treatments-xml/data/03/DF/BE/03DFBE05E05EFFB1FF64D7EDFAA7FBF1.xml
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<document id="001B95B7B6F1869CE3BC4BC8E347CA04" ID-CLB-Dataset="8721" ID-DOI="10.11646/zootaxa.4823.1.1" ID-GBIF-Dataset="a43d0118-9df5-4970-a316-0dae5c9caebf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401916" ID-ZooBank="3DF06F26-FD35-452E-9056-1D05B49946EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609340622775" checkinUser="plazi" docAuthor="Dombrow, Holger E. &amp; Colville, Jonathan F." docDate="2020" docId="03DFBE05E05EFFB1FF64D7EDFAA7FBF1" docLanguage="en" docName="zootaxa.4823.1.1.pdf" docOrigin="Zootaxa 4823" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Beckhoplia bicolor Dombrow 2020, new species" docType="treatment" docUuid="4CC61FC4-17AB-4996-98E5-0BE7A186C663" docUuidSource="ZooBank" docVersion="9" lastPageNumber="69" masterDocId="FFE6C67DE01CFFF5FFF3D41BFFBDFF8D" masterDocTitle="Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="67" updateTime="1698849823920" updateUser="ExternalLinkService">
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<mods:title id="946EAF642B0F2FD6FA573B36A5AA7A83">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
</mods:titleInfo>
<mods:name id="6B2A7FF0D6F1EA6D311DF88DA40006CF" type="personal">
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<mods:namePart id="93115AD009711EDE7DA5E61984BACA3F">Dombrow, Holger E.</mods:namePart>
<mods:nameIdentifier id="AD4B4D4ECA02010430F67AB3D3B664E1" type="ORCID">0000-0003-2176-3077</mods:nameIdentifier>
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<mods:namePart id="1950D094B2CD65554CB92C3B24DE801C">Colville, Jonathan F.</mods:namePart>
<mods:nameIdentifier id="E46EF4E9CA5A00485F104C76EC3B0760" type="ORCID">0000-0003-2176-3077</mods:nameIdentifier>
<mods:nameIdentifier id="6CC74A113DF16672EED2F71F89865850" type="email">jonathan.colville@gmail.com</mods:nameIdentifier>
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<mods:title id="6461C99FB3718754577142F29E9B9A3B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0AE0EADE33969A6EB1CFF00EDF1C9740">
<mods:date id="281145983A63F6863ABE64C0499019A0">2020</mods:date>
<mods:detail id="DCC6A5D31EE5C8518900D8F69AC568E0" type="pubDate">
<mods:number id="D6AA578087F3769A9EB202417E0D146E">2020-08-10</mods:number>
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<mods:number id="35D320DB847CB79D1053DFBE20E3ABD0">4823</mods:number>
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<mods:classification id="F19C9B91492052FB03CFECDF7EB3255B">journal article</mods:classification>
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<mods:identifier id="EE3A2039FF9D9850E549EB673E2B9902" type="DOI">10.11646/zootaxa.4823.1.1</mods:identifier>
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<treatment id="03DFBE05E05EFFB1FF64D7EDFAA7FBF1" ID-DOI="http://doi.org/10.5281/zenodo.4455309" ID-GBIF-Taxon="177013515" ID-Zenodo-Dep="4455309" LSID="urn:lsid:zoobank.org:act:4CC61FC4-17AB-4996-98E5-0BE7A186C663" httpUri="http://treatment.plazi.org/id/03DFBE05E05EFFB1FF64D7EDFAA7FBF1" lastPageId="68" lastPageNumber="69" pageId="66" pageNumber="67">
<subSubSection id="C36C5C98E05EFFB7FF64D7EDFD20FB9D" box="[151,669,1014,1041]" pageId="66" pageNumber="67" type="nomenclature">
<paragraph id="8BC90F13E05EFFB7FF64D7EDFD20FB9D" blockId="66.[151,862,1014,1112]" box="[151,669,1014,1041]" pageId="66" pageNumber="67">
<heading id="D081B87FE05EFFB7FF64D7EDFD20FB9D" bold="true" box="[151,669,1014,1041]" fontSize="11" level="2" pageId="66" pageNumber="67" reason="4">
<emphasis id="B902D301E05EFFB7FF64D7EDFD20FB9D" bold="true" box="[151,669,1014,1041]" pageId="66" pageNumber="67">
<taxonomicName id="4C767490E05EFFB7FF64D7EDFDBDFB9D" authority="Dombrow" authorityName="Dombrow" authorityYear="2020" box="[151,512,1014,1041]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="B902D301E05EFFB7FF64D7EDFEC1FB9C" bold="true" box="[151,380,1014,1041]" italics="true" pageId="66" pageNumber="67">Beckhoplia bicolor</emphasis>
Dombrow
</taxonomicName>
,
<taxonomicNameLabel id="A2316E7AE05EFFB7FDFED7EDFD20FB9D" box="[525,669,1014,1040]" pageId="66" pageNumber="67" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E05EFFB7FF64D001FCE3FBD5" pageId="66" pageNumber="67" type="description">
<paragraph id="8BC90F13E05EFFB7FF64D001FE3BFBB9" blockId="66.[151,862,1014,1112]" box="[151,390,1050,1077]" pageId="66" pageNumber="67">
(
<figureCitation id="134D1396E05EFFB7FF6CD001FF43FBB9" box="[159,254,1050,1076]" captionStart="FIGURES 3940" captionStartId="23.[151,264,749,773]" captionTargetBox="[432,1159,536,689]" captionTargetId="figure-79@23.[423,1164,532,700]" captionTargetPageId="23" captionText="FIGURES 3940. General drawings of the tip of pygidium of Beckhoplia Dombrow, 2005 species in posterior view ♂♂. 39, tip linear; 40, tip rounded. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401942" httpUri="https://zenodo.org/record/4401942/files/figure.png" pageId="66" pageNumber="67">Figs. 40</figureCitation>
,
<figureCitation id="134D1396E05EFFB7FEFAD001FEECFBB8" box="[265,337,1050,1077]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="66" pageNumber="67">9495</figureCitation>
,
<figureCitation id="134D1396E05EFFB7FEAED001FE3BFBB9" box="[349,390,1050,1076]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="66" pageNumber="67">116</figureCitation>
</paragraph>
<paragraph id="8BC90F13E05EFFB7FF64D05BFCE3FBD5" blockId="66.[151,862,1014,1112]" box="[151,862,1088,1112]" pageId="66" pageNumber="67">
http://zoobank.org/
<uuid id="FFD035C6E05EFFB7FE9BD05BFCE3FBD5" box="[360,862,1088,1112]" pageId="66" pageNumber="67">4CC61FC4-17AB-4996-98E5-0BE7A186C663</uuid>
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E05EFFB7FF64D093FD3FFA6B" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="8BC90F13E05EFFB7FF64D093FC1DFB2F" blockId="66.[151,928,1159,1187]" box="[151,928,1159,1187]" pageId="66" pageNumber="67">
<materialsCitation id="3B1E054EE05EFFB7FF64D093FC21FB2F" ID-GBIF-Occurrence="3022772312" box="[151,924,1159,1187]" collectionCode="NCP" country="South Africa" location="RSA" pageId="66" pageNumber="67" specimenCode="R357" specimenCount="1" typeStatus="holotype">
<emphasis id="B902D301E05EFFB7FF64D093FE8BFB2F" bold="true" box="[151,310,1160,1186]" pageId="66" pageNumber="67">
<typeStatus id="54CDB1B1E05EFFB7FF64D093FF6FFB2F" box="[151,210,1160,1186]" pageId="66" pageNumber="67">Type</typeStatus>
locality.
</emphasis>
[
<collectingCountry id="F3614F83E05EFFB7FEB0D093FEC2FB2F" box="[323,383,1160,1186]" name="South Africa" pageId="66" pageNumber="67">RSA</collectingCountry>
,
<collectionCode id="ED6797D6E05EFFB7FE79D09CFE7AFB2F" box="[394,455,1159,1186]" country="United Kingdom" name="National Collection of Passiflora" pageId="66" pageNumber="67" type="Herbarium">NCP</collectionCode>
]:
<specimenCode id="DBD0A768E05EFFB7FE2AD093FDA5FB2E" box="[473,536,1160,1187]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="66" pageNumber="67">R357</specimenCode>
, Nieuwoudtville to Loeriesfontein
</materialsCitation>
.
</paragraph>
<paragraph id="8BC90F13E05EFFB7FF64D0CBFD70FAF7" blockId="66.[151,1437,1231,1402]" pageId="66" pageNumber="67">
<emphasis id="B902D301E05EFFB7FF64D0CBFEF4FB67" bold="true" box="[151,329,1232,1258]" pageId="66" pageNumber="67">Type material.</emphasis>
[
<materialsCitation id="3B1E054EE05EFFB7FEADD0D4FD70FAF7" ID-GBIF-Occurrence="3022772307" collectingDate="2012-08-27" collectionCode="NCP, SAMC" collectorName="Dombrow" country="South Africa" determinerName="H. Dombrow" latitude="-31.294167" location="Nieuwoudtville" longLatPrecision="20" longitude="19.135555" pageId="66" pageNumber="67" specimenCode="R357" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
1]
<emphasis id="B902D301E05EFFB7FE73D0CBFE50FB67" bold="true" box="[384,493,1232,1258]" pageId="66" pageNumber="67">
<typeStatus id="54CDB1B1E05EFFB7FE73D0CBFE50FB67" box="[384,493,1232,1258]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D70C49AE05EFFB7FE0AD0CBFDACFB67" box="[505,529,1232,1258]" pageId="66" pageNumber="67" type="male"></specimenCount>
: white label pr «
<collectingCountry id="F3614F83E05EFFB7FD28D0CBFCABFB67" box="[731,790,1232,1258]" name="South Africa" pageId="66" pageNumber="67">RSA</collectingCountry>
,
<collectionCode id="ED6797D6E05EFFB7FCD6D0CBFCE6FB67" box="[805,859,1232,1258]" country="United Kingdom" name="National Collection of Passiflora" pageId="66" pageNumber="67" type="Herbarium">NCP</collectionCode>
,
<specimenCode id="DBD0A768E05EFFB7FC98D0CBFC15FB66" box="[875,936,1232,1259]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="66" pageNumber="67">R357</specimenCode>
to Loeriesfont. |
<quantity id="4C8EA2F6E05EFFB7FB8BD0CBFB0BFB67" box="[1144,1206,1232,1259]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="66" pageNumber="67" unit="km" value="5.0">5 km</quantity>
E
<location id="8EA959C8E05EFFB7FB2ED0CBFA36FB67" LSID="urn:lsid:plazi:treatment:03DFBE05E05EFFB1FF64D7EDFAA7FBF1:8EA959C8E05EFFB7FB2ED0CBFA36FB67" box="[1245,1419,1232,1258]" country="South Africa" latitude="-31.294167" longLatPrecision="20" longitude="19.135555" name="Nieuwoudtville" pageId="66" pageNumber="67">Nieuwoudtville</location>
|
<geoCoordinate id="EE4269D4E05EFFB7FF64D0EFFE90FA83" box="[151,301,1268,1294]" degrees="31" direction="south" minutes="17" orientation="latitude" pageId="66" pageNumber="67" precision="15" seconds="39" value="-31.294167">31°1739S</geoCoordinate>
<geoCoordinate id="EE4269D4E05EFFB7FEDED0EFFE05FA83" box="[301,440,1268,1294]" degrees="19" direction="east" minutes="08" orientation="longitude" pageId="66" pageNumber="67" precision="15" seconds="08" value="19.135555">19°0808E</geoCoordinate>
|
<date id="FFC829D3E05EFFB7FE3DD0EFFDFCFA83" box="[462,577,1268,1294]" pageId="66" pageNumber="67" value="2012-08-27">
<collectingDate id="EF8CD03BE05EFFB7FE3DD0EFFDFCFA83" box="[462,577,1268,1294]" pageId="66" pageNumber="67" value="2012-08-27">27.8.2012</collectingDate>
</date>
, lg. Colville/Melin» || green label pr «Host plant: |
<taxonomicName id="4C767490E05EFFB7FB7BD0EFFA87FA83" authorityName="Aiton" authorityYear="1789" box="[1160,1338,1268,1294]" class="Adenophorea" family="Linhomoeidae" genus="Didelta" kingdom="Animalia" order="Monhysterida" pageId="66" pageNumber="67" phylum="Nematoda" rank="species" species="spinosa">
<emphasis id="B902D301E05EFFB7FB7BD0EFFA87FA83" box="[1160,1338,1268,1294]" italics="true" pageId="66" pageNumber="67">Didelta spinosa</emphasis>
</taxonomicName>
» || blue label pr «Bioregion: | Sandstone Fynbos | Vegetation Type: FFs1 | Bokkeveld Sandstone Fynbos» || yellow label hw «50» pr «447 | coll.
<collectorName id="26836AC5E05EFFB7FE6DD127FDAFFADB" box="[414,530,1340,1366]" pageId="66" pageNumber="67">Dombrow</collectorName>
» || yellow label pr «SAMCOL | A071218» || red label pr «
<typeStatus id="54CDB1B1E05EFFB7FB51D127FAB1FADB" box="[1186,1292,1340,1366]" pageId="66" pageNumber="67" type="holotype">Holotype</typeStatus>
|
<taxonomicName id="4C767490E05EFFB7FAEDD127FA21FADB" authorityName="Dombrow" authorityYear="2005" box="[1310,1436,1340,1366]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E05EFFB7FAEDD127FA21FADB" box="[1310,1436,1340,1366]" italics="true" pageId="66" pageNumber="67">Beckhoplia</emphasis>
</taxonomicName>
|
<taxonomicName id="4C767490E05EFFB7FF57D17BFF48FAF7" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[164,245,1376,1402]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="B902D301E05EFFB7FF57D17BFF48FAF7" box="[164,245,1376,1402]" italics="true" pageId="66" pageNumber="67">bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE05EFFB7FF0FD17BFE8CFAF7" box="[252,305,1376,1402]" pageId="66" pageNumber="67" rank="species">sp.n.</taxonomicNameLabel>
| det.:
<determinerName id="7E15E3D7E05EFFB7FE8FD17BFDADFAF7" box="[380,528,1376,1402]" pageId="66" pageNumber="67">H. Dombrow</determinerName>
2019» (
<collectionCode id="ED6797D6E05EFFB7FD9FD17BFD7FFAF7" box="[620,706,1376,1402]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="66" pageNumber="67" type="Museum">SAMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BC90F13E05EFFB7FF64D1B3FD3FFA6B" blockId="66.[151,1437,1448,1510]" pageId="66" pageNumber="67">
<emphasis id="B902D301E05EFFB7FF64D1B3FE18FA4F" bold="true" box="[151,421,1448,1474]" pageId="66" pageNumber="67">
Condition of
<typeStatus id="54CDB1B1E05EFFB7FEC4D1B3FE1DFA4F" box="[311,416,1448,1474]" pageId="66" pageNumber="67" type="holotype">holotype</typeStatus>
.
</emphasis>
Good. Glued on a card. Dissected, parameres and spiculum gastrale glued behind beetle. Left mesothoracic tibia with tarsus missing.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E05EFFB7FF64D20FFC66F9DF" pageId="66" pageNumber="67" type="discussion">
<paragraph id="8BC90F13E05EFFB7FF64D20FFC66F9DF" blockId="66.[151,1436,1556,1619]" pageId="66" pageNumber="67">
<emphasis id="B902D301E05EFFB7FF64D20FFEBDF9A3" bold="true" box="[151,256,1556,1582]" pageId="66" pageNumber="67">Remark.</emphasis>
The locality description requires an adjustment, as the GPS coordinate on the locality label actually indicates a locality ~
<quantity id="4C8EA2F6E05EFFB7FEA7D223FE26F9DF" box="[340,411,1592,1618]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="66" pageNumber="67" unit="km" value="10.0">10 km</quantity>
NE Nieuwoudtville on the R357 to Loeriesfontein.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E05EFFB7FF64D29BFBD2F8FF" pageId="66" pageNumber="67" type="diagnosis">
<paragraph id="8BC90F13E05EFFB7FF64D29BFBD2F8FF" blockId="66.[151,1437,1664,1906]" pageId="66" pageNumber="67">
<emphasis id="B902D301E05EFFB7FF64D29BFE20F917" bold="true" box="[151,413,1664,1690]" pageId="66" pageNumber="67">Differential diagnosis.</emphasis>
Differs from
<taxonomicName id="4C767490E05EFFB7FDC5D29AFCEDF917" authority=": Elytra" authorityName="Elytra" box="[566,848,1664,1690]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dolichiocnemis">
<emphasis id="B902D301E05EFFB7FDC5D29AFD42F917" box="[566,767,1664,1690]" italics="true" pageId="66" pageNumber="67">B. dolichiocnemis</emphasis>
: Elytra
</taxonomicName>
of
<taxonomicName id="4C767490E05EFFB7FC86D29AFC58F917" authorityName="Dombrow" authorityYear="2020" box="[885,997,1664,1690]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B902D301E05EFFB7FC86D29AFC58F917" box="[885,997,1664,1690]" italics="true" pageId="66" pageNumber="67">B. bicolor</emphasis>
</taxonomicName>
partly covering
<taxonomicName id="4C767490E05EFFB7FB6FD29BFA8DF917" authorityName="Bocchino" authorityYear="1964" box="[1180,1328,1664,1690]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="66" pageNumber="70" phylum="Chordata" rank="genus">propygidium</taxonomicName>
. Elytra of
<taxonomicName id="4C767490E05EFFB7FF64D2BEFEDFF933" box="[151,354,1700,1726]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dolichiocnemis">
<emphasis id="B902D301E05EFFB7FF64D2BEFEDFF933" box="[151,354,1700,1726]" italics="true" pageId="66" pageNumber="67">B. dolichiocnemis</emphasis>
</taxonomicName>
protruding
<taxonomicName id="4C767490E05EFFB7FE1AD2BFFDC0F933" authorityName="Bocchino" authorityYear="1964" box="[489,637,1700,1726]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="66" pageNumber="70" phylum="Chordata" rank="genus">propygidium</taxonomicName>
. Elytra of
<taxonomicName id="4C767490E05EFFB7FD05D2BEFCDAF933" authorityName="Dombrow" authorityYear="2020" box="[758,871,1700,1726]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B902D301E05EFFB7FD05D2BEFCDAF933" box="[758,871,1700,1726]" italics="true" pageId="66" pageNumber="67">B. bicolor</emphasis>
</taxonomicName>
transparently pale-brown with broadly blackened lateral margins. Elytra of
<taxonomicName id="4C767490E05EFFB7FE44D2D2FD3DF96F" box="[439,640,1736,1762]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dolichiocnemis">
<emphasis id="B902D301E05EFFB7FE44D2D2FD3DF96F" box="[439,640,1736,1762]" italics="true" pageId="66" pageNumber="67">B. dolichiocnemis</emphasis>
</taxonomicName>
transparently pale red brown with narrowly blackened lateral margins. Interno-posterior margin of parameres of
<taxonomicName id="4C767490E05EFFB7FD9BD2F6FD64F88B" authorityName="Dombrow" authorityYear="2020" box="[616,729,1772,1798]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B902D301E05EFFB7FD9BD2F6FD64F88B" box="[616,729,1772,1798]" italics="true" pageId="66" pageNumber="67">B. bicolor</emphasis>
</taxonomicName>
u-shaped in dorsal view (
<figureCitation id="134D1396E05EFFB7FC09D2F7FBEAF88B" box="[1018,1111,1772,1798]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="66" pageNumber="67">Figs. 94</figureCitation>
,
<figureCitation id="134D1396E05EFFB7FB90D2F7FB2DF88A" box="[1123,1168,1772,1799]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="66" pageNumber="67">116</figureCitation>
). Interno-posterior margin of parameres of
<taxonomicName id="4C767490E05EFFB7FE89D30AFDFEF8A7" box="[378,579,1808,1834]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dolichiocnemis">
<emphasis id="B902D301E05EFFB7FE89D30AFDFEF8A7" box="[378,579,1808,1834]" italics="true" pageId="66" pageNumber="67">B. dolichiocnemis</emphasis>
</taxonomicName>
parabola-shaped in dorsal view (
<figureCitation id="134D1396E05EFFB7FC4BD30BFBA8F8A7" box="[952,1045,1808,1834]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="66" pageNumber="67">Figs. 96</figureCitation>
,
<figureCitation id="134D1396E05EFFB7FBD2D30BFBF3F8A7" box="[1057,1102,1808,1834]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="66" pageNumber="67">117</figureCitation>
). Externo-posterior margin of parameres of
<taxonomicName id="4C767490E05EFFB7FEDDD32EFE20F8C3" authorityName="Dombrow" authorityYear="2020" box="[302,413,1844,1870]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B902D301E05EFFB7FEDDD32EFE20F8C3" box="[302,413,1844,1870]" italics="true" pageId="66" pageNumber="67">B. bicolor</emphasis>
</taxonomicName>
concave with a broad convex median extension in dorsal view (
<figureCitation id="134D1396E05EFFB7FB90D32FFB02F8C3" box="[1123,1215,1844,1870]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="66" pageNumber="67">Figs. 94</figureCitation>
,
<figureCitation id="134D1396E05EFFB7FB38D32FFB45F8C2" box="[1227,1272,1844,1871]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="66" pageNumber="67">116</figureCitation>
). Externo-posterior margin of parameres of
<taxonomicName id="4C767490E05EFFB7FE14D342FD0DF8FF" box="[487,688,1880,1906]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="dolichiocnemis">
<emphasis id="B902D301E05EFFB7FE14D342FD0DF8FF" box="[487,688,1880,1906]" italics="true" pageId="66" pageNumber="67">B. dolichiocnemis</emphasis>
</taxonomicName>
undulate in dorsal view (
<figureCitation id="134D1396E05EFFB7FC3FD343FB97F8FF" box="[972,1066,1880,1906]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="66" pageNumber="67">Figs. 96</figureCitation>
,
<figureCitation id="134D1396E05EFFB7FBC5D343FBDEF8FF" box="[1078,1123,1880,1906]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="66" pageNumber="67">117</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E05FFFB1FF64D48DFED7FCEB" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68" type="description">
<paragraph id="8BC90F13E05FFFB6FF64D48DFE95FF3D" blockId="67.[151,1437,150,2015]" box="[151,296,150,176]" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF64D48DFE95FF3D" bold="true" box="[151,296,150,176]" pageId="67" pageNumber="68">Description</emphasis>
</paragraph>
<paragraph id="8BC90F13E05FFFB6FF34D4A2FD44FF59" blockId="67.[151,1437,150,2015]" box="[199,761,185,212]" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF34D4A2FE35FF5E" bold="true" box="[199,392,185,211]" pageId="67" pageNumber="68">Measurements:</emphasis>
length
<quantity id="4C8EA2F6E05FFFB6FE11D4A2FDFDFF59" box="[482,576,185,212]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="67" pageNumber="68" unit="mm" value="8.3">8.3 mm</quantity>
; width
<quantity id="4C8EA2F6E05FFFB6FD64D4A2FD49FF59" box="[663,756,185,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="67" pageNumber="68" unit="mm" value="3.8">3.8 mm</quantity>
.
</paragraph>
<paragraph id="8BC90F13E05FFFB6FF34D4C7FEAAFE94" blockId="67.[151,1437,150,2015]" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF34D4C7FEDFFF7B" bold="true" box="[199,354,220,246]" pageId="67" pageNumber="68">Integument:</emphasis>
Black; elytra transparently yellowish pale-brown, lateral and posterior margin broadly blackened.
</paragraph>
<paragraph id="8BC90F13E05FFFB6FF34D53AFA9FFA1C" blockId="67.[151,1437,150,2015]" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF34D53AFEF0FEB6" bold="true" box="[199,333,289,315]" pageId="67" pageNumber="68">Tegument:</emphasis>
<emphasis id="B902D301E05FFFB6FEABD53AFE0BFEB6" box="[344,438,289,315]" italics="true" pageId="67" pageNumber="68">Clypeus</emphasis>
and lateral margin bare; inner side of reflexed anterior margin sparse setose with short, erect, black setae.
<emphasis id="B902D301E05FFFB6FE30D55FFDBFFED3" box="[451,514,324,350]" italics="true" pageId="67" pageNumber="68">Head</emphasis>
loose setose with medium-long, erect, black setae.
<emphasis id="B902D301E05FFFB6FB82D55EFABCFED3" box="[1137,1281,324,350]" italics="true" pageId="67" pageNumber="68">Eye canthus</emphasis>
with a loose setose ciliary of long, black setae.
<taxonomicName id="4C767490E05FFFB6FDC8D57CFD33FE0C" authorityName="Norman" authorityYear="1875" box="[571,654,359,385]" genus="Scapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="68" phylum="Cnidaria" rank="genus">
<emphasis id="B902D301E05FFFB6FDC8D57CFD33FE0C" box="[571,654,359,385]" italics="true" pageId="67" pageNumber="68">Scapus</emphasis>
</taxonomicName>
loose setose with long, black setae, pale-brown ventrally, sparse and medium-long on
<emphasis id="B902D301E05FFFB6FE6FD592FD23FE2E" box="[412,670,393,419]" italics="true" pageId="67" pageNumber="68">pedicellus; funiculum</emphasis>
with few very short, erect, brown and black setae, shorter and semi-erect on
<emphasis id="B902D301E05FFFB6FEB3D5B7FE36FE4B" box="[320,395,428,454]" italics="true" pageId="67" pageNumber="68">clavus</emphasis>
.
<emphasis id="B902D301E05FFFB6FE6FD5B6FDADFE4B" box="[412,528,429,454]" italics="true" pageId="67" pageNumber="68">Pronotum</emphasis>
loose setose with long, erect, black setae, shorter towards posterior margin; with loose setose ciliary of distinctly long, black setae on lateral margin.
<taxonomicName id="4C767490E05FFFB6FBFED5D4FB3CFE64" box="[1037,1153,463,489]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E05FFFB6FBFED5D4FB3CFE64" box="[1037,1153,463,489]" italics="true" pageId="67" pageNumber="68">Scutellum</emphasis>
</taxonomicName>
loose setose with short and medium-long, semi-erect, brown setae.
<emphasis id="B902D301E05FFFB6FD6CD5EAFD54FD86" box="[671,745,497,523]" italics="true" pageId="67" pageNumber="68">Elytra</emphasis>
without pattern. Loose setose with short, reclinate, brown setae; medium-long, erect, black setae interspersed; few short, reclinate, whitish setae below apical torus; with two loose, longitudinal rows of long, erect, black setae: a juxta-sutural row from anterior margin crossing scutellar torus towards apical torus, shorter in posterior third; a juxta-humeral row from anterior margin between humerus and scutellar torus towards apical torus. Suture with a loose row of short and long, robust, semi-erect, black setae.
<emphasis id="B902D301E05FFFB6FE48D684FDB2FD34" box="[443,527,671,697]" italics="true" pageId="67" pageNumber="68">Thorax</emphasis>
and
<emphasis id="B902D301E05FFFB6FDBDD684FD0BFD34" box="[590,694,671,697]" italics="true" pageId="67" pageNumber="68">pleurites</emphasis>
loose pilose with long, black hairs.
<emphasis id="B902D301E05FFFB6FB81D684FABFFD35" box="[1138,1282,671,696]" italics="true" pageId="67" pageNumber="68">Ventrite IV</emphasis>
loose setose with short, reclinate, white and medium-long, black setae; long, semi-erect, black setae interspersed.
<taxonomicName id="4C767490E05FFFB6FA9BD6D9FEB9FD73" authorityName="Bocchino" authorityYear="1964" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="67" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="B902D301E05FFFB6FA9BD6D9FEB9FD73" italics="true" pageId="67" pageNumber="68">Propygidium</emphasis>
</taxonomicName>
loose squamose with reclinate, narrowly-lanceolate, greyish scales; long, semi-erect, black setae interspersed.
<taxonomicName id="4C767490E05FFFB6FEC6D71CFE18FCAC" box="[309,421,775,801]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="67" pageNumber="68" phylum="Chordata" rank="genus">
<emphasis id="B902D301E05FFFB6FEC6D71CFE18FCAC" box="[309,421,775,801]" italics="true" pageId="67" pageNumber="68">Pygidium</emphasis>
</taxonomicName>
sparse setose with long, erect, black setae; tip with a loose setose ciliary of long, black setae.
<emphasis id="B902D301E05FFFB6FF10D732FE02FCCE" box="[227,447,809,835]" italics="true" pageId="67" pageNumber="68">Prothoracic femur</emphasis>
loose pilose with long, black hairs; bare internally.
<emphasis id="B902D301E05FFFB6FBC5D732FABEFCCE" box="[1078,1283,809,835]" italics="true" pageId="67" pageNumber="68">Prothoracic tibia</emphasis>
loose setose with short, reclinate, greyish setae; medium-long, semi-erect, black setae interspersed; long ventrally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; with a loose ciliary of mediumlong, black and whitish setae on internal ridge; a sparse ciliary of short pale-brown setae on external ridge; a setaceous ciliary of short, black setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth.
<emphasis id="B902D301E05FFFB6FB08D7CCFF66FB9E" italics="true" pageId="67" pageNumber="68">Mesothoracic femur</emphasis>
loose pilose with long, black hairs; distinctly long ventrally; bare internally.
<emphasis id="B902D301E05FFFB6FB98D7E2FAF6FB9E" box="[1131,1355,1017,1043]" italics="true" pageId="67" pageNumber="68">Mesothoracic tibia</emphasis>
sparse setose with short, reclinate, white setae; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of short, black bristles; lengthening dorsally; missing internally.
<emphasis id="B902D301E05FFFB6FEE4D07AFDBEFBF6" box="[279,515,1121,1147]" italics="true" pageId="67" pageNumber="68">Metathoracic femur</emphasis>
loose setose with long, semi-erect, black setae; bare internally.
<emphasis id="B902D301E05FFFB6FB0DD07AFF70FB13" italics="true" pageId="67" pageNumber="68">Metathoracic tibia</emphasis>
sparse setose with short, reclinate, brown and black setae; medium-long, semi-erect, robust, black setae interspersed; long and fine internally. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally, more robust ventrally.
<emphasis id="B902D301E05FFFB6FBF1D0D2FAE4FB6E" box="[1026,1369,1225,1251]" italics="true" pageId="67" pageNumber="68">Prothoracic tarsomeres IIV</emphasis>
bare.
<emphasis id="B902D301E05FFFB6FF64D0F7FE84FA8B" box="[151,313,1260,1286]" italics="true" pageId="67" pageNumber="68">Mesothoracic</emphasis>
and
<emphasis id="B902D301E05FFFB6FE84D0F7FD66FA8B" box="[375,731,1260,1286]" italics="true" pageId="67" pageNumber="68">metathoracic tarsomeres IIV</emphasis>
sparse setose with very short, reclinate, pale-brown setae.
<emphasis id="B902D301E05FFFB6FF64D114FEF0FAA5" box="[151,333,1295,1320]" italics="true" pageId="67" pageNumber="68">Tarsomere IIV</emphasis>
of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae, lengthening towards
<emphasis id="B902D301E05FFFB6FF0ED12AFD99FAC6" box="[253,548,1329,1355]" italics="true" pageId="67" pageNumber="68">metathoracic tarsomeres</emphasis>
, fine dorsally and more robust ventrally.
<emphasis id="B902D301E05FFFB6FBE6D129FB08FAC6" box="[1045,1205,1330,1355]" italics="true" pageId="67" pageNumber="68">Pretarsomere</emphasis>
of all thoracic legs with few, medium-long, semi-erect, black setae at apical margin, shorter and dark-brown ventrally.
<emphasis id="B902D301E05FFFB6FADDD14FFF6EFA1D" italics="true" pageId="67" pageNumber="68">Mesothoracic</emphasis>
and
<emphasis id="B902D301E05FFFB6FEFDD16CFDF3FA1D" box="[270,590,1399,1425]" italics="true" pageId="67" pageNumber="68">metathoracic pretarsomere</emphasis>
additionally sparse setose with short, reclinate, brown setae.
</paragraph>
<paragraph id="8BC90F13E05FFFB6FF34D182FD18F996" blockId="67.[151,1437,150,2015]" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF34D182FEAEFA3E" bold="true" box="[199,275,1433,1459]" pageId="67" pageNumber="68">Head:</emphasis>
Suture between head and clypeus undulate, crenated. Head dense punctate with small, shallow, roundish, partly merging punctures, micropunctate within. Funicle I cylindrical, slightly thickened medially; funicle II cylindrical; funicle III short asymmetric frustum-shaped; clavus long elliptic, slightly shorter than
<taxonomicName id="4C767490E05FFFB6FF64D219FF54F991" authorityName="Norman" authorityYear="1875" box="[151,233,1538,1564]" genus="Scapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="67" pageNumber="70" phylum="Cnidaria" rank="genus">scapus</taxonomicName>
, pedicellus and funiculum combined.
</paragraph>
<paragraph id="8BC90F13E05FFFB6FF34D23FFDE4F92B" blockId="67.[151,1437,150,2015]" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF34D23FFEF2F9B3" bold="true" box="[199,335,1572,1598]" pageId="67" pageNumber="68">Pronotum:</emphasis>
Widest in anterior two fifth; loose punctate with shallow, very small and small, roundish punctures, dense laterally and posteriorly and within posterior half of medial furrow. Anterior corner obtuse. Lateral margin convex. With an indistinct, shortened, shallow medial furrow from posterior margin terminating on disk. Posterior corner rounded.
</paragraph>
<paragraph id="8BC90F13E05FFFB6FF34D2B5FDBEF8BC" blockId="67.[151,1437,150,2015]" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF34D2B5FEF3F945" bold="true" box="[199,334,1710,1736]" pageId="67" pageNumber="68">
<taxonomicName id="4C767490E05FFFB6FF34D2B5FEF4F945" box="[199,329,1710,1736]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="67" pageNumber="70" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
:
</emphasis>
Buckler-shaped; as broad as long; dense punctate with shallow, very small and small, roundish punctures, partly merging, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior fifth. Lateral margin linearly attenuated in anterior quarter, stronger apically. Tip rounded.
</paragraph>
<paragraph id="8BC90F13E05FFFB1FF34D322FE45FF59" blockId="67.[151,1437,150,2015]" lastBlockId="68.[151,1437,151,872]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
<emphasis id="B902D301E05FFFB6FF34D322FE9CF8DE" bold="true" box="[199,289,1849,1875]" pageId="67" pageNumber="68">Elytra:</emphasis>
Ovate; longer than broad; widest in anterior third; partly covering
<taxonomicName id="4C767490E05FFFB6FBB6D322FB5FF8DE" authorityName="Bocchino" authorityYear="1964" box="[1093,1250,1849,1875]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="67" pageNumber="70" phylum="Chordata" rank="genus">propygidium</taxonomicName>
. Lateral margin slightly convex attenuate apically. Sculpture indistinct; almost complanate laterally; humerus indistinct; with a broad roundish torus beside
<taxonomicName id="4C767490E05FFFB6FDE2D364FD35F814" box="[529,648,1919,1945]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="67" pageNumber="70" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
; lateral trench missing; complanate along
<taxonomicName id="4C767490E05FFFB6FB64D364FAB7F814" box="[1175,1290,1919,1945]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="67" pageNumber="70" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
and suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; with two loose, longitudinal rows of small, deep, roundish punctures: a juxta-sutural row from anterior margin crossing scutellar torus to apical torus; a supra-humeral row from anterior margin between humerus and scutellar torus to apical torus.
</paragraph>
<paragraph id="8BC90F13E058FFB1FF34D4C7FA26FEB6" blockId="68.[151,1437,151,872]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF34D4C7FEF7FF7B" bold="true" box="[199,330,220,246]" pageId="68" pageNumber="69">Abdomen:</emphasis>
Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally.
<taxonomicName id="4C767490E058FFB1FABED4C6FF56FE94" authorityName="Bocchino" authorityYear="1964" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="68" pageNumber="70" phylum="Chordata" rank="genus">Propygidium</taxonomicName>
with a slightly projected stoma at posterior corner.
<taxonomicName id="4C767490E058FFB1FCBBD4E4FC06FE94" box="[840,955,255,281]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="68" pageNumber="70" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
slightly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin linearly attenuate apically; tip rounded.
</paragraph>
<paragraph id="8BC90F13E058FFB1FF34D55FFE17FD86" blockId="68.[151,1437,151,872]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF34D55FFE2BFED3" bold="true" box="[199,406,324,350]" pageId="68" pageNumber="69">Prothoracic leg:</emphasis>
Femur and tibia slender. Tibia smooth ventrally. Median tooth robust. Basal tooth rudimentary. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere IIV combined extremely short; equal in length; slightly more than
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in
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length of tibia. Tarsomere IV with a single ventral extension. Pretarsomere distinctly thickened; smooth ventrally; slightly more than
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<subScript id="17F20D56E058FFB1FAEED5A5FA93FE41" attach="left" box="[1309,1326,446,460]" fontSize="6" pageId="68" pageNumber="69">10</subScript>
in
</quantity>
length of tarsomere IIV combined. Internal claw as long as pretarsomere. External claw slightly less than
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<quantity id="4C8EA2F6E058FFB1FA98D5FAFA21FE64" box="[1387,1436,463,495]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="68" pageNumber="69" unit="in" value="10.0">
<subScript id="17F20D56E058FFB1FA98D5FAFAC1FE62" attach="left" box="[1387,1404,481,495]" fontSize="6" pageId="68" pageNumber="69">10</subScript>
in
</quantity>
length of internal claw.
</paragraph>
<paragraph id="8BC90F13E058FFB1FF34D60FFB57FD1B" blockId="68.[151,1437,151,872]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF34D60FFE15FDA3" bold="true" box="[199,424,532,558]" pageId="68" pageNumber="69">Mesothoracic leg:</emphasis>
Tibia slightly more than
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<subScript id="17F20D56E058FFB1FD18D63DFD4EFDB9" attach="left" box="[747,755,550,564]" fontSize="6" pageId="68" pageNumber="69">5</subScript>
in
</quantity>
length of femur. Tarsomere IIV combined extremely short; slightly less than
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<quantity id="4C8EA2F6E058FFB1FE37D652FE56FDDC" box="[452,491,567,599]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="68" pageNumber="69" unit="in" value="5.0">
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in
</quantity>
length of tibia. Pretarsomere with a shortened smooth ridge ventrally, followed by a roundish cutting apically; slightly less than
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more in length of tarsomere IIV combined. External claw slightly less than
<superScript id="7C03A25BE058FFB1FE54D667FE12FD07" attach="right" box="[423,431,636,650]" fontSize="6" pageId="68" pageNumber="69">9</superScript>
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<quantity id="4C8EA2F6E058FFB1FE4BD695FE55FD1B" box="[440,488,636,668]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="68" pageNumber="69" unit="in" value="10.0">
<subScript id="17F20D56E058FFB1FE4BD695FE74FD11" attach="left" box="[440,457,654,668]" fontSize="6" pageId="68" pageNumber="69">10</subScript>
in
</quantity>
length of pretarsomere. Internal claw shorter than external claw.
</paragraph>
<paragraph id="8BC90F13E058FFB1FF34D685FC4BFCAC" blockId="68.[151,1437,151,872]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF34D685FE1BFD34" bold="true" box="[199,422,670,697]" pageId="68" pageNumber="69">Metathoracic leg:</emphasis>
Tibia slightly longer than femur. Tarsomere IIV combined distinctly short; slightly more than
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<subScript id="17F20D56E058FFB1FED9D6CFFE86FD6F" attach="left" box="[298,315,724,738]" fontSize="6" pageId="68" pageNumber="69">10</subScript>
in
</quantity>
length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than
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<quantity id="4C8EA2F6E058FFB1FEE4D6EDFE83FD73" box="[279,318,740,772]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="68" pageNumber="69" unit="in" value="2.0">
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in
</quantity>
length of tarsomere IIV combined. External claw slightly less than
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more in length of pretarsomere. Internal claw slightly more than
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<quantity id="4C8EA2F6E058FFB1FD43D702FD6AFCAC" box="[688,727,775,807]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="68" pageNumber="69" unit="in" value="5.0">
<subScript id="17F20D56E058FFB1FD43D702FD05FCAA" attach="left" box="[688,696,793,807]" fontSize="6" pageId="68" pageNumber="69">5</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E058FFB1FF34D732FDFCFCC9" blockId="68.[151,1437,151,872]" box="[199,577,809,836]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF34D732FEEAFCCE" bold="true" box="[199,343,809,835]" pageId="68" pageNumber="69">Parameres:</emphasis>
(
<figureCitation id="134D1396E058FFB1FE9AD732FE44FCC9" box="[361,505,809,836]" captionStart="FIGURES 6297" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 6297. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 6263, B. nigra Dombrow, new species; 6465, B. pallidibrunnea Dombrow, new species; 6667, B. fusca Dombrow, new species; 6869, B. setosa Dombrow, new species; 7071, B. castanea Dombrow, new species; 7273, B. colvillei Dombrow, 2005; 7475, B. pulchra Dombrow, new species; 7677, B. nigrofasciata Dombrow, new species; 7879, B. caliginosa Dombrow, new species; 8081, B. pumila Dombrow, new species; 8283, B. gifbergensis Dombrow, new species; 8485, B. julianae Dombrow, 2005; 8687, B. elkeae Dombrow, new species; 8889, B. suturalis Dombrow, new species; 9091, B. nigrosetosa Dombrow, new species; 9293, B. occidentalis Dombrow, 2005; 9495, B. bicolor Dombrow, new species; 9697, B. dolichiocnemis Dombrow, new species; 9899, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="68" pageNumber="69">Figs. 9495</figureCitation>
,
<figureCitation id="134D1396E058FFB1FDF6D732FD89FCC9" box="[517,564,809,836]" captionStart="FIGURES 100117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="68" pageNumber="69">116</figureCitation>
).
</paragraph>
<paragraph id="8BC90F13E058FFB1FF34D756FED7FCEB" blockId="68.[151,1437,151,872]" box="[199,362,844,872]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF34D756FF5BFCE5" bold="true" box="[199,230,845,872]" pageId="68" pageNumber="69">
<specimenCount id="9D70C49AE058FFB1FF34D756FF62FCE5" box="[199,223,845,872]" pageId="68" pageNumber="69" type="female"></specimenCount>
:
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E058FFB1FF64D78AFC88FC7C" pageId="68" pageNumber="69" type="distribution">
<paragraph id="8BC90F13E058FFB1FF64D78AFB37FC26" blockId="68.[151,1162,913,940]" box="[151,1162,913,940]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF64D78AFE84FC26" bold="true" box="[151,313,913,939]" pageId="68" pageNumber="69">Distribution.</emphasis>
<collectingRegion id="49B2C1F1E058FFB1FEB2D78AFDDCFC26" box="[321,609,913,939]" country="South Africa" name="Northern Cape" pageId="68" pageNumber="69">Northern Cape Province</collectingRegion>
(
<figureCitation id="134D1396E058FFB1FD82D78AFD64FC26" box="[625,729,913,939]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="68" pageNumber="69">Fig. 140</figureCitation>
), only known from the
<typeStatus id="54CDB1B1E058FFB1FC03D789FB9EFC21" box="[1008,1059,914,940]" pageId="68" pageNumber="69">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BC90F13E058FFB1FF64D7CDFC88FC7C" blockId="68.[151,821,982,1009]" box="[151,821,982,1009]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF64D7CDFED5FC7C" bold="true" box="[151,360,982,1009]" pageId="68" pageNumber="69">
Vegetation
<typeStatus id="54CDB1B1E058FFB1FED7D7CCFEDEFC7C" box="[292,355,983,1009]" pageId="68" pageNumber="69">Type</typeStatus>
.
</emphasis>
Bokkeveld Sandstone Fynbos [FFs 1].
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E058FFB1FF64D007FCD6FBBB" box="[151,875,1052,1079]" pageId="68" pageNumber="69" type="biology_ecology">
<paragraph id="8BC90F13E058FFB1FF64D007FCD6FBBB" blockId="68.[151,875,1052,1079]" box="[151,875,1052,1079]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF64D007FE8DFBBB" bold="true" box="[151,304,1052,1078]" pageId="68" pageNumber="69">Host Plants.</emphasis>
<taxonomicName id="4C767490E058FFB1FECBD007FE02FBBB" box="[312,447,1052,1078]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="68" pageNumber="69" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
:
<taxonomicName id="4C767490E058FFB1FE39D007FD38FBBB" authorityName="Aiton" authorityYear="1789" box="[458,645,1052,1078]" class="Adenophorea" family="Linhomoeidae" genus="Didelta" kingdom="Animalia" order="Monhysterida" pageId="68" pageNumber="69" phylum="Nematoda" rank="species" species="spinosa">
<emphasis id="B902D301E058FFB1FE39D007FD38FBBB" box="[458,645,1052,1078]" italics="true" pageId="68" pageNumber="69">Didelta spinosa</emphasis>
</taxonomicName>
(L. f.) W. T. Aiton.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E058FFB1FF64D07AFAA7FBF1" box="[151,1306,1121,1148]" pageId="68" pageNumber="69" type="etymology">
<paragraph id="8BC90F13E058FFB1FF64D07AFAA7FBF1" blockId="68.[151,1306,1121,1148]" box="[151,1306,1121,1148]" pageId="68" pageNumber="69">
<emphasis id="B902D301E058FFB1FF64D07AFE99FBF6" bold="true" box="[151,292,1121,1147]" pageId="68" pageNumber="69">Etymology.</emphasis>
The name derives from the bicoloured elytra [Latin: bi = two; Latin: color = colour].
</paragraph>
</subSubSection>
</treatment>
</document>