pageNumber="62">SVL HDL</th>
<th id="76AB5DF2BC2D0078FEF2FB57FE10FA99" box="[340,511,1248,1313]" gridcol="1" gridrow="0" pageId="5" pageNumber="62">20.7 20.2 8.2 7.9</th>
<th id="76AB5DF2BC2D0078FDD5FB57FD4DFA99" box="[627,674,1248,1313]" gridcol="2" gridrow="0" pageId="5" pageNumber="62">22.8 8.6</th>
<th id="76AB5DF2BC2D0078FCB3FB57FCABFA99" box="[789,836,1248,1313]" gridcol="3" gridrow="0" pageId="5" pageNumber="62">21.8 8.2</th>
<th id="76AB5DF2BC2D0078FC11FB57FC0AFA99" box="[951,997,1248,1313]" gridcol="4" gridrow="0" pageId="5" pageNumber="62">19.8 7.6</th>
<th id="76AB5DF2BC2D0078FBFCFB57FB66FA99" box="[1114,1161,1248,1313]" gridcol="5" gridrow="0" pageId="5" pageNumber="62">20.1 7.7</th>
<th id="76AB5DF2BC2D0078FAAEFB57FA75FA99" box="[1288,1434,1248,1313]" gridcol="6" gridrow="0" pageId="5" pageNumber="62">19.5 7.7</th>
</tr>
<tr id="357A348EBC2D0078FF31FA82FA75FAF4" box="[151,1434,1333,1356]" gridrow="1" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31FA82FF31FAF4" box="[151,222,1333,1356]" gridcol="0" gridrow="1" pageId="5" pageNumber="62">HDW</th>
<td id="76AB5DF2BC2D0078FEF2FA82FE10FAF4" box="[340,511,1333,1356]" gridcol="1" gridrow="1" pageId="5" pageNumber="62">7.5 7.4</td>
<td id="76AB5DF2BC2D0078FDD5FA82FD4DFAF4" box="[627,674,1333,1356]" gridcol="2" gridrow="1" pageId="5" pageNumber="62">8.2</td>
<td id="76AB5DF2BC2D0078FCB3FA82FCABFAF4" box="[789,836,1333,1356]" gridcol="3" gridrow="1" pageId="5" pageNumber="62">7.6</td>
<td id="76AB5DF2BC2D0078FC11FA82FC0AFAF4" box="[951,997,1333,1356]" gridcol="4" gridrow="1" pageId="5" pageNumber="62">7.3</td>
<td id="76AB5DF2BC2D0078FBFCFA82FB66FAF4" box="[1114,1161,1333,1356]" gridcol="5" gridrow="1" pageId="5" pageNumber="62">7.4</td>
<td id="76AB5DF2BC2D0078FAAEFA82FA75FAF4" box="[1288,1434,1333,1356]" gridcol="6" gridrow="1" pageId="5" pageNumber="62">7.0</td>
</tr>
<tr id="357A348EBC2D0078FF31FAD7FA75FA19" box="[151,1434,1376,1441]" gridrow="2" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31FAD7FF31FA19" box="[151,222,1376,1441]" gridcol="0" gridrow="2" pageId="5" pageNumber="62">SNT EYE</th>
<td id="76AB5DF2BC2D0078FEF2FAD7FE10FA19" box="[340,511,1376,1441]" gridcol="1" gridrow="2" pageId="5" pageNumber="62">3.3 3.2 2.8 2.6</td>
<td id="76AB5DF2BC2D0078FDD5FAD7FD4DFA19" box="[627,674,1376,1441]" gridcol="2" gridrow="2" pageId="5" pageNumber="62">3.2 3.1</td>
<td id="76AB5DF2BC2D0078FCB3FAD7FCABFA19" box="[789,836,1376,1441]" gridcol="3" gridrow="2" pageId="5" pageNumber="62">3.2 2.8</td>
<td id="76AB5DF2BC2D0078FC11FAD7FC0AFA19" box="[951,997,1376,1441]" gridcol="4" gridrow="2" pageId="5" pageNumber="62">3.1 2.3</td>
<td id="76AB5DF2BC2D0078FBFCFAD7FB66FA19" box="[1114,1161,1376,1441]" gridcol="5" gridrow="2" pageId="5" pageNumber="62">3.0 2.7</td>
<td id="76AB5DF2BC2D0078FAAEFAD7FA75FA19" box="[1288,1434,1376,1441]" gridcol="6" gridrow="2" pageId="5" pageNumber="62">3.1 2.6</td>
</tr>
<tr id="357A348EBC2D0078FF31FA02FA75FA74" box="[151,1434,1461,1484]" gridrow="3" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31FA02FF31FA74" box="[151,222,1461,1484]" gridcol="0" gridrow="3" pageId="5" pageNumber="62">IOD</th>
<td id="76AB5DF2BC2D0078FEF2FA02FE10FA74" box="[340,511,1461,1484]" gridcol="1" gridrow="3" pageId="5" pageNumber="62">2.9 2.5</td>
<td id="76AB5DF2BC2D0078FDD5FA02FD4DFA74" box="[627,674,1461,1484]" gridcol="2" gridrow="3" pageId="5" pageNumber="62">2.9</td>
<td id="76AB5DF2BC2D0078FCB3FA02FCABFA74" box="[789,836,1461,1484]" gridcol="3" gridrow="3" pageId="5" pageNumber="62">2.8</td>
<td id="76AB5DF2BC2D0078FC11FA02FC0AFA74" box="[951,997,1461,1484]" gridcol="4" gridrow="3" pageId="5" pageNumber="62">2.5</td>
<td id="76AB5DF2BC2D0078FBFCFA02FB66FA74" box="[1114,1161,1461,1484]" gridcol="5" gridrow="3" pageId="5" pageNumber="62">2.4</td>
<td id="76AB5DF2BC2D0078FAAEFA02FA75FA74" box="[1288,1434,1461,1484]" gridcol="6" gridrow="3" pageId="5" pageNumber="62">2.4</td>
</tr>
<tr id="357A348EBC2D0078FF31FA57FA75F999" box="[151,1434,1504,1569]" gridrow="4" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31FA57FF31F999" box="[151,222,1504,1569]" gridcol="0" gridrow="4" pageId="5" pageNumber="62">TMP TEY</th>
<td id="76AB5DF2BC2D0078FEF2FA57FE10F999" box="[340,511,1504,1569]" gridcol="1" gridrow="4" pageId="5" pageNumber="62">1.5 1.1 0.7 0.7</td>
<td id="76AB5DF2BC2D0078FDD5FA57FD4DF999" box="[627,674,1504,1569]" gridcol="2" gridrow="4" pageId="5" pageNumber="62">1.3 1.0</td>
<td id="76AB5DF2BC2D0078FCB3FA57FCABF999" box="[789,836,1504,1569]" gridcol="3" gridrow="4" pageId="5" pageNumber="62">1.2 0.7</td>
<td id="76AB5DF2BC2D0078FC11FA57FC0AF999" box="[951,997,1504,1569]" gridcol="4" gridrow="4" pageId="5" pageNumber="62">1.3 0.7</td>
<td id="76AB5DF2BC2D0078FBFCFA57FB66F999" box="[1114,1161,1504,1569]" gridcol="5" gridrow="4" pageId="5" pageNumber="62">1.2 0.6</td>
<td id="76AB5DF2BC2D0078FAAEFA57FA75F999" box="[1288,1434,1504,1569]" gridcol="6" gridrow="4" pageId="5" pageNumber="62">1.0 0.7</td>
</tr>
<tr id="357A348EBC2D0078FF31F982FA75F9F4" box="[151,1434,1589,1612]" gridrow="5" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31F982FF31F9F4" box="[151,222,1589,1612]" gridcol="0" gridrow="5" pageId="5" pageNumber="62">TIB</th>
<td id="76AB5DF2BC2D0078FEF2F982FE10F9F4" box="[340,511,1589,1612]" gridcol="1" gridrow="5" pageId="5" pageNumber="62">9.9 9.2</td>
<td id="76AB5DF2BC2D0078FDD5F982FD4DF9F4" box="[627,674,1589,1612]" gridcol="2" gridrow="5" pageId="5" pageNumber="62">10.4</td>
<td id="76AB5DF2BC2D0078FCB3F982FCABF9F4" box="[789,836,1589,1612]" gridcol="3" gridrow="5" pageId="5" pageNumber="62">10.1</td>
<td id="76AB5DF2BC2D0078FC11F982FC0AF9F4" box="[951,997,1589,1612]" gridcol="4" gridrow="5" pageId="5" pageNumber="62">9.1</td>
<td id="76AB5DF2BC2D0078FBFCF982FB66F9F4" box="[1114,1161,1589,1612]" gridcol="5" gridrow="5" pageId="5" pageNumber="62">9.6</td>
<td id="76AB5DF2BC2D0078FAAEF982FA75F9F4" box="[1288,1434,1589,1612]" gridcol="6" gridrow="5" pageId="5" pageNumber="62">9.4</td>
</tr>
<tr id="357A348EBC2D0078FF31F9D7FA75F919" box="[151,1434,1632,1697]" gridrow="6" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31F9D7FF31F919" box="[151,222,1632,1697]" gridcol="0" gridrow="6" pageId="5" pageNumber="62">ML PL</th>
<td id="76AB5DF2BC2D0078FEF2F9D7FE10F919" box="[340,511,1632,1697]" gridcol="1" gridrow="6" pageId="5" pageNumber="62">4.8 4.6 9.1 8.9</td>
<td id="76AB5DF2BC2D0078FDD5F9D7FD4DF919" box="[627,674,1632,1697]" gridcol="2" gridrow="6" pageId="5" pageNumber="62">4.8 9.2</td>
<td id="76AB5DF2BC2D0078FCB3F9D7FCABF919" box="[789,836,1632,1697]" gridcol="3" gridrow="6" pageId="5" pageNumber="62">5.0 9.1</td>
<td id="76AB5DF2BC2D0078FC11F9D7FC0AF919" box="[951,997,1632,1697]" gridcol="4" gridrow="6" pageId="5" pageNumber="62">4.5 8.9</td>
<td id="76AB5DF2BC2D0078FBFCF9D7FB66F919" box="[1114,1161,1632,1697]" gridcol="5" gridrow="6" pageId="5" pageNumber="62">4.8 9.1</td>
<td id="76AB5DF2BC2D0078FAAEF9D7FA75F919" box="[1288,1434,1632,1697]" gridcol="6" gridrow="6" pageId="5" pageNumber="62">4.6 8.7</td>
</tr>
<tr id="357A348EBC2D0078FF31F902FA75F974" box="[151,1434,1717,1740]" gridrow="7" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31F902FF31F974" box="[151,222,1717,1740]" gridcol="0" gridrow="7" pageId="5" pageNumber="62">F1L</th>
<td id="76AB5DF2BC2D0078FEF2F902FE10F974" box="[340,511,1717,1740]" gridcol="1" gridrow="7" pageId="5" pageNumber="62">1.6 1.7</td>
<td id="76AB5DF2BC2D0078FDD5F902FD4DF974" box="[627,674,1717,1740]" gridcol="2" gridrow="7" pageId="5" pageNumber="62">1.7</td>
<td id="76AB5DF2BC2D0078FCB3F902FCABF974" box="[789,836,1717,1740]" gridcol="3" gridrow="7" pageId="5" pageNumber="62">1.8</td>
<td id="76AB5DF2BC2D0078FC11F902FC0AF974" box="[951,997,1717,1740]" gridcol="4" gridrow="7" pageId="5" pageNumber="62">1.7</td>
<td id="76AB5DF2BC2D0078FBFCF902FB66F974" box="[1114,1161,1717,1740]" gridcol="5" gridrow="7" pageId="5" pageNumber="62">1.5</td>
<td id="76AB5DF2BC2D0078FAAEF902FA75F974" box="[1288,1434,1717,1740]" gridcol="6" gridrow="7" pageId="5" pageNumber="62">1.6</td>
</tr>
<tr id="357A348EBC2D0078FF31F957FA75F899" box="[151,1434,1760,1825]" gridrow="8" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31F957FF31F899" box="[151,222,1760,1825]" gridcol="0" gridrow="8" pageId="5" pageNumber="62">F2L F3L</th>
<td id="76AB5DF2BC2D0078FEF2F957FE10F899" box="[340,511,1760,1825]" gridcol="1" gridrow="8" pageId="5" pageNumber="62">1.8 1.8 4.0 3.8</td>
<td id="76AB5DF2BC2D0078FDD5F957FD4DF899" box="[627,674,1760,1825]" gridcol="2" gridrow="8" pageId="5" pageNumber="62">2.1 3.9</td>
<td id="76AB5DF2BC2D0078FCB3F957FCABF899" box="[789,836,1760,1825]" gridcol="3" gridrow="8" pageId="5" pageNumber="62">2.0 4.1</td>
<td id="76AB5DF2BC2D0078FC11F957FC0AF899" box="[951,997,1760,1825]" gridcol="4" gridrow="8" pageId="5" pageNumber="62">2.0 3.5</td>
<td id="76AB5DF2BC2D0078FBFCF957FB66F899" box="[1114,1161,1760,1825]" gridcol="5" gridrow="8" pageId="5" pageNumber="62">1.8 3.5</td>
<td id="76AB5DF2BC2D0078FAAEF957FA75F899" box="[1288,1434,1760,1825]" gridcol="6" gridrow="8" pageId="5" pageNumber="62">1.8 3.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF536CCBC2DFF82FF39F882FEEEF861" pageId="5" pageNumber="62">
<table id="F94AC46CBC2D0078FF31F882FA75F861" box="[151,1434,1845,2009]" gridcols="6" gridrows="3" pageId="5" pageNumber="62">
<tr id="357A348EBC2D0078FF31F882FA75F8F4" box="[151,1434,1845,1868]" gridrow="0" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31F882FE10F8F4" box="[151,511,1845,1868]" gridcol="0" gridrow="0" pageId="5" pageNumber="62">TIB:SVL 0.48 0.46</th>
<th id="76AB5DF2BC2D0078FDD5F882FD4DF8F4" box="[627,674,1845,1868]" gridcol="1" gridrow="0" pageId="5" pageNumber="62">0.46</th>
<th id="76AB5DF2BC2D0078FCB3F882FCABF8F4" box="[789,836,1845,1868]" gridcol="2" gridrow="0" pageId="5" pageNumber="62">0.46</th>
<th id="76AB5DF2BC2D0078FC11F882FC0AF8F4" box="[951,997,1845,1868]" gridcol="3" gridrow="0" pageId="5" pageNumber="62">0.46</th>
<th id="76AB5DF2BC2D0078FBFCF882FB66F8F4" box="[1114,1161,1845,1868]" gridcol="4" gridrow="0" pageId="5" pageNumber="62">0.48</th>
<th id="76AB5DF2BC2D0078FAAEF882FA75F8F4" box="[1288,1434,1845,1868]" gridcol="5" gridrow="0" pageId="5" pageNumber="62">0.48</th>
</tr>
<tr id="357A348EBC2D0078FF31F8D7FA75F819" box="[151,1434,1888,1953]" gridrow="1" pageId="5" pageNumber="62">
<th id="76AB5DF2BC2D0078FF31F8D7FE10F819" box="[151,511,1888,1953]" gridcol="0" gridrow="1" pageId="5" pageNumber="62">HDL:SVL 0.40 0.39 HDL:HDW 1.09 1.07</th>
<td id="76AB5DF2BC2D0078FDD5F8D7FD4DF819" box="[627,674,1888,1953]" gridcol="1" gridrow="1" pageId="5" pageNumber="62">0.38 1.05</td>
<td id="76AB5DF2BC2D0078FCB3F8D7FCABF819" box="[789,836,1888,1953]" gridcol="2" gridrow="1" pageId="5" pageNumber="62">0.38 1.08</td>
<td id="76AB5DF2BC2D0078FC11F8D7FC0AF819" box="[951,997,1888,1953]" gridcol="3" gridrow="1" pageId="5" pageNumber="62">0.38 1.04</td>
<td id="76AB5DF2BC2D0078FBFCF8D7FB66F819" box="[1114,1161,1888,1953]" gridcol="4" gridrow="1" pageId="5" pageNumber="62">0.38 1.04</td>
<td id="76AB5DF2BC2D0078FAAEF8D7FA75F819" box="[1288,1434,1888,1953]" gridcol="5" gridrow="1" pageId="5" pageNumber="62">0.39 1.10</td>
</tr>
<tr id="357A348EBC2D0078FF31F875FA75F861" box="[151,1434,1986,2009]" gridrow="2" pageId="5" pageNumber="62" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76AB5DF2BC2D0078FF31F875FE10F861" box="[151,511,1986,2009]" gridcol="0" gridrow="2" pageId="5" pageNumber="62">*holotype</th>
</tr>
</table>
</paragraph>
<caption id="DF356644BC2EFF81FF31F9E0FD7DF936" httpUri="https://zenodo.org/record/197321/files/figure.png" pageId="6" pageNumber="63" targetBox="[151,1422,207,1601]" targetPageId="6">
<paragraph id="8BF536CCBC2EFF81FF31F9E0FD7DF936" blockId="6.[151,1436,1623,1679]" pageId="6" pageNumber="63">
<emphasis id="B93EEADEBC2EFF81FF31F9E0FEF2F9D7" bold="true" box="[151,285,1623,1647]" pageId="6" pageNumber="63">FIGURE 3.</emphasis>
(A) Dorsal view, (B) ventral view and (C) lateral view of head of preserved holotype (MVZ 258198) of 
<taxonomicName id="4C4A4D4FBC2EFF81FF31F9C0FE8EF936" box="[151,353,1655,1678]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="63" phylum="Chordata" rank="species" species="melicus" status="sp. nov.">
<emphasis id="B93EEADEBC2EFF81FF31F9C0FE8EF936" box="[151,353,1655,1678]" italics="true" pageId="6" pageNumber="63">Leptolalax melicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57A5BC2EFF81FEC1F9C0FE52F937" box="[359,445,1655,1679]" pageId="6" pageNumber="63" rank="species">
<emphasis id="B93EEADEBC2EFF81FEC1F9C0FE52F937" bold="true" box="[359,445,1655,1679]" pageId="6" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="8BF536CCBC2EFF80FF63F90BFEC9FE9A" blockId="6.[151,1437,1724,2012]" lastBlockId="7.[151,1437,152,290]" lastPageId="7" lastPageNumber="64" pageId="6" pageNumber="63">
<emphasis id="B93EEADEBC2EFF81FF63F90BFE54F96E" bold="true" box="[197,443,1724,1750]" pageId="6" pageNumber="63">Advertisement call:</emphasis>
Advertisement call descriptions are based on the calls of the 
<typeStatus id="54F1886EBC2EFF81FB2BF90BFB1BF96E" box="[1165,1268,1724,1750]" pageId="6" pageNumber="63" type="holotype">holotype</typeStatus>
MVZ 258198 and 
<typeStatus id="54F1886EBC2EFF81FF6BF955FED6F944" box="[205,313,1762,1788]" pageId="6" pageNumber="63" type="paratype">paratype</typeStatus>
MVZ 258199, taken at ambient temperatures of 26.1ºC and 26.2ºC respectively. In both recorded calls, call intensity and call rate were highly variable (
<figureCitation id="13712A49BC2EFF81FC25F8B0FC06F899" box="[899,1001,1799,1825]" captionStart="FIGURE 4" captionStartId="8.[151,259,1291,1315]" captionTargetBox="[155,1431,384,1264]" captionTargetId="figure@8.[151,1436,381,1270]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement call of Leptolalax melicus sp. nov. (holotype MVZ 258198) recorded at ambient air temperature of 26.1 º C. (A) 60 s waveform of relative amplitude over time, (B) waveform and (C) corresponding spectrogram of single representative call expanded from section shown in A, and (D) power spectrum (relative amplitude vs. frequency) of introductory note and (E) click." httpUri="https://zenodo.org/record/197322/files/figure.png" pageId="6" pageNumber="63">Figure 4</figureCitation>
A). Although both the amplitude and rate of calling increased and decreased during recordings, calls did not form distinct call groups (in up to 120 s of calling). Individual calls were consistent in overall structure, consisting of a single introductory note of a variable length (24–132 ms), containing 8–50 pulses, followed by 4–11 short (2–24 ms) notes, or clicks, of 1– 2 pulses repeated at a rate of approximately 15–26 notes per second (
<tableCitation id="C6C80377BC2EFF81FC79F82BFBD2F80E" box="[991,1085,1948,1974]" captionStart="TABLE 2" captionStartId="7.[151,239,332,356]" captionTargetBox="[151,1436,416,1136]" captionTargetPageId="7" captionText="TABLE 2. Measurements of advertisement call parameters for Leptolalax melicus sp. nov. Parameter values are given as" httpUri="http://table.plazi.org/id/DF356644BC2FFF80FF31FEFBFA73FEDC" pageId="6" pageNumber="63" tableUuid="DF356644BC2FFF80FF31FEFBFA73FEDC">Table 2</tableCitation>
, 
<figureCitation id="13712A49BC2EFF81FBEDF82BFB5DF80E" box="[1099,1202,1948,1974]" captionStart="FIGURE 4" captionStartId="8.[151,259,1291,1315]" captionTargetBox="[155,1431,384,1264]" captionTargetId="figure@8.[151,1436,381,1270]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement call of Leptolalax melicus sp. nov. (holotype MVZ 258198) recorded at ambient air temperature of 26.1 º C. (A) 60 s waveform of relative amplitude over time, (B) waveform and (C) corresponding spectrogram of single representative call expanded from section shown in A, and (D) power spectrum (relative amplitude vs. frequency) of introductory note and (E) click." httpUri="https://zenodo.org/record/197322/files/figure.png" pageId="6" pageNumber="63">Figure 4</figureCitation>
B–C). Pulses in the introductory notes were of less than half the amplitude of clicks, and peak amplitude was usually towards the middle of the click series. Both note 
<typeStatus id="54F1886EBC2FFF80FDF6FF2FFD60FF0A" box="[592,655,152,178]" pageId="7" pageNumber="64">types</typeStatus>
had similar dominant frequencies of 2584–3962.1 Hz (
<figureCitation id="13712A49BC2FFF80FA82FF2FFA62FF0A" box="[1316,1421,152,178]" captionStart="FIGURE 4" captionStartId="8.[151,259,1291,1315]" captionTargetBox="[155,1431,384,1264]" captionTargetId="figure@8.[151,1436,381,1270]" captionTargetPageId="8" captionText="FIGURE 4. Advertisement call of Leptolalax melicus sp. nov. (holotype MVZ 258198) recorded at ambient air temperature of 26.1 º C. (A) 60 s waveform of relative amplitude over time, (B) waveform and (C) corresponding spectrogram of single representative call expanded from section shown in A, and (D) power spectrum (relative amplitude vs. frequency) of introductory note and (E) click." httpUri="https://zenodo.org/record/197322/files/figure.png" pageId="7" pageNumber="64">Figure 4</figureCitation>
), with faint harmonics at approximately 7752 Hz. To the human ear, the call of 
<taxonomicName id="4C4A4D4FBC2FFF80FBECFF0AFB2AFF6E" box="[1098,1221,189,214]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2FFF80FBECFF0AFB2AFF6E" box="[1098,1221,189,214]" italics="true" pageId="7" pageNumber="64">L. melicus</emphasis>
</taxonomicName>
contains a barely discernable ‘squeak’ (introductory note), followed by a faint clicking, and resembles the call of an orthopteran.
</paragraph>
<caption id="DF356644BC2FFF80FF31FEFBFA73FEDC" ID-Table-UUID="DF356644BC2FFF80FF31FEFBFA73FEDC" box="[151,1436,332,356]" httpUri="http://table.plazi.org/id/DF356644BC2FFF80FF31FEFBFA73FEDC" pageId="7" pageNumber="64" targetBox="[151,1436,416,1136]" targetIsTable="true" targetPageId="7">
<paragraph id="8BF536CCBC2FFF80FF31FEFBFA73FEDC" blockId="7.[151,1436,332,388]" box="[151,1436,332,356]" pageId="7" pageNumber="64">
<emphasis id="B93EEADEBC2FFF80FF31FEFBFEE7FEDC" bold="true" box="[151,264,332,356]" pageId="7" pageNumber="64">TABLE 2.</emphasis>
Measurements of advertisement call parameters for 
<taxonomicName id="4C4A4D4FBC2FFF80FC90FEFAFC10FEDC" box="[822,1023,333,356]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="melicus" status="sp. nov.">
<emphasis id="B93EEADEBC2FFF80FC90FEFAFC10FEDC" box="[822,1023,333,356]" italics="true" pageId="7" pageNumber="64">Leptolalax melicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57A5BC2FFF80FBA0FEFBFBB4FEDC" box="[1030,1115,332,356]" pageId="7" pageNumber="64" rank="species">
<emphasis id="B93EEADEBC2FFF80FBA0FEFBFBB4FEDC" bold="true" box="[1030,1115,332,356]" pageId="7" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
Parameter values are given as
</paragraph>
</caption>
<paragraph id="8BF536CCBC2FFF80FF31FEDAFE81FE3C" blockId="7.[151,1436,332,388]" box="[151,366,365,388]" pageId="7" pageNumber="64">means (and ranges).</paragraph>
<paragraph id="8BF536CCBC2FFF80FF39FE17FEEEFBC8" pageId="7" pageNumber="64">
<table id="F94AC46CBC2F0078FF31FE17FA73FBC8" box="[151,1436,416,1136]" gridcols="3" gridrows="11" pageId="7" pageNumber="64">
<tr id="357A348EBC2F0078FF31FE17FA73FE59" box="[151,1436,416,481]" gridrow="0" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FE17FD7FFE59" box="[151,656,416,481]" gridcol="0" gridrow="0" pageId="7" pageNumber="64">Recording Temperature (ºC)</th>
<th id="76AB5DF2BC2F0078FD57FE17FC1DFE59" box="[753,1010,416,481]" gridcol="1" gridrow="0" pageId="7" pageNumber="64">MVZ 258198* 26.1</th>
<th id="76AB5DF2BC2F0078FBE0FE17FA73FE59" box="[1094,1436,416,481]" gridcol="2" gridrow="0" pageId="7" pageNumber="64">MVZ 258199 26.2</th>
</tr>
<tr id="357A348EBC2F0078FF31FE42FA73FDB4" box="[151,1436,501,524]" gridrow="1" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FE42FD7FFDB4" box="[151,656,501,524]" gridcol="0" gridrow="1" pageId="7" pageNumber="64">Number of notes</th>
<td id="76AB5DF2BC2F0078FD57FE42FC1DFDB4" box="[753,1010,501,524]" gridcol="1" gridrow="1" pageId="7" pageNumber="64">268</td>
<td id="76AB5DF2BC2F0078FBE0FE42FA73FDB4" box="[1094,1436,501,524]" gridcol="2" gridrow="1" pageId="7" pageNumber="64">251</td>
</tr>
<tr id="357A348EBC2F0078FF31FD97FA73FDD9" box="[151,1436,544,609]" gridrow="2" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FD97FD7FFDD9" box="[151,656,544,609]" gridcol="0" gridrow="2" pageId="7" pageNumber="64">Call duration (ms) Intercall interval (ms)</th>
<td id="76AB5DF2BC2F0078FD57FD97FC1DFDD9" box="[753,1010,544,609]" gridcol="1" gridrow="2" pageId="7" pageNumber="64">292 (176–484) 920 (214–3821)</td>
<td id="76AB5DF2BC2F0078FBE0FD97FA73FDD9" box="[1094,1436,544,609]" gridcol="2" gridrow="2" pageId="7" pageNumber="64">278 (168–427) 1248 (226–3647)</td>
</tr>
<tr id="357A348EBC2F0078FF31FDC2FA73FD34" box="[151,1436,629,652]" gridrow="3" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FDC2FD7FFD34" box="[151,656,629,652]" gridcol="0" gridrow="3" pageId="7" pageNumber="64">Notes/call</th>
<td id="76AB5DF2BC2F0078FD57FDC2FC1DFD34" box="[753,1010,629,652]" gridcol="1" gridrow="3" pageId="7" pageNumber="64">6.7 (4–11)</td>
<td id="76AB5DF2BC2F0078FBE0FDC2FA73FD34" box="[1094,1436,629,652]" gridcol="2" gridrow="3" pageId="7" pageNumber="64">6.2 (4–11)</td>
</tr>
<tr id="357A348EBC2F0078FF31FD17FA73FD59" box="[151,1436,672,737]" gridrow="4" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FD17FD7FFD59" box="[151,656,672,737]" gridcol="0" gridrow="4" pageId="7" pageNumber="64">Internote interval (ms) Call repetition rate (calls/s)</th>
<td id="76AB5DF2BC2F0078FD57FD17FC1DFD59" box="[753,1010,672,737]" gridcol="1" gridrow="4" pageId="7" pageNumber="64">33.7 (12–82) 0.8</td>
<td id="76AB5DF2BC2F0078FBE0FD17FA73FD59" box="[1094,1436,672,737]" gridcol="2" gridrow="4" pageId="7" pageNumber="64">31.6 (12–62) 0.7</td>
</tr>
<tr id="357A348EBC2F0078FF31FD42FA73FCD9" box="[151,1436,757,865]" gridrow="5" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FD42FD7FFCD9" box="[151,656,757,865]" gridcol="0" gridrow="5" pageId="7" pageNumber="64">Introductory Note duration (ms) note Pulses/note Dominant frequency (Hz).</th>
<td id="76AB5DF2BC2F0078FD57FD42FC1DFCD9" box="[753,1010,757,865]" gridcol="1" gridrow="5" pageId="7" pageNumber="64">58 (24–106) 22.7 (11–40) 3561.6 (2928.5–3789.8)</td>
<td id="76AB5DF2BC2F0078FBE0FD42FA73FCD9" box="[1094,1436,757,865]" gridcol="2" gridrow="5" pageId="7" pageNumber="64">70 (25–132) 26.5 (8–50) 3772.6 (3445.3–3789.8)</td>
</tr>
<tr id="357A348EBC2F0078FF31FCC2FA73FC34" box="[151,1436,885,908]" gridrow="6" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FCC2FD7FFC34" box="[151,656,885,908]" gridcol="0" gridrow="6" pageId="7" pageNumber="64">Pulse repetition rate (pulses/s)</th>
<td id="76AB5DF2BC2F0078FD57FCC2FC1DFC34" box="[753,1010,885,908]" gridcol="1" gridrow="6" pageId="7" pageNumber="64">394 (368–429)</td>
<td id="76AB5DF2BC2F0078FBE0FCC2FA73FC34" box="[1094,1436,885,908]" gridcol="2" gridrow="6" pageId="7" pageNumber="64">363 (326–427)</td>
</tr>
<tr id="357A348EBC2F0078FF31FC17FA73FC59" box="[151,1436,928,993]" gridrow="7" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FC17FD7FFC59" box="[151,656,928,993]" gridcol="0" gridrow="7" pageId="7" pageNumber="64">Clicks Note duration (ms) Pulses/note</th>
<td id="76AB5DF2BC2F0078FD57FC17FC1DFC59" box="[753,1010,928,993]" gridcol="1" gridrow="7" pageId="7" pageNumber="64">7.2 (2–23) 1.2 (1–7)</td>
<td id="76AB5DF2BC2F0078FBE0FC17FA73FC59" box="[1094,1436,928,993]" gridcol="2" gridrow="7" pageId="7" pageNumber="64">7.8 (3–24) 1.2 (1–7)</td>
</tr>
<tr id="357A348EBC2F0078FF31FC42FA73FBB4" box="[151,1436,1013,1036]" gridrow="8" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FC42FD7FFBB4" box="[151,656,1013,1036]" gridcol="0" gridrow="8" pageId="7" pageNumber="64">Dominant frequency (Hz)</th>
<td id="76AB5DF2BC2F0078FD57FC42FC1DFBB4" box="[753,1010,1013,1036]" gridcol="1" gridrow="8" pageId="7" pageNumber="64">3560.4 (2584–3798.8)</td>
<td id="76AB5DF2BC2F0078FBE0FC42FA73FBB4" box="[1094,1436,1013,1036]" gridcol="2" gridrow="8" pageId="7" pageNumber="64">3602.9 (2756.2–3962.1)</td>
</tr>
<tr id="357A348EBC2F0078FF31FB97FA73FB8F" box="[151,1436,1056,1079]" gridrow="9" pageId="7" pageNumber="64">
<th id="76AB5DF2BC2F0078FF31FB97FD7FFB8F" box="[151,656,1056,1079]" gridcol="0" gridrow="9" pageId="7" pageNumber="64">Note repetition rate (notes/s)</th>
<td id="76AB5DF2BC2F0078FD57FB97FC1DFB8F" box="[753,1010,1056,1079]" gridcol="1" gridrow="9" pageId="7" pageNumber="64">24 (16–26)</td>
<td id="76AB5DF2BC2F0078FBE0FB97FA73FB8F" box="[1094,1436,1056,1079]" gridcol="2" gridrow="9" pageId="7" pageNumber="64">23 (15–26)</td>
</tr>
<tr id="357A348EBC2F0078FF31FBEEFA73FBC8" box="[151,1436,1113,1136]" gridrow="10" pageId="7" pageNumber="64" rowspan-1="1" rowspan-2="1">
<th id="76AB5DF2BC2F0078FF31FBEEFD7FFBC8" box="[151,656,1113,1136]" gridcol="0" gridrow="10" pageId="7" pageNumber="64">*holotype</th>
</tr>
</table>
</paragraph>
<paragraph id="8BF536CCBC2FFF80FF63FB15FE78FAEA" blockId="7.[151,1437,1186,2034]" pageId="7" pageNumber="64">
<emphasis id="B93EEADEBC2FFF80FF63FB15FE36FB04" bold="true" box="[197,473,1186,1212]" pageId="7" pageNumber="64">Sequence divergence:</emphasis>
Uncorrected sequence divergences between 
<taxonomicName id="4C4A4D4FBC2FFF80FC51FB14FB9CFB04" box="[1015,1139,1187,1212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2FFF80FC51FB14FB9CFB04" box="[1015,1139,1187,1212]" italics="true" pageId="7" pageNumber="64">L. melicus</emphasis>
</taxonomicName>
(MVZ 258197–258199) and 
<taxonomicName id="4C4A4D4FBC2FFF80FF6CFB7FFEBEFB59" box="[202,337,1224,1249]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B93EEADEBC2FFF80FF6CFB7FFEBEFB59" box="[202,337,1224,1249]" italics="true" pageId="7" pageNumber="64">L. applebyi</emphasis>
</taxonomicName>
(
<typeStatus id="54F1886EBC2FFF80FEC7FB7FFE24FB5A" box="[353,459,1224,1250]" pageId="7" pageNumber="64" type="holotype">holotype</typeStatus>
AMS R 171703, 
<typeStatus id="54F1886EBC2FFF80FD07FB7FFCE8FB5A" box="[673,775,1224,1250]" pageId="7" pageNumber="64" type="paratype">paratype</typeStatus>
AMS R171704) collected from approximately 
<quantity id="4CB29B29BC2FFF80FAE7FB7FFA73FB5A" box="[1345,1436,1224,1250]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="64" unit="km" value="100.0">100 km</quantity>
away were 6.1% at the 16S rRNA gene. This degree of pairwise divergence in the 16S rRNA gene in frogs usually represents differentiation at the species level (
<bibRefCitation id="EFDB4B3DBC2FFF80FCAEFAA5FC0BFA94" author="Vences" box="[776,996,1298,1324]" pageId="7" pageNumber="64" refString="Vences, M., Thomas, M., Bonett, R. M. &amp; Vieites, D. R. (2005) Deciphering amphibian diversity through DNA barcoding: chances and challenges. Philosophical Transactions of the Royal Society London, Ser. B, 360, 1859 - 1868." type="journal article" year="2005">
Vences 
<emphasis id="B93EEADEBC2FFF80FCC2FAA4FC70FA94" box="[868,927,1299,1324]" italics="true" pageId="7" pageNumber="64">et al.</emphasis>
2005
</bibRefCitation>
). There was no intraspecific variation in this gene fragment.
</paragraph>
<paragraph id="8BF536CCBC2FFF80FF63FAEAFECBF9B4" blockId="7.[151,1437,1186,2034]" pageId="7" pageNumber="64">
<emphasis id="B93EEADEBC2FFF80FF63FAEAFED7FACF" bold="true" box="[197,312,1373,1399]" pageId="7" pageNumber="64">Ecology:</emphasis>
All specimens were found in evergreen forest between 
<quantity id="4CB29B29BC2FFF80FC52FAEAFB6EFACF" box="[1012,1153,1373,1399]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="8.5" metricValueMin="6.0" pageId="7" pageNumber="64" unit="m" value="725.0" valueMax="850.0" valueMin="600.0">600–850 m</quantity>
elevation. Males were observed calling on and between rocks and on tree roots, or under leaf litter, less than 
<quantity id="4CB29B29BC2FFF80FB29FA35FB53FA24" box="[1167,1212,1410,1436]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="64" unit="m" value="1.0">1 m</quantity>
from small (&lt;
<quantity id="4CB29B29BC2FFF80FAC9FA35FA73FA24" box="[1391,1436,1410,1436]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="64" unit="m" value="5.0">5 m</quantity>
wide) rocky streams. 
<taxonomicName id="4C4A4D4FBC2FFF80FE32FA1FFD9CFA79" box="[404,627,1448,1473]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2FFF80FE32FA1FFD9CFA79" box="[404,627,1448,1473]" italics="true" pageId="7" pageNumber="64">Leptolalax melicus</emphasis>
</taxonomicName>
was heard calling in both June and October, during and shortly after rainfall, suggesting that breeding in 
<taxonomicName id="4C4A4D4FBC2FFF80FDE7FA7AFD55FA5E" box="[577,698,1485,1510]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2FFF80FDE7FA7AFD55FA5E" box="[577,698,1485,1510]" italics="true" pageId="7" pageNumber="64">L. melicus</emphasis>
</taxonomicName>
occurs throughout the monsoon season (approximately April– November).
</paragraph>
<paragraph id="8BF536CCBC2FFF80FF63F9A0FD83F8AA" blockId="7.[151,1437,1186,2034]" pageId="7" pageNumber="64">
<emphasis id="B93EEADEBC2FFF80FF63F9A0FE26F989" bold="true" box="[197,457,1559,1585]" pageId="7" pageNumber="64">Conservation status:</emphasis>
The seven 
<typeStatus id="54F1886EBC2FFF80FDF5F9AFFD69F98A" box="[595,646,1560,1586]" pageId="7" pageNumber="64">type</typeStatus>
specimens are the only known representatives of the new species. Given the available information, we suggest the species should be considered Data Deficient following IUCN’s Red List categories (IUCN 2001). Being known only from Virachey National Park, its extent of occurrence is unknown but probably extends further into adjoining areas of the Kon Tum Plateau. Virachey National Park is bordered by two other protected areas; Dong Amphan National Protected Area in 
<collectingCountry id="F35D765CBC2FFF80FA96F91AFA85F97F" box="[1328,1386,1709,1735]" name="Laos" pageId="7" pageNumber="64">Laos</collectingCountry>
and Mom Ray Nature Reserve in 
<collectingCountry id="F35D765CBC2FFF80FE53F965FDB3F954" box="[501,604,1746,1772]" name="Vietnam" pageId="7" pageNumber="64">Vietnam</collectingCountry>
, both of which contain evergreen forest at similar elevations that may represent suitable habitat for 
<taxonomicName id="4C4A4D4FBC2FFF80FE4AF94FFD8AF8A9" box="[492,613,1784,1809]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2FFF80FE4AF94FFD8AF8A9" box="[492,613,1784,1809]" italics="true" pageId="7" pageNumber="64">L. melicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF536CCBC2FFF8FFF63F8AAFC15FEFF" blockId="7.[151,1437,1186,2034]" lastBlockId="8.[151,1436,152,327]" lastPageId="8" lastPageNumber="65" pageId="7" pageNumber="64">
<emphasis id="B93EEADEBC2FFF80FF63F8AAFE99F88F" bold="true" box="[197,374,1821,1847]" pageId="7" pageNumber="64">Comparisons:</emphasis>
<taxonomicName id="4C4A4D4FBC2FFF80FED8F8AAFDB0F88E" box="[382,607,1821,1846]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC2FFF80FED8F8AAFDB0F88E" box="[382,607,1821,1846]" italics="true" pageId="7" pageNumber="64">Leptolalax melicus</emphasis>
</taxonomicName>
is distinguished from all other species in the genus by having an offwhite to pale pink ventral surface with diffuse dark brown blotches and distinct white speckling on the chest, belly and throat (
<emphasis id="B93EEADEBC2FFF80FED2F8DFFC7EF839" box="[372,913,1896,1921]" italics="true" pageId="7" pageNumber="64">
<taxonomicName id="4C4A4D4FBC2FFF80FED2F8DFFE1EF839" box="[372,497,1896,1921]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FE59F8DFFD97F839" box="[511,632,1896,1921]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FD23F8DFFCFDF839" box="[645,786,1896,1921]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="bourreti">L. bourreti</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FC87F8DFFC7EF839" box="[801,913,1896,1921]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
</emphasis>
, 
<emphasis id="B93EEADEBC2FFF80FC04F8DFFEFCF81E" italics="true" pageId="7" pageNumber="64">
<taxonomicName id="4C4A4D4FBC2FFF80FC04F8DFFBBBF839" box="[930,1108,1896,1921]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="fuliginosus">L. fuliginosus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FBC4F8DFFB05F839" box="[1122,1258,1896,1921]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FB5EF8DFFA99F839" box="[1272,1398,1896,1921]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FA25F8DFFEFCF81E" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FE87F83AFE4CF81E" box="[289,419,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B93EEADEBC2FFF80FE87F83AFE4CF81E" box="[289,419,1933,1958]" italics="true" pageId="7" pageNumber="64">L. lateralis</emphasis>
</taxonomicName>
, 
<emphasis id="B93EEADEBC2FFF80FE17F83AFBC6F81E" box="[433,1065,1933,1958]" italics="true" pageId="7" pageNumber="64">
<taxonomicName id="4C4A4D4FBC2FFF80FE17F83AFE15F81E" box="[433,506,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FDA0F83AFD54F81E" box="[518,699,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FD61F83AFC84F81E" box="[711,875,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FCD0F83AFBC6F81E" box="[886,1065,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
</emphasis>
, 
<emphasis id="B93EEADEBC2FFF80FB9EF83AFF0DF874" italics="true" pageId="7" pageNumber="64">
<taxonomicName id="4C4A4D4FBC2FFF80FB9EF83AFB4FF81E" box="[1080,1184,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FB0AF83AFAE4F81E" box="[1196,1291,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FAB1F83AFA97F81E" box="[1303,1400,1933,1958]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC2FFF80FA22F83AFF0DF874" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="tamdil">L. tamdil</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C4A4D4FBC2FFF80FEBAF804FE42F874" box="[284,429,1971,1996]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B93EEADEBC2FFF80FEBAF804FE42F874" box="[284,429,1971,1996]" italics="true" pageId="7" pageNumber="64">L. tuberosus</emphasis>
</taxonomicName>
have mostly white or pale grey venters, without white speckling and with or without dark spots or mottling; 
<taxonomicName id="4C4A4D4FBC2FFF80FE14F86FFDD5F849" box="[434,570,2008,2033]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B93EEADEBC2FFF80FE14F86FFDD5F849" box="[434,570,2008,2033]" italics="true" pageId="7" pageNumber="64">L. applebyi</emphasis>
</taxonomicName>
has a dark brownish pink ventral surface with white speckling over the entire ventral surface, including tibiotarsus and both lower and upper arms; 
<taxonomicName id="4C4A4D4FBC20FF8FFBA8FF2FFB7CFF09" box="[1038,1171,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B93EEADEBC20FF8FFBA8FF2FFB7CFF09" box="[1038,1171,152,177]" italics="true" pageId="8" pageNumber="65">L. pluvialis</emphasis>
</taxonomicName>
has a grey venter with dark grey marbling, uniform pale grey throat with speckling around the border; 
<taxonomicName id="4C4A4D4FBC20FF8FFBDDFF0AFAA9FF6E" box="[1147,1350,189,214]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="B93EEADEBC20FF8FFBDDFF0AFAA9FF6E" box="[1147,1350,189,214]" italics="true" pageId="8" pageNumber="65">L. melanoleucus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4A4D4FBC20FF8FFA22FF0AFEBDFF43" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B93EEADEBC20FF8FFA22FF0AFEBDFF43" italics="true" pageId="8" pageNumber="65">L. ventripunctatus</emphasis>
</taxonomicName>
display large patches of distinct black and white marbling, 
<taxonomicName id="4C4A4D4FBC20FF8FFB8EFF55FB51FF43" box="[1064,1214,226,251]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B93EEADEBC20FF8FFB8EFF55FB51FF43" box="[1064,1214,226,251]" italics="true" pageId="8" pageNumber="65">L. heteropus</emphasis>
</taxonomicName>
has a grey venter, speckled with black; 
<taxonomicName id="4C4A4D4FBC20FF8FFE29FEBFFDE9FE99" box="[399,518,264,289]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B93EEADEBC20FF8FFE29FEBFFDE9FE99" box="[399,518,264,289]" italics="true" pageId="8" pageNumber="65">L. maurus</emphasis>
</taxonomicName>
has a black or dark grey brown venter, with indistinct small light areas, and 
<taxonomicName id="4C4A4D4FBC20FF8FFA22FEBFFF20FEFE" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B93EEADEBC20FF8FFA22FEBFFF20FEFE" italics="true" pageId="8" pageNumber="65">L. kecil</emphasis>
</taxonomicName>
has a uniformly dark venter with large, dark orange pectoral glands).
</paragraph>
<caption id="DF356644BC20FF8FFF31FABCFD4CFA3A" httpUri="https://zenodo.org/record/197322/files/figure.png" pageId="8" pageNumber="65" targetBox="[155,1431,384,1264]" targetPageId="8">
<paragraph id="8BF536CCBC20FF8FFF31FABCFD4CFA3A" blockId="8.[151,1437,1291,1410]" pageId="8" pageNumber="65">
<emphasis id="B93EEADEBC20FF8FFF31FABCFECEFA9B" bold="true" box="[151,289,1291,1315]" pageId="8" pageNumber="65">FIGURE 4.</emphasis>
Advertisement call of 
<taxonomicName id="4C4A4D4FBC20FF8FFD8CFABCFCECFA9A" box="[554,771,1291,1314]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="melicus" status="sp. nov.">
<emphasis id="B93EEADEBC20FF8FFD8CFABCFCECFA9A" box="[554,771,1291,1314]" italics="true" pageId="8" pageNumber="65">Leptolalax melicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57A5BC20FF8FFCAAFABCFC87FA9B" box="[780,872,1291,1315]" pageId="8" pageNumber="65" rank="species">
<emphasis id="B93EEADEBC20FF8FFCAAFABCFC87FA9B" bold="true" box="[780,872,1291,1315]" pageId="8" pageNumber="65">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype MVZ 258198) recorded at ambient air temperature of 26.1º C. (A) 60 s waveform of relative amplitude over time, (B) waveform and (C) corresponding spectrogram of single representative call expanded from section shown in A, and (D) power spectrum (relative amplitude vs. frequency) of introductory note and (E) click.
</paragraph>
</caption>
<paragraph id="8BF536CCBC20FF8FFF63FA06FB1BF9D8" blockId="8.[151,1437,1457,2005]" pageId="8" pageNumber="65">
In having finger I &lt;II, 
<taxonomicName id="4C4A4D4FBC20FF8FFE75FA06FDA4FA72" box="[467,587,1457,1482]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC20FF8FFE75FA06FDA4FA72" box="[467,587,1457,1482]" italics="true" pageId="8" pageNumber="65">L. melicus</emphasis>
</taxonomicName>
differs from 
<emphasis id="B93EEADEBC20FF8FFD40FA06FD8EFA57" italics="true" pageId="8" pageNumber="65">
<taxonomicName id="4C4A4D4FBC20FF8FFD40FA06FCBBFA72" box="[742,852,1457,1482]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFCC6FA06FC34FA72" box="[864,987,1457,1482]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFC41FA06FB94FA72" box="[999,1147,1457,1482]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFB21FA06FADDFA72" box="[1159,1330,1457,1482]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="kajangensis">L. kajangensis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFA98FA06FA78FA72" box="[1342,1431,1457,1482]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFF31FA61FEA2FA57" box="[151,333,1494,1519]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFEFFFA61FE2DFA57" box="[345,450,1494,1519]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
, and 
<taxonomicName id="4C4A4D4FBC20FF8FFDA7FA61FD8EFA57" box="[513,609,1494,1519]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
</emphasis>
, which have fingers I and II equal, and from 
<taxonomicName id="4C4A4D4FBC20FF8FFBD1FA61FB3DFA57" box="[1143,1234,1494,1519]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="solus">
<emphasis id="B93EEADEBC20FF8FFBD1FA61FB3DFA57" box="[1143,1234,1494,1519]" italics="true" pageId="8" pageNumber="65">L. solus</emphasis>
</taxonomicName>
, in which Finger I&gt; II. In lacking webbing and dermal fringes on fingers and having only slight basal webbing and no dermal fringes on toes, 
<taxonomicName id="4C4A4D4FBC20FF8FFEF2F996FE22F982" box="[340,461,1569,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC20FF8FFEF2F996FE22F982" box="[340,461,1569,1594]" italics="true" pageId="8" pageNumber="65">L. melicus</emphasis>
</taxonomicName>
differs from 
<emphasis id="B93EEADEBC20FF8FFDCCF996FEC4F9E7" italics="true" pageId="8" pageNumber="65">
<taxonomicName id="4C4A4D4FBC20FF8FFDCCF996FD32F982" box="[618,733,1569,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFD4FF996FC84F982" box="[745,875,1569,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="bourreti">L. bourreti</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFCD1F996FBF4F982" box="[887,1051,1569,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="fuliginosus">L. fuliginosus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFB80F996FB54F982" box="[1062,1211,1569,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFB6EF996FACDF982" box="[1224,1314,1569,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFA88F996FA97F982" box="[1326,1400,1569,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFA22F996FEC4F9E7" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
</emphasis>
, and 
<taxonomicName id="4C4A4D4FBC20FF8FFECDF9F1FE26F9E7" box="[363,457,1606,1631]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B93EEADEBC20FF8FFECDF9F1FE26F9E7" box="[363,457,1606,1631]" italics="true" pageId="8" pageNumber="65">L. sungi</emphasis>
</taxonomicName>
, all of which have lateral fringes or more extensive webbing on toes.
</paragraph>
<paragraph id="8BF536CCBC20FF8FFF63F9DCFC53F833" blockId="8.[151,1437,1457,2005]" pageId="8" pageNumber="65">
The small size of male 
<taxonomicName id="4C4A4D4FBC20FF8FFE7FF9DBFD56F93D" box="[473,697,1644,1669]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC20FF8FFE7FF9DBFD56F93D" box="[473,697,1644,1669]" italics="true" pageId="8" pageNumber="65">Leptolalax melicus</emphasis>
</taxonomicName>
(
<quantity id="4CB29B29BC20FF8FFD6EF9DCFC99F93D" box="[712,886,1643,1669]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" metricValueMax="2.27" metricValueMin="1.95" pageId="8" pageNumber="65" unit="mm" value="21.1" valueMax="22.7" valueMin="19.5">19.5–22.7 mm</quantity>
) further distinguishes it from all but 
<taxonomicName id="4C4A4D4FBC20FF8FFA8AF9DBFA73F93D" box="[1324,1436,1644,1669]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="alpinis">
<emphasis id="B93EEADEBC20FF8FFA8AF9DBFA73F93D" box="[1324,1436,1644,1669]" italics="true" pageId="8" pageNumber="65">L. alpinis</emphasis>
</taxonomicName>
(24.0– 
<quantity id="4CB29B29BC20FF8FFF44F926FEA0F913" box="[226,335,1681,1707]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="8" pageNumber="65" unit="mm" value="26.4">26.4 mm</quantity>
), 
<taxonomicName id="4C4A4D4FBC20FF8FFEC2F926FE07F912" box="[356,488,1681,1706]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B93EEADEBC20FF8FFEC2F926FE07F912" box="[356,488,1681,1706]" italics="true" pageId="8" pageNumber="65">L. applebyi</emphasis>
</taxonomicName>
(
<quantity id="4CB29B29BC20FF8FFE51F926FD47F913" box="[503,680,1681,1707]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" metricValueMax="2.08" metricValueMin="1.96" pageId="8" pageNumber="65" unit="mm" value="20.2" valueMax="20.8" valueMin="19.6">19.6–20.8 mm</quantity>
), 
<taxonomicName id="4C4A4D4FBC20FF8FFD1AF926FCFDF912" box="[700,786,1681,1706]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B93EEADEBC20FF8FFD1AF926FCFDF912" box="[700,786,1681,1706]" italics="true" pageId="8" pageNumber="65">L. kecil</emphasis>
</taxonomicName>
(
<quantity id="4CB29B29BC20FF8FFC84F926FC3DF913" box="[802,978,1681,1707]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.05" metricValueMin="1.93" pageId="8" pageNumber="65" unit="mm" value="19.9" valueMax="20.5" valueMin="19.3">19.3–20.5 mm</quantity>
), 
<taxonomicName id="4C4A4D4FBC20FF8FFC40F926FB6EF912" box="[998,1153,1681,1706]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="khasiorum">
<emphasis id="B93EEADEBC20FF8FFC40F926FB6EF912" box="[998,1153,1681,1706]" italics="true" pageId="8" pageNumber="65">L. khasiorum</emphasis>
</taxonomicName>
(
<quantity id="4CB29B29BC20FF8FFB37F926FAAEF913" box="[1169,1345,1681,1707]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" metricValueMax="2.73" metricValueMin="2.45" pageId="8" pageNumber="65" unit="mm" value="25.9" valueMax="27.3" valueMin="24.5">24.5–27.3 mm</quantity>
), 
<taxonomicName id="4C4A4D4FBC20FF8FFAF3F926FA74F912" box="[1365,1435,1681,1706]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="liui">
<emphasis id="B93EEADEBC20FF8FFAF3F926FA74F912" box="[1365,1435,1681,1706]" italics="true" pageId="8" pageNumber="65">L. liui</emphasis>
</taxonomicName>
(23.0– 
<quantity id="4CB29B29BC20FF8FFF44F901FEBFF968" box="[226,336,1718,1744]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="8" pageNumber="65" unit="mm" value="28.7">28.7 mm</quantity>
), and 
<taxonomicName id="4C4A4D4FBC20FF8FFE30F901FDF5F977" box="[406,538,1718,1743]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B93EEADEBC20FF8FFE30F901FDF5F977" box="[406,538,1718,1743]" italics="true" pageId="8" pageNumber="65">L. pluvialis</emphasis>
</taxonomicName>
(
<quantity id="4CB29B29BC20FF8FFD8DF901FD33F968" box="[555,732,1718,1744]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" metricValueMax="2.23" metricValueMin="2.13" pageId="8" pageNumber="65" unit="mm" value="21.8" valueMax="22.3" valueMin="21.3">21.3–22.3 mm</quantity>
). All other congeners have distinctly larger male body sizes (
<taxonomicName id="4C4A4D4FBC20FF8FFF07F96BFEE1F94D" box="[161,270,1756,1781]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="arayai">
<emphasis id="B93EEADEBC20FF8FFF07F96BFEE1F94D" box="[161,270,1756,1781]" italics="true" pageId="8" pageNumber="65">L. arayai</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFEB3F96CFE6DF94D" box="[277,386,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="8" pageNumber="65" unit="mm" value="29.6">29.6 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFE29F96BFDE0F94D" box="[399,527,1756,1781]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="B93EEADEBC20FF8FFE29F96BFDE0F94D" box="[399,527,1756,1781]" italics="true" pageId="8" pageNumber="65">L. bourreti</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFDB1F96CFD6CF94D" box="[535,643,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="8" pageNumber="65" unit="mm" value="36.2">36.2 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFD36F96BFD17F94D" box="[656,760,1756,1781]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B93EEADEBC20FF8FFD36F96BFD17F94D" box="[656,760,1756,1781]" italics="true" pageId="8" pageNumber="65">L. dringi</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFCA7F96CFC5EF94D" box="[769,945,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.03" metricValueMin="2.87" pageId="8" pageNumber="65" unit="mm" value="29.5" valueMax="30.3" valueMin="28.7">28.7–30.3 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFC1AF96BFBB1F94D" box="[956,1118,1756,1781]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B93EEADEBC20FF8FFC1AF96BFBB1F94D" box="[956,1118,1756,1781]" italics="true" pageId="8" pageNumber="65">L. fuliginosus</emphasis>
</taxonomicName>
28.2–30.0 mm; 
<taxonomicName id="4C4A4D4FBC20FF8FFA84F96BFA73F94D" box="[1314,1436,1756,1781]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B93EEADEBC20FF8FFA84F96BFA73F94D" box="[1314,1436,1756,1781]" italics="true" pageId="8" pageNumber="65">L. gracilis</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFF31F8B6FEF5F8A3" box="[151,282,1793,1819]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.6" metricValueMin="3.0" pageId="8" pageNumber="65" unit="mm" value="33.0" valueMax="36.0" valueMin="30.0">30–36 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFE80F8B6FE77F8A2" box="[294,408,1793,1818]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B93EEADEBC20FF8FFE80F8B6FE77F8A2" box="[294,408,1793,1818]" italics="true" pageId="8" pageNumber="65">L. hamidi</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFE06F8B6FDA0F8A3" box="[416,591,1793,1819]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.13" metricValueMin="2.87" pageId="8" pageNumber="65" unit="mm" value="30.0" valueMax="31.3" valueMin="28.7">28.7–31.3 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFDFAF8B6FD00F8A2" box="[604,751,1793,1818]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B93EEADEBC20FF8FFDFAF8B6FD00F8A2" box="[604,751,1793,1818]" italics="true" pageId="8" pageNumber="65">L. heteropus</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFD50F8B6FCA1F8A3" box="[758,846,1793,1819]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="65" unit="mm" value="33.0">33 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFCFCF8B6FBEAF8A2" box="[858,1029,1793,1818]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="B93EEADEBC20FF8FFCFCF8B6FBEAF8A2" box="[858,1029,1793,1818]" italics="true" pageId="8" pageNumber="65">L. kajangensis</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFBABF8B6FB60F8A3" box="[1037,1167,1793,1819]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="8" pageNumber="65" unit="mm" value="34.5" valueMax="35.0" valueMin="34.0">34–35 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFB3AF8B6FAF1F8A2" box="[1180,1310,1793,1818]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B93EEADEBC20FF8FFB3AF8B6FAF1F8A2" box="[1180,1310,1793,1818]" italics="true" pageId="8" pageNumber="65">L. lateralis</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFA80F8B6FF27F8F8" metricMagnitude="-2" metricUnit="m" metricValue="2.76" metricValueMax="2.83" metricValueMin="2.69" pageId="8" pageNumber="65" unit="mm" value="27.6" valueMax="28.3" valueMin="26.9">26.9–28.3 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFF72F891FEA3F887" box="[212,332,1830,1855]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B93EEADEBC20FF8FFF72F891FEA3F887" box="[212,332,1830,1855]" italics="true" pageId="8" pageNumber="65">L. maurus</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFEF5F891FE50F8F8" box="[339,447,1830,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="8" pageNumber="65" unit="mm" value="26.1">26.1 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFE6DF891FD63F887" box="[459,652,1830,1855]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="B93EEADEBC20FF8FFE6DF891FD63F887" box="[459,652,1830,1855]" italics="true" pageId="8" pageNumber="65">L. melanoleucus</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFD35F891FCAEF8F8" box="[659,833,1830,1856]" metricMagnitude="-2" metricUnit="m" metricValue="2.77" metricValueMax="2.88" metricValueMin="2.66" pageId="8" pageNumber="65" unit="mm" value="27.7" valueMax="28.8" valueMin="26.6">26.6–28.8 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFCEBF891FC10F887" box="[845,1023,1830,1855]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="B93EEADEBC20FF8FFCEBF891FC10F887" box="[845,1023,1830,1855]" italics="true" pageId="8" pageNumber="65">L. nahangensis</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFBA1F891FB9CF8F8" box="[1031,1139,1830,1856]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="8" pageNumber="65" unit="mm" value="40.8">40.8 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFBD9F891FAF1F887" box="[1151,1310,1830,1855]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B93EEADEBC20FF8FFBD9F891FAF1F887" box="[1151,1310,1830,1855]" italics="true" pageId="8" pageNumber="65">L. oshanensis</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFA81F891FF26F8DD" metricMagnitude="-2" metricUnit="m" metricValue="2.8649999999999998" metricValueMax="3.07" metricValueMin="2.66" pageId="8" pageNumber="65" unit="mm" value="28.65" valueMax="30.7" valueMin="26.6">26.6–30.7 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFF73F8FBFE65F8DD" box="[213,394,1868,1893]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B93EEADEBC20FF8FFF73F8FBFE65F8DD" box="[213,394,1868,1893]" italics="true" pageId="8" pageNumber="65">L. pelodytoides</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFE34F8FCFE06F8DD" box="[402,489,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="65" unit="mm" value="37.0">37 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFE53F8FBFDB4F8DD" box="[501,603,1868,1893]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B93EEADEBC20FF8FFE53F8FBFDB4F8DD" box="[501,603,1868,1893]" italics="true" pageId="8" pageNumber="65">L. pictus</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFDC2F8FCFD08F8DD" box="[612,743,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="8" pageNumber="65" unit="mm" value="32.5" valueMax="34.0" valueMin="31.0">31–34 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFD55F8FBFCA0F8DD" box="[755,847,1868,1893]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="solus">
<emphasis id="B93EEADEBC20FF8FFD55F8FBFCA0F8DD" box="[755,847,1868,1893]" italics="true" pageId="8" pageNumber="65">L. solus</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFCFEF8FCFC2BF8DD" box="[856,964,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="8" pageNumber="65" unit="mm" value="27.6">27.6 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFC77F8FBFBDEF8DD" box="[977,1073,1868,1893]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B93EEADEBC20FF8FFC77F8FBFBDEF8DD" box="[977,1073,1868,1893]" italics="true" pageId="8" pageNumber="65">L. sungi</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFB9FF8FCFB06F8DD" box="[1081,1257,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.27" metricValueMin="4.83" pageId="8" pageNumber="65" unit="mm" value="50.5" valueMax="52.7" valueMin="48.3">48.3–52.7 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFB53F8FBFA8FF8DD" box="[1269,1376,1868,1893]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B93EEADEBC20FF8FFB53F8FBFA8FF8DD" box="[1269,1376,1868,1893]" italics="true" pageId="8" pageNumber="65">L. tamdil</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFACEF8FCFF27F833" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="8" pageNumber="65" unit="mm" value="32.3">32.3 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFF72F8C6FE8BF832" box="[212,356,1905,1930]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B93EEADEBC20FF8FFF72F8C6FE8BF832" box="[212,356,1905,1930]" italics="true" pageId="8" pageNumber="65">L. tuberosus</emphasis>
</taxonomicName>
<quantity id="4CB29B29BC20FF8FFECDF8C6FDF7F833" box="[363,536,1905,1931]" metricMagnitude="-2" metricUnit="m" metricValue="2.695" metricValueMax="2.95" metricValueMin="2.44" pageId="8" pageNumber="65" unit="mm" value="26.95" valueMax="29.5" valueMin="24.4">24.4–29.5 mm</quantity>
; 
<taxonomicName id="4C4A4D4FBC20FF8FFD82F8C6FD15F832" box="[548,762,1905,1930]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B93EEADEBC20FF8FFD82F8C6FD15F832" box="[548,762,1905,1930]" italics="true" pageId="8" pageNumber="65">L. ventripunctatus</emphasis>
</taxonomicName>
25.5–28.0 mm).
</paragraph>
<paragraph id="8BF536CCBC20FF8EFF63F821FBE5FF6F" blockId="8.[151,1437,1457,2005]" lastBlockId="9.[151,1437,152,738]" lastPageId="9" lastPageNumber="66" pageId="8" pageNumber="65">
<taxonomicName id="4C4A4D4FBC20FF8FFF63F821FE57F817" box="[197,440,1942,1967]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC20FF8FFF63F821FE57F817" box="[197,440,1942,1967]" italics="true" pageId="8" pageNumber="65">Leptolalax melicus</emphasis>
</taxonomicName>
is also differentiated from 
<emphasis id="B93EEADEBC20FF8FFCBCF821FD85F86D" italics="true" pageId="8" pageNumber="65">
<taxonomicName id="4C4A4D4FBC20FF8FFCBCF821FC77F817" box="[794,920,1942,1967]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFC00F821FBB5F817" box="[934,1114,1942,1967]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="fuliginosus">L. fuliginosus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFBCEF821FAFAF817" box="[1128,1301,1942,1967]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFA84F821FA9BF817" box="[1314,1396,1942,1967]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFA25F821FEF4F86D" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFE83F80BFE34F86D" box="[293,475,1980,2005]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC20FF8FFE40F80BFD85F86D" box="[486,618,1980,2005]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="pluvialis">L. pluvialis</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C4A4D4FBC20FF8FFD04F80BFCE4F86D" box="[674,779,1980,2005]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="65" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B93EEADEBC20FF8FFD04F80BFCE4F86D" box="[674,779,1980,2005]" italics="true" pageId="8" pageNumber="65">L. tamdil</emphasis>
</taxonomicName>
by the absence of ventrolateral glandular lines and from 
<emphasis id="B93EEADEBC21FF8EFF31FF2FFB8AFF09" box="[151,1125,152,177]" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFF31FF2FFEE6FF09" box="[151,265,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFEB0FF2FFE56FF09" box="[278,441,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFE63FF2FFDA2FF09" box="[453,589,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="lateralis">L. lateralis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFDFCFF2FFD37FF09" box="[602,728,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="maurus">L. maurus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFD43FF2FFCA9FF09" box="[741,838,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFCF5FF2FFC2BFF09" box="[851,964,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="tamdil">L. tamdil</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFC76FF2FFB8AFF09" box="[976,1125,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="tuberosus">L. tuberosus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C4A4D4FBC21FF8EFB07FF2FFA93FF09" box="[1185,1404,152,177]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B93EEADEBC21FF8EFB07FF2FFA93FF09" box="[1185,1404,152,177]" italics="true" pageId="9" pageNumber="66">L. ventripunctatus</emphasis>
</taxonomicName>
in having mostly smooth (versus tuberculate) skin texture with no skin ridges.
</paragraph>
<paragraph id="8BF536CCBC21FF8EFF63FF55FA8CFE2A" blockId="9.[151,1437,152,738]" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFF63FF55FE5CFF43" box="[197,435,226,251]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC21FF8EFF63FF55FE5CFF43" box="[197,435,226,251]" italics="true" pageId="9" pageNumber="66">Leptolalax melicus</emphasis>
</taxonomicName>
can be further distinguished from the most morphologically similar species, 
<emphasis id="B93EEADEBC21FF8EFA25FF55FEE9FE99" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFA25FF55FEEDFE99" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
,
</emphasis>
in having a more distinct dorsal patterning, significantly larger pectoral glands (
<taxonomicName id="4C4A4D4FBC21FF8EFB71FEBFFABDFE99" box="[1239,1362,264,289]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC21FF8EFB71FEBFFABDFE99" box="[1239,1362,264,289]" italics="true" pageId="9" pageNumber="66">L. melicus</emphasis>
</taxonomicName>
mean diameter 
<quantity id="4CB29B29BC21FF8EFEA1FE9AFE8CFEFF" box="[263,355,301,327]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="66" unit="mm" value="0.9">0.9 mm</quantity>
± 
<geoCoordinate id="EE7E500BBC21FF8EFE22FE9AFE3BFEFF" box="[388,468,301,327]" direction="south" orientation="latitude" pageId="9" pageNumber="66" precision="555" value="-0.07">0.07 S</geoCoordinate>
. E., 
<taxonomicName id="4C4A4D4FBC21FF8EFDACFE9AFD7DFEFE" box="[522,658,301,326]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B93EEADEBC21FF8EFDACFE9AFD7DFEFE" box="[522,658,301,326]" italics="true" pageId="9" pageNumber="66">L. applebyi</emphasis>
</taxonomicName>
mean diameter 
<quantity id="4CB29B29BC21FF8EFCF5FE9AFC40FEFF" box="[851,943,301,327]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="66" unit="mm" value="0.4">0.4 mm</quantity>
± 
<geoCoordinate id="EE7E500BBC21FF8EFC76FE9AFBF0FEFF" box="[976,1055,301,327]" direction="south" orientation="latitude" pageId="9" pageNumber="66" precision="555" value="-0.03">0.03 S</geoCoordinate>
. E.; Mann-Whitney 
<emphasis id="B93EEADEBC21FF8EFAB3FE9AFAC5FEFE" box="[1301,1322,301,326]" italics="true" pageId="9" pageNumber="66">U</emphasis>
-test, 
<emphasis id="B93EEADEBC21FF8EFACBFE9AFA7DFEFE" box="[1389,1426,301,326]" italics="true" pageId="9" pageNumber="66">Z=</emphasis>
- 2.847, 
<emphasis id="B93EEADEBC21FF8EFF4EFEE5FF18FED3" box="[232,247,338,363]" italics="true" pageId="9" pageNumber="66">p</emphasis>
= 0.004; N=12), and significantly larger femoral glands (
<taxonomicName id="4C4A4D4FBC21FF8EFC3DFEE5FBFCFED3" box="[923,1043,338,363]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC21FF8EFC3DFEE5FBFCFED3" box="[923,1043,338,363]" italics="true" pageId="9" pageNumber="66">L. melicus</emphasis>
</taxonomicName>
mean diameter 
<quantity id="4CB29B29BC21FF8EFB76FEE5FAC5FED4" box="[1232,1322,338,364]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="66" unit="mm" value="0.9">0.9 mm</quantity>
± 
<geoCoordinate id="EE7E500BBC21FF8EFAEFFEE5FA78FED4" box="[1353,1431,338,364]" direction="south" orientation="latitude" pageId="9" pageNumber="66" precision="555" value="-0.08">0.08 S</geoCoordinate>
. E., 
<taxonomicName id="4C4A4D4FBC21FF8EFF19FECFFEACFE29" box="[191,323,376,401]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B93EEADEBC21FF8EFF19FECFFEACFE29" box="[191,323,376,401]" italics="true" pageId="9" pageNumber="66">L. applebyi</emphasis>
</taxonomicName>
mean diameter 
<quantity id="4CB29B29BC21FF8EFE58FECFFDB7FE2A" box="[510,600,376,402]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="66" unit="mm" value="0.5">0.5 mm</quantity>
± 
<geoCoordinate id="EE7E500BBC21FF8EFDD8FECFFD24FE2A" box="[638,715,376,402]" direction="south" orientation="latitude" pageId="9" pageNumber="66" precision="555" value="-0.05">0.05 S</geoCoordinate>
. E.; Mann-Whitney 
<emphasis id="B93EEADEBC21FF8EFC1DFECFFC3FFE29" box="[955,976,376,401]" italics="true" pageId="9" pageNumber="66">U</emphasis>
-test, 
<emphasis id="B93EEADEBC21FF8EFBB7FECFFBDAFE29" box="[1041,1077,376,401]" italics="true" pageId="9" pageNumber="66">Z=</emphasis>
-2.847, 
<emphasis id="B93EEADEBC21FF8EFB29FECFFB71FE29" box="[1167,1182,376,401]" italics="true" pageId="9" pageNumber="66">p</emphasis>
= 0.004; N=12).
</paragraph>
<paragraph id="8BF536CCBC21FF8EFF63FE2AFB0FFD5A" blockId="9.[151,1437,152,738]" pageId="9" pageNumber="66">
The advertisement call of 
<taxonomicName id="4C4A4D4FBC21FF8EFDB7FE2AFD7BFE0E" box="[529,660,413,438]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC21FF8EFDB7FE2AFD7BFE0E" box="[529,660,413,438]" italics="true" pageId="9" pageNumber="66">L. melicus</emphasis>
</taxonomicName>
differs structurally from all fourteen 
<taxonomicName id="4C4A4D4FBC21FF8EFBD4FE2AFB16FE0E" box="[1138,1273,413,438]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="B93EEADEBC21FF8EFBD4FE2AFB16FE0E" box="[1138,1273,413,438]" italics="true" pageId="9" pageNumber="66">Leptolalax</emphasis>
</taxonomicName>
species with described calls; 
<emphasis id="B93EEADEBC21FF8EFEF2FE75FDB8FE63" box="[340,599,450,475]" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFEF2FE75FE29FE63" box="[340,454,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFE75FE75FDB8FE63" box="[467,599,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
</emphasis>
, 
<emphasis id="B93EEADEBC21FF8EFDC0FE75FD12FDB9" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFDC0FE75FD3AFE63" box="[614,725,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFD47FE75FCA5FE63" box="[737,842,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFCF1FE75FC01FE63" box="[855,1006,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="fuliginous">L. fuliginous</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFC5DFE75FB99FE63" box="[1019,1142,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFB25FE75FB19FE63" box="[1155,1270,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFAA4FE75FA78FE63" box="[1282,1431,450,475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFF31FE5FFF1DFDB9" box="[151,242,488,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFF58FE5FFE2BFDB9" box="[254,452,488,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFE77FE5FFD66FDB9" box="[465,649,488,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFD30FE5FFD12FDB9" box="[662,765,488,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C4A4D4FBC21FF8EFCAAFE5FFC40FDB9" box="[780,943,488,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B93EEADEBC21FF8EFCAAFE5FFC40FDB9" box="[780,943,488,513]" italics="true" pageId="9" pageNumber="66">L. oshanensis</emphasis>
</taxonomicName>
and 
<emphasis id="B93EEADEBC21FF8EFC4CFE5FFBA1FDB9" box="[1002,1102,488,513]" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFC4CFE5FFBA6FDB9" box="[1002,1097,488,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
,
</emphasis>
and also from 
<taxonomicName id="4C4A4D4FBC21FF8EFAA5FE5FFA7AFDB9" box="[1283,1429,488,513]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B93EEADEBC21FF8EFAA5FE5FFA7AFDB9" box="[1283,1429,488,513]" italics="true" pageId="9" pageNumber="66">L. tuberosus</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C4A4D4FBC21FF8EFF31FDBAFE93FD9E" box="[151,380,525,550]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B93EEADEBC21FF8EFF31FDBAFE93FD9E" box="[151,380,525,550]" italics="true" pageId="9" pageNumber="66">Leptolalax melicus</emphasis>
</taxonomicName>
is the only species of 
<taxonomicName id="4C4A4D4FBC21FF8EFD2BFDBAFCE1FD9E" box="[653,782,525,550]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="B93EEADEBC21FF8EFD2BFDBAFCE1FD9E" box="[653,782,525,550]" italics="true" pageId="9" pageNumber="66">Leptolalax</emphasis>
</taxonomicName>
with a call containing a single long introductory note containing 8–50 pulses, followed by a series of predominantly single-pulse notes or clicks. All other described calls of 
<taxonomicName id="4C4A4D4FBC21FF8EFF5FFDEFFE94FDC9" box="[249,379,600,625]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="B93EEADEBC21FF8EFF5FFDEFFE94FDC9" box="[249,379,600,625]" italics="true" pageId="9" pageNumber="66">Leptolalax</emphasis>
</taxonomicName>
species contain notes that are similar in structure and duration. The only species with noticeably different first notes in the advertisement call are 
<taxonomicName id="4C4A4D4FBC21FF8EFC38FDCAFBF5FD2E" box="[926,1050,637,662]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B93EEADEBC21FF8EFC38FDCAFBF5FD2E" box="[926,1050,637,662]" italics="true" pageId="9" pageNumber="66">L. hamidi</emphasis>
</taxonomicName>
and 
<emphasis id="B93EEADEBC21FF8EFBFCFDCAFAEDFD2E" box="[1114,1282,637,662]" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFBFCFDCAFB12FD2E" box="[1114,1277,637,662]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
,
</emphasis>
which have introductory notes of a slightly longer duration but similar amplitude to successive notes, and 
<emphasis id="B93EEADEBC21FF8EFB7DFD15FAA2FD03" box="[1243,1357,674,699]" italics="true" pageId="9" pageNumber="66">
<taxonomicName id="4C4A4D4FBC21FF8EFB7DFD15FAA7FD03" box="[1243,1352,674,699]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="66" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
,
</emphasis>
which has introductory notes of a slightly shorter duration but similar amplitude to successive notes.
</paragraph>
</subSubSection>
</treatment>
</document>