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<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" approvalRequired="186" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="100" approvalRequired_for_treatments="80" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy &amp; Divljan, Anja" docDate="2017" docId="03DD87C8FF86730A189EFD80FBA494DB" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Thylogale eugenii subsp. decres Troughton 1941" docType="treatment" docVersion="3" lastPageNumber="340" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="340" updateTime="1674153641407" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parnaby, Harry E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ingleby, Sandy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Divljan, Anja</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-10-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>5</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>277</mods:start>
<mods:end>420</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3853/j.2201-4349.69.2017.1653</mods:identifier>
<mods:identifier type="ISSN">2201-4349</mods:identifier>
<mods:identifier type="Zenodo-Dep">5237800</mods:identifier>
<mods:identifier type="ZooBank">68F315FF-3FEB-410E-96EC-5F494510F440</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5237980" ID-Zenodo-Dep="5237980" LSID="urn:lsid:plazi:treatment:03DD87C8FF86730A189EFD80FBA494DB" httpUri="http://treatment.plazi.org/id/03DD87C8FF86730A189EFD80FBA494DB" lastPageNumber="340" pageId="63" pageNumber="340">
<subSubSection pageId="63" pageNumber="340" type="nomenclature">
<paragraph blockId="63.[172,698,619,646]" box="[172,698,619,646]" pageId="63" pageNumber="340">
<heading box="[172,698,619,646]" centered="true" fontSize="11" level="2" pageId="63" pageNumber="340" reason="2">
<taxonomicName authority="Troughton, 1941" authorityName="Troughton" authorityYear="1941" box="[172,698,619,646]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="63" pageNumber="340" phylum="Chordata" rank="subSpecies" species="eugenii" subSpecies="decres">
<emphasis bold="true" box="[172,698,619,646]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[172,480,619,646]" italics="true" pageId="63" pageNumber="340">Thylogale eugenii decres</emphasis>
<bibRefCitation author="Troughton, E" box="[487,698,619,645]" pageId="63" pageNumber="340" refId="ref147799" refString="Troughton, E. [L. G.] 1941. Furred Animals of Australia. First edition. Sydney: Angus &amp; Robertson. xxvii + 374 pp." type="book" year="1941">Troughton, 1941</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="63.[151,718,665,714]" pageId="63" pageNumber="340">
<emphasis box="[151,347,665,686]" italics="true" pageId="63" pageNumber="340">Furred Animals of</emphasis>
<materialsCitation collectingDate="1941-11-20" country="Australia" location="Australia" pageId="63" pageNumber="340" specimenCount="1">
<emphasis box="[357,453,665,686]" italics="true" pageId="63" pageNumber="340">
<collectingCountry box="[357,453,665,686]" name="Australia" pageId="63" pageNumber="340">Australia</collectingCountry>
</emphasis>
, 1st edition, p. 194. (
<date pageId="63" pageNumber="340" value="1941-11-20">
<collectingDate pageId="63" pageNumber="340" value="1941-11-20">20 November 1941</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="340" type="description">
<paragraph blockId="63.[121,497,735,760]" box="[121,497,735,760]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[121,295,735,760]" pageId="63" pageNumber="340">Common name</emphasis>
. Tammar Wallaby.
</paragraph>
<paragraph blockId="63.[119,750,781,1039]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[121,281,781,805]" pageId="63" pageNumber="340">Current name</emphasis>
.
<taxonomicName authority="eugenii (Desmarest, 1817 a)" authorityName="eugenii (Desmarest" authorityYear="1817" baseAuthorityName="Desmarest" baseAuthorityYear="1817" class="Mammalia" family="Macropodidae" genus="Notamacropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="63" pageNumber="340" phylum="Chordata" rank="subSpecies" species="eugenii" subSpecies="eugenii">
<emphasis box="[293,615,782,805]" italics="true" pageId="63" pageNumber="340">Notamacropus eugenii eugenii</emphasis>
(
<bibRefCitation author="Desmarest, A" pageId="63" pageNumber="340" pagination="28 - 44" refId="ref127066" refString="Desmarest, A. G. 1817 a. Kanguroo. In Nouveau Dictionnaire d'Histoire naturelle, appliquee aux arts, a l'agriculture, a l'economie rurale et domestique, a la medecine, etc. Par une societe de natualistes et d'agriculteurs. Nouvelle Edition presqu'entierement refondue et considerablement augmentee. Volume 17, ed. C. S. Sonnini, pp. 28 - 44. Paris: Deterville." type="book chapter" year="1817">Desmarest, 1817a</bibRefCitation>
)
</taxonomicName>
, following Eldridge &amp; Coulson (2015). Jackson &amp; Groves (2015) elevated
<taxonomicName box="[379,535,840,863]" class="Mammalia" family="Macropodidae" genus="Notamacropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="63" pageNumber="340" phylum="Chordata" rank="genus">
<emphasis box="[379,535,840,863]" italics="true" pageId="63" pageNumber="340">Notamacropus</emphasis>
</taxonomicName>
to generic rank and highlighted the need for further taxonomic resolution of the status of names included within
<taxonomicName baseAuthorityName="Desmarest" baseAuthorityYear="1817" box="[471,581,899,922]" class="Mammalia" family="Macropodidae" genus="Notamacropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="63" pageNumber="340" phylum="Chordata" rank="species" species="eugenii">
<emphasis box="[471,581,899,922]" italics="true" pageId="63" pageNumber="340">N. eugenii</emphasis>
</taxonomicName>
.
<bibRefCitation author="Eldridge, M &amp; Miller, L &amp; Neaves, K" pageId="63" pageNumber="340" pagination="0172777" refId="ref128002" refString="Eldridge, M. D. B., E. J. Miller, L. E. Neaves, K. R. Zenger, and C. A. Herbert 2017. Extensive genetic differentiation detected within a model marsupial, the tammar wallaby (Notamacropus eugenii). PLoS ONE 12 (3): e 0172777. https: // doi. org / 10.1371 / journal. pone. 0172777" type="journal article" year="2017">
Eldridge
<emphasis box="[695,750,898,922]" italics="true" pageId="63" pageNumber="340">et al.</emphasis>
(2017)
</bibRefCitation>
demonstrated that the Kangaroo Island population was genetically distinct, but its taxonomic status remains unresolved as they were unable to include definitive samples from extinct South Australian mainland populations.
</paragraph>
<paragraph blockId="63.[119,751,1061,1349]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[121,223,1061,1085]" pageId="63" pageNumber="340">
<typeStatus box="[121,223,1061,1085]" pageId="63" pageNumber="340">Syntypes</typeStatus>
</emphasis>
. (5, by subsequent determination):
<emphasis bold="true" box="[591,677,1061,1085]" pageId="63" pageNumber="340">M.2821</emphasis>
, male, skull, study skin, collected
<date box="[416,609,1090,1114]" pageId="63" pageNumber="340" value="1920-02-13">13 February 1920</date>
<materialsCitation collectingDate="1920-07-31" collectorName="E. Le G. Troughton" county="Deep Creek" location="Kangaroo Island" municipality="Kingscote" pageId="63" pageNumber="340" specimenCount="1" stateProvince="South Australia" typeStatus="syntype">
by
<collectorName pageId="63" pageNumber="340">E. Le G. Troughton</collectorName>
,
<collectingCounty box="[242,369,1119,1143]" pageId="63" pageNumber="340">Deep Creek</collectingCounty>
,
<quantity box="[377,465,1119,1144]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="63" pageNumber="340" unit="mi" value="20.0">20 miles</quantity>
from
<collectingMunicipality box="[526,635,1119,1143]" pageId="63" pageNumber="340">Kingscote</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF86730A189EFD80FBA494DB:8EAB6005FF86730A1AB6FB8BFF6E93A9" county="Deep Creek" municipality="Kingscote" name="Kangaroo Island" pageId="63" pageNumber="340" stateProvince="South Australia">Kangaroo Island</location>
,
<collectingRegion box="[200,370,1149,1173]" country="Australia" name="South Australia" pageId="63" pageNumber="340">South Australia</collectingRegion>
, registered
<date box="[496,642,1149,1173]" pageId="63" pageNumber="340" value="1920-07-31">
<collectingDate box="[496,642,1149,1173]" pageId="63" pageNumber="340" value="1920-07-31">31 July 1920</collectingDate>
</date>
</materialsCitation>
;
<emphasis bold="true" box="[659,742,1149,1173]" pageId="63" pageNumber="340">M.2820</emphasis>
, male, subadult, skull, study skin, collected
<date box="[569,744,1178,1202]" pageId="63" pageNumber="340" value="1920-02-08">8 February 1920</date>
,
<materialsCitation collectingDate="1933-03-16" county="Taronga Zoo" location="Kangaroo Island" municipality="Sydney" pageId="63" pageNumber="340" specimenCount="1" stateProvince="South Australia" typeStatus="syntype">
other details as per M.2821;
<emphasis bold="true" box="[436,519,1207,1231]" pageId="63" pageNumber="340">M.5332</emphasis>
, female, adult, skull, study skin, (ex
<collectingCounty box="[276,410,1237,1261]" pageId="63" pageNumber="340">Taronga Zoo</collectingCounty>
,
<collectingMunicipality box="[419,501,1237,1261]" pageId="63" pageNumber="340">Sydney</collectingMunicipality>
), from
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF86730A189EFD80FBA494DB:8EAB6005FF86730A1A0EFB3EFD3E93D1" box="[572,747,1237,1261]" county="Taronga Zoo" municipality="Sydney" name="Kangaroo Island" pageId="63" pageNumber="340" stateProvince="South Australia">Kangaroo Island</location>
, registered
<date box="[233,403,1266,1290]" pageId="63" pageNumber="340" value="1933-03-16">
<collectingDate box="[233,403,1266,1290]" pageId="63" pageNumber="340" value="1933-03-16">16 March 1933</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="1928-06-18" collectorName="A. S. Le Souef." location="Kangaroo Island" pageId="63" pageNumber="340" specimenCount="1" stateProvince="South Australia" typeStatus="syntype">
<emphasis bold="true" box="[419,494,1266,1290]" pageId="63" pageNumber="340">S.1808</emphasis>
and
<emphasis bold="true" box="[550,625,1266,1290]" pageId="63" pageNumber="340">S.1809</emphasis>
, both male skulls, collection dates not given,
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF86730A189EFD80FBA494DB:8EAB6005FF86730A19E2FAFBFDAB921B" box="[464,638,1295,1320]" name="Kangaroo Island" pageId="63" pageNumber="340" stateProvince="South Australia">Kangaroo Island</location>
, registered
<date box="[119,261,1325,1349]" pageId="63" pageNumber="340" value="1928-06-18">
<collectingDate box="[119,261,1325,1349]" pageId="63" pageNumber="340" value="1928-06-18">18 June 1928</collectingDate>
</date>
collected by
<collectorName box="[403,568,1325,1349]" pageId="63" pageNumber="340">A. S. Le Souef.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="63.[119,749,1370,1658]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[121,235,1370,1394]" pageId="63" pageNumber="340">Condition</emphasis>
.
<emphasis bold="true" box="[252,337,1370,1394]" pageId="63" pageNumber="340">M.2821</emphasis>
: Cranium missing last erupted upper right molar and right alisphenoid bone, hole in left maxilla in the orbit area; right dentary missing coronoid and condylar processes. Study skin complete, in good condition.
<emphasis bold="true" box="[654,739,1458,1482]" pageId="63" pageNumber="340">M.2820</emphasis>
: Cranium and dentaries complete, in good condition; study skin missing tail tip and scrotum.
<emphasis bold="true" box="[503,588,1517,1541]" pageId="63" pageNumber="340">M.5332</emphasis>
: Cranium and dentaries complete. Study skin in good condition, slither of skin missing from the right ear.
<emphasis bold="true" box="[443,516,1575,1599]" pageId="63" pageNumber="340">S.1808</emphasis>
: Cranium missing left 3rd upper incisor; both dentaries complete.
<emphasis bold="true" box="[568,642,1605,1629]" pageId="63" pageNumber="340">S.1809</emphasis>
: Cranium and both dentaries complete.
</paragraph>
<paragraph blockId="63.[119,751,1679,2000]" lastBlockId="63.[792,1422,151,442]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[121,447,1679,1704]" pageId="63" pageNumber="340">Cranial measurements (mm)</emphasis>
.
<emphasis bold="true" box="[460,545,1679,1703]" pageId="63" pageNumber="340">M.2821</emphasis>
: GL, 90.64; ConL, 84.87; BasL, 79.59; NasL, 31.12; NasB, 14.85; DIL, 17.82; APV, 3.95; PPV, —; PAL, —; UPM (alv.), 4.42; UMR (alv.), 16.76 (M
<superScript attach="left" box="[221,229,1766,1780]" fontSize="6" pageId="63" pageNumber="340">1</superScript>
to M
<superScript attach="left" box="[288,296,1766,1780]" fontSize="6" pageId="63" pageNumber="340">3</superScript>
); ZB, 49.07; POC, 17.10; MB, 37.19; DL (condyl.), 59.58 (left dentary); LPM (alv.), 2.82; LMR (alv.), 16.57 (M
<subScript attach="left" box="[220,228,1839,1853]" fontSize="6" pageId="63" pageNumber="340">1</subScript>
to M
<subScript attach="left" box="[286,294,1839,1853]" fontSize="6" pageId="63" pageNumber="340">3</subScript>
).
<emphasis bold="true" box="[316,401,1826,1850]" pageId="63" pageNumber="340">M.5332</emphasis>
: GL, 97.83; ConL, 92.45; BasL, 87.86; NasL, 31.46; NasB, 17.07; DIL, 23.89; APV, 4.84; PPV, 12.69; PAL, 55.59; UPM (alv.), 4.13; UMR (alv.), 23.12 (M
<superScript attach="left" box="[221,229,1913,1927]" fontSize="6" pageId="63" pageNumber="340">1</superScript>
to M
<superScript attach="left" box="[288,296,1913,1927]" fontSize="6" pageId="63" pageNumber="340">4</superScript>
); ZB, 51.20; POC, 16.22; MB, 40.71; DL (condyl.), 70.23; LPM (alv.), 3.61; LMR (alv.), 22.41 (M
<subScript attach="left" box="[714,722,1956,1970]" fontSize="6" pageId="63" pageNumber="340">1</subScript>
to M
<subScript attach="right" box="[145,153,1986,2000]" fontSize="6" pageId="63" pageNumber="340">4</subScript>
).
<emphasis bold="true" box="[174,249,1973,1997]" pageId="63" pageNumber="340">S.1808</emphasis>
: GL, 101.79; ConL, 96.42; BasL, 90.36; NasL, 33.68; NasB, 18.63; DIL, 22.85; APV, 4.09; PPV, 11.86; PAL, 57.02; UPM (alv.), 4.64; UMR (alv.), 17.61 (M
<superScript attach="left" box="[1335,1343,179,193]" fontSize="6" pageId="63" pageNumber="340">1</superScript>
to M
<superScript attach="left" box="[1397,1405,179,193]" fontSize="6" pageId="63" pageNumber="340">3</superScript>
); ZB, 54.24; POC, 16.34; MB, 40.21; DL (condyl.), 71.51; LPM (alv.), 3.88; LMR (alv.), 16.55 (M
<subScript attach="left" box="[1234,1242,252,266]" fontSize="6" pageId="63" pageNumber="340">1</subScript>
to M
<subScript attach="left" box="[1306,1314,252,266]" fontSize="6" pageId="63" pageNumber="340">3</subScript>
).
<emphasis bold="true" box="[1339,1414,239,263]" pageId="63" pageNumber="340">S.1809</emphasis>
: GL, 101.95; ConL, 96.88; BasL, 92.54; NasL, 36.62* (tips worn); NasB, 17.51; DIL, 21.74; APV, 5.45; PPV, 10.51; PAL, 56.95* (posterior margin slightly eroded); UPM (alv.), 5.17; UMR (alv.), 17.08 (M
<superScript attach="left" box="[1082,1090,355,369]" fontSize="6" pageId="63" pageNumber="340">1</superScript>
to M
<superScript attach="left" box="[1144,1152,355,369]" fontSize="6" pageId="63" pageNumber="340">3</superScript>
); ZB, 53.16; POC, 18.20; MB, 42.06; DL (condyl.), 71.05; LPM (alv.), 4.12; LMR (alv.), 16.60 (M
<subScript attach="left" box="[962,970,428,442]" fontSize="6" pageId="63" pageNumber="340">1</subScript>
to M
<subScript attach="left" box="[1028,1036,428,442]" fontSize="6" pageId="63" pageNumber="340">3</subScript>
). [* = estimate].
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="340" type="materials_examined">
<paragraph blockId="63.[794,1421,461,544]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[794,963,461,485]" pageId="63" pageNumber="340">Type localities</emphasis>
.
<materialsCitation county="Deep Creek" location="Kangaroo Island" municipality="Kingscote" pageId="63" pageNumber="340" specimenCount="1" stateProvince="South Australia" typeStatus="syntype">
<collectingCounty box="[981,1117,461,485]" pageId="63" pageNumber="340">Deep Creek</collectingCounty>
,
<quantity box="[1132,1229,461,485]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="63" pageNumber="340" unit="mi" value="20.0">20 miles</quantity>
from
<collectingMunicipality box="[1303,1417,461,485]" pageId="63" pageNumber="340">Kingscote</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF86730A189EFD80FBA494DB:8EAB6005FF86730A1B28FE00FC1B953E" box="[794,974,490,515]" county="Deep Creek" municipality="Kingscote" name="Kangaroo Island" pageId="63" pageNumber="340" stateProvince="South Australia">Kangaroo Island</location>
,
<collectingRegion box="[984,1153,490,514]" country="Australia" name="South Australia" pageId="63" pageNumber="340">South Australia</collectingRegion>
.
<location LSID="urn:lsid:plazi:treatment:03DD87C8FF86730A189EFD80FBA494DB:8EAB6005FF86730A1CB9FE01FB1D953E" box="[1163,1224,490,514]" county="Deep Creek" municipality="Kingscote" name="Other" pageId="63" pageNumber="340" stateProvince="South Australia">Other</location>
<typeStatus box="[1230,1323,491,515]" pageId="63" pageNumber="340">syntypes</typeStatus>
are from “Kangaroo Island”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="340" type="discussion">
<paragraph blockId="63.[792,1422,565,999]" pageId="63" pageNumber="340">
<emphasis bold="true" box="[794,915,565,589]" pageId="63" pageNumber="340">Comments</emphasis>
.
<bibRefCitation author="Troughton, E" box="[926,1111,565,589]" pageId="63" pageNumber="340" refId="ref147799" refString="Troughton, E. [L. G.] 1941. Furred Animals of Australia. First edition. Sydney: Angus &amp; Robertson. xxvii + 374 pp." type="book" year="1941">Troughton (1941)</bibRefCitation>
stated that the subspecies was based on “specimens in the Australian Museum, including a male and female collected by the author in 1920” but does not cite registration or field numbers nor indicate the number of specimens in the type series. Calaby &amp; Richardson (1988) listed registration numbers of the
<specimenCount box="[1133,1286,711,735]" pageId="63" pageNumber="340" type="generic">five specimens</specimenCount>
listed above, which they recognize as
<typeStatus box="[1039,1132,741,765]" pageId="63" pageNumber="340">syntypes</typeStatus>
. Troughton did not nominate a
<typeStatus box="[808,899,770,794]" pageId="63" pageNumber="340">holotype</typeStatus>
in his description, but the specimen tag of M.2821 is labelled
<typeStatus box="[916,1013,799,823]" pageId="63" pageNumber="340">holotype</typeStatus>
, M.5332 as
<typeStatus box="[1156,1243,799,823]" pageId="63" pageNumber="340">allotype</typeStatus>
and M.2820 as
<typeStatus box="[794,887,829,853]" pageId="63" pageNumber="340">paratype</typeStatus>
, all in Troughtons characteristic handwriting; they are listed here as
<typeStatus box="[979,1070,859,883]" pageId="63" pageNumber="340">syntypes</typeStatus>
because Troughtons attributions remain unpublished. Specimen index cards for S.18089 are not marked as
<typeStatus box="[946,1046,917,941]" pageId="63" pageNumber="340">paratypes</typeStatus>
but they are recognized as
<typeStatus box="[1329,1421,917,941]" pageId="63" pageNumber="340">syntypes</typeStatus>
because they were available to Troughton in 1941. Publication date drawn from
<bibRefCitation author="Whitley, G" box="[970,1133,975,999]" pageId="63" pageNumber="340" pagination="219 - 238" refId="ref149696" refString="Whitley, G. P. 1975. Memorial notice and bibliography. Ellis Le Geyt Troughton. Australian Zoologist 18 (3): 219 - 238." type="journal article" year="1975">Whitley (1975)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>