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<document ID-DOI="http://doi.org/10.5281/zenodo.4752929" ID-GBIF-Dataset="f0f8f265-10f1-448f-8b15-6c56fc140166" ID-GBIF-Taxon="182067787" ID-ISSN="1854-0392" ID-Zenodo-Dep="4752929" ID-ZooBank="8BFBE3F7-E0EF-4EDD-BA82-9EA087F74688" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1620834297829" checkinUser="felipe" docAuthor="Grubbs, Scott A." docDate="2015" docId="03DD8784FF99A71AFCB55F0F65E9FE67" docLanguage="en" docName="Illiesia.11.12.147-166.pdf" docOrigin="Illiesia 11 (12)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Leuctra tenuis" docType="key" docVersion="5" lastPageId="2" lastPageNumber="149" masterDocId="FFE4FFFCFF98A718FFF65E23604AFFEB" masterDocTitle="Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa" masterLastPageNumber="166" masterPageNumber="147" pageId="1" pageNumber="148" updateTime="1632012907386" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa</mods:title>
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Key to males of the
<taxonomicName baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[1076,1177,300,324]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis bold="true" box="[1076,1177,300,324]" italics="true" pageId="1" pageNumber="148">L. tenuis</emphasis>
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species group
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1 Dorsal process on 7
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abdominal tergum tapering to a single, narrowly-rounded posterior lobe, overall shape triangular (
<figureCitation captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="1" pageNumber="148">Figs. 10</figureCitation>
11, 2325) ..……………….………...........…… 2
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1 Dorsal process on 7
<superScript attach="left" box="[1163,1178,516,530]" fontSize="6" pageId="1" pageNumber="148">th</superScript>
abdominal tergum narrowing little posteriorly, overall shape variable (
<figureCitation box="[984,1088,588,612]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="1" pageNumber="148">Figs. 13</figureCitation>
,
<figureCitation box="[1099,1208,588,612]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs 49" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStart-6="Figs" captionStart-7="Figs" captionStartId-0="8.[189,241,1152,1176]" captionStartId-1="2.[189,241,1373,1397]" captionStartId-2="3.[189,238,1552,1576]" captionStartId-3="4.[189,241,1588,1612]" captionStartId-4="6.[189,241,1153,1177]" captionStartId-5="10.[189,241,1584,1608]" captionStartId-6="11.[189,241,1592,1616]" captionStartId-7="13.[189,241,1706,1730]" captionTargetBox-0="[249,1383,254,1105]" captionTargetBox-1="[249,1383,470,1327]" captionTargetBox-2="[249,1383,218,1505]" captionTargetBox-3="[249,1383,254,1542]" captionTargetBox-4="[249,1383,254,1107]" captionTargetBox-5="[249,1383,254,1538]" captionTargetBox-6="[249,1383,254,1546]" captionTargetBox-7="[249,1383,362,1658]" captionTargetId-0="figure-432@8.[249,1383,254,1105]" captionTargetId-1="figure-268@2.[249,1383,470,1327]" captionTargetId-2="figure-210@3.[249,1383,218,1505]" captionTargetId-3="figure-193@4.[249,1383,254,1542]" captionTargetId-4="figure-422@6.[249,1383,254,1107]" captionTargetId-5="figure-197@10.[249,1383,254,1538]" captionTargetId-6="figure-151@11.[249,1383,254,1546]" captionTargetId-7="figure-160@13.[249,1383,362,1659]" captionTargetPageId-0="8" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="6" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="13" captionText-0="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." captionText-1="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-2="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." captionText-3="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." captionText-4="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." captionText-5="Figs. 2327. Leuctra tenuis. 23, 7th abdominal process, dorsal view, 350X, Maryland, Middle Fork; 24, 7th abdominal process, dorsal view, 350X, Maryland, Youghioghenny River; 25, 7th abdominal process, dorsal view, 350X, Massachusetts, Stillwater River; 26, paraprocts, caudal view, 500X, Indiana, tributary to West Fork Mosquito Creek; 27, paraprocts, caudal view, 1000X, Indiana, tributary to West Fork Mosquito Creek. sp = specillum, sa = subanal lobe." captionText-6="Figs. 2833. Leuctra tenuis. 28, paraprocts, lateral view, 500X, Alabama, Hillabee Creek; 29, paraprocts, lateral view, 500X, Massachusetts, Stillwater River; 30, paraprocts, lateral view, 500X, Maryland, Middle Fork; 31, paraprocts, lateral view, 2000X, Maryland, Middle Fork; 32, paraprocts, lateral view, 350X, Tennessee, Cosby Creek; 33, paraprocts, lateral view, 2000X, Tennessee, Cosby Creek. sp = specillum, sa = subanal lobe." captionText-7="Figs. 3438. Leuctra triloba. 34, 7th abdominal process, dorsal view, 350X, Alabama, tributary to West Fork Hatchett Creek; 35, 7th abdominal process, dorsal view, 350X, North Carolina, Wine Spring Creek; 36, L. triloba, 7th abdominal process, dorsal view, 500X, Tennessee, tributary to South Suck Creek; 37, paraprocts, caudal view, 750X, North Carolina, Wine Spring Creek; 38, L. triloba, paraprocts, caudal view, 1500X, North Carolina, Wine Spring Creek. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752780" figureDoi-1="http://doi.org/10.5281/zenodo.4752770" figureDoi-2="http://doi.org/10.5281/zenodo.4752774" figureDoi-3="http://doi.org/10.5281/zenodo.4752776" figureDoi-4="http://doi.org/10.5281/zenodo.4752778" figureDoi-5="http://doi.org/10.5281/zenodo.4752784" figureDoi-6="http://doi.org/10.5281/zenodo.4752786" figureDoi-7="http://doi.org/10.5281/zenodo.4752788" httpUri-0="https://zenodo.org/record/4752780/files/figure.png" httpUri-1="https://zenodo.org/record/4752770/files/figure.png" httpUri-2="https://zenodo.org/record/4752774/files/figure.png" httpUri-3="https://zenodo.org/record/4752776/files/figure.png" httpUri-4="https://zenodo.org/record/4752778/files/figure.png" httpUri-5="https://zenodo.org/record/4752784/files/figure.png" httpUri-6="https://zenodo.org/record/4752786/files/figure.png" httpUri-7="https://zenodo.org/record/4752788/files/figure.png" pageId="1" pageNumber="148">19, 3436</figureCitation>
,
<figureCitation box="[1219,1248,588,612]" captionStart="Figs" captionStartId="15.[189,241,1594,1618]" captionTargetBox="[249,1383,254,1548]" captionTargetId="figure-163@15.[249,1383,254,1548]" captionTargetPageId="15" captionText="Figs. 4347. Leuctra usdi, Tennessee, spring into Pinewood Branch. 43, 7th abdominal process, dorsal view, 350X; 44, paraprocts, caudal view, 500X; 45, paraprocts, caudal view, 2000X; 46, paraprocts, lateral view, 350X; 47, L. usdi, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi="http://doi.org/10.5281/zenodo.4752792" httpUri="https://zenodo.org/record/4752792/files/figure.png" pageId="1" pageNumber="148">43</figureCitation>
,
<figureCitation box="[1259,1286,588,612]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="1" pageNumber="148">48</figureCitation>
).……….… 3
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2 Subanal lobe ca. ¾ length of the specillum (
<figureCitation captionStart="Figs" captionStartId="11.[189,241,1592,1616]" captionTargetBox="[249,1383,254,1546]" captionTargetId="figure-151@11.[249,1383,254,1546]" captionTargetPageId="11" captionText="Figs. 2833. Leuctra tenuis. 28, paraprocts, lateral view, 500X, Alabama, Hillabee Creek; 29, paraprocts, lateral view, 500X, Massachusetts, Stillwater River; 30, paraprocts, lateral view, 500X, Maryland, Middle Fork; 31, paraprocts, lateral view, 2000X, Maryland, Middle Fork; 32, paraprocts, lateral view, 350X, Tennessee, Cosby Creek; 33, paraprocts, lateral view, 2000X, Tennessee, Cosby Creek. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752786" httpUri="https://zenodo.org/record/4752786/files/figure.png" pageId="1" pageNumber="148">Figs. 2830, 32</figureCitation>
); specillum simple, lacking spurs or tubercles distally (
<figureCitation box="[1160,1284,696,720]" captionStart="Figs" captionStartId="10.[189,241,1584,1608]" captionTargetBox="[249,1383,254,1538]" captionTargetId="figure-197@10.[249,1383,254,1538]" captionTargetPageId="10" captionText="Figs. 2327. Leuctra tenuis. 23, 7th abdominal process, dorsal view, 350X, Maryland, Middle Fork; 24, 7th abdominal process, dorsal view, 350X, Maryland, Youghioghenny River; 25, 7th abdominal process, dorsal view, 350X, Massachusetts, Stillwater River; 26, paraprocts, caudal view, 500X, Indiana, tributary to West Fork Mosquito Creek; 27, paraprocts, caudal view, 1000X, Indiana, tributary to West Fork Mosquito Creek. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752784" httpUri="https://zenodo.org/record/4752784/files/figure.png" pageId="1" pageNumber="148">Figs. 27</figureCitation>
,
<figureCitation box="[1333,1434,696,720]" captionStart="Figs" captionStartId="11.[189,241,1592,1616]" captionTargetBox="[249,1383,254,1546]" captionTargetId="figure-151@11.[249,1383,254,1546]" captionTargetPageId="11" captionText="Figs. 2833. Leuctra tenuis. 28, paraprocts, lateral view, 500X, Alabama, Hillabee Creek; 29, paraprocts, lateral view, 500X, Massachusetts, Stillwater River; 30, paraprocts, lateral view, 500X, Maryland, Middle Fork; 31, paraprocts, lateral view, 2000X, Maryland, Middle Fork; 32, paraprocts, lateral view, 350X, Tennessee, Cosby Creek; 33, paraprocts, lateral view, 2000X, Tennessee, Cosby Creek. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752786" httpUri="https://zenodo.org/record/4752786/files/figure.png" pageId="1" pageNumber="148">31, 33</figureCitation>
) ...…………………………...….…
<taxonomicName authority="(Pictet)" baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[1246,1439,732,756]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis bold="true" box="[1246,1348,732,756]" italics="true" pageId="1" pageNumber="148">L. tenuis</emphasis>
(Pictet)
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2 Subanal lobe ca. equal length to specillum (
<figureCitation box="[882,1027,804,828]" captionStart-0="Figs" captionStart-1="Figs 49" captionStart-2="Figs" captionStartId-0="2.[189,241,1373,1397]" captionStartId-1="3.[189,238,1552,1576]" captionStartId-2="4.[189,241,1588,1612]" captionTargetBox-0="[249,1383,470,1327]" captionTargetBox-1="[249,1383,218,1505]" captionTargetBox-2="[249,1383,254,1542]" captionTargetId-0="figure-268@2.[249,1383,470,1327]" captionTargetId-1="figure-210@3.[249,1383,218,1505]" captionTargetId-2="figure-193@4.[249,1383,254,1542]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-1="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." captionText-2="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752770" figureDoi-1="http://doi.org/10.5281/zenodo.4752774" figureDoi-2="http://doi.org/10.5281/zenodo.4752776" httpUri-0="https://zenodo.org/record/4752770/files/figure.png" httpUri-1="https://zenodo.org/record/4752774/files/figure.png" httpUri-2="https://zenodo.org/record/4752776/files/figure.png" pageId="1" pageNumber="148">Figs. 1415</figureCitation>
); specillum with a prominent, laterally-directed spur (
<figureCitation box="[1255,1433,840,864]" captionStart="Figs" captionStartId="4.[189,241,1588,1612]" captionTargetBox="[249,1383,254,1542]" captionTargetId="figure-193@4.[249,1383,254,1542]" captionTargetPageId="4" captionText="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752776" httpUri="https://zenodo.org/record/4752776/files/figure.png" pageId="1" pageNumber="148">Figs. 1213</figureCitation>
) ………………………...
<emphasis bold="true" box="[1139,1347,875,900]" pageId="1" pageNumber="148">
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[1139,1275,875,899]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis bold="true" box="[1139,1275,875,899]" italics="true" pageId="1" pageNumber="148">L. schusteri</emphasis>
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<taxonomicNameLabel box="[1283,1347,876,900]" pageId="1" pageNumber="148" rank="species">sp. n.</taxonomicNameLabel>
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Grubbs
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3 Posterior margin of 7
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abdominal dorsal process terminating along the same plane, overall shape symmetrically-trapezoidal (
<figureCitation captionStart="Figs" captionStartId="15.[189,241,1594,1618]" captionTargetBox="[249,1383,254,1548]" captionTargetId="figure-163@15.[249,1383,254,1548]" captionTargetPageId="15" captionText="Figs. 4347. Leuctra usdi, Tennessee, spring into Pinewood Branch. 43, 7th abdominal process, dorsal view, 350X; 44, paraprocts, caudal view, 500X; 45, paraprocts, caudal view, 2000X; 46, paraprocts, lateral view, 350X; 47, L. usdi, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi="http://doi.org/10.5281/zenodo.4752792" httpUri="https://zenodo.org/record/4752792/files/figure.png" pageId="1" pageNumber="148">Figs. 43</figureCitation>
,
<figureCitation box="[913,939,1020,1044]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="1" pageNumber="148">48</figureCitation>
) .......................................…………………... 4
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3 Dorsal process on 7
<superScript attach="left" box="[1163,1178,1056,1070]" fontSize="6" pageId="1" pageNumber="148">th</superScript>
abdominal tergum variable, overall shape variable (
<figureCitation box="[1282,1392,1092,1116]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="1" pageNumber="148">Figs. 13</figureCitation>
,
<figureCitation captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs 49" captionStart-3="Figs" captionStart-4="Figs" captionStart-5="Figs" captionStart-6="Figs" captionStart-7="Figs" captionStartId-0="8.[189,241,1152,1176]" captionStartId-1="2.[189,241,1373,1397]" captionStartId-2="3.[189,238,1552,1576]" captionStartId-3="4.[189,241,1588,1612]" captionStartId-4="6.[189,241,1153,1177]" captionStartId-5="10.[189,241,1584,1608]" captionStartId-6="11.[189,241,1592,1616]" captionStartId-7="13.[189,241,1706,1730]" captionTargetBox-0="[249,1383,254,1105]" captionTargetBox-1="[249,1383,470,1327]" captionTargetBox-2="[249,1383,218,1505]" captionTargetBox-3="[249,1383,254,1542]" captionTargetBox-4="[249,1383,254,1107]" captionTargetBox-5="[249,1383,254,1538]" captionTargetBox-6="[249,1383,254,1546]" captionTargetBox-7="[249,1383,362,1658]" captionTargetId-0="figure-432@8.[249,1383,254,1105]" captionTargetId-1="figure-268@2.[249,1383,470,1327]" captionTargetId-2="figure-210@3.[249,1383,218,1505]" captionTargetId-3="figure-193@4.[249,1383,254,1542]" captionTargetId-4="figure-422@6.[249,1383,254,1107]" captionTargetId-5="figure-197@10.[249,1383,254,1538]" captionTargetId-6="figure-151@11.[249,1383,254,1546]" captionTargetId-7="figure-160@13.[249,1383,362,1659]" captionTargetPageId-0="8" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="6" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="13" captionText-0="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." captionText-1="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-2="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." captionText-3="Figs. 1015. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1011, 7th abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe." captionText-4="Figs. 1618. Leuctra schusteri sp. n., Kentucky, spring into Smyrna Creek. 1617, subgenital plate, ventral view, 350X; 18, vesicle, ventral view, 650X." captionText-5="Figs. 2327. Leuctra tenuis. 23, 7th abdominal process, dorsal view, 350X, Maryland, Middle Fork; 24, 7th abdominal process, dorsal view, 350X, Maryland, Youghioghenny River; 25, 7th abdominal process, dorsal view, 350X, Massachusetts, Stillwater River; 26, paraprocts, caudal view, 500X, Indiana, tributary to West Fork Mosquito Creek; 27, paraprocts, caudal view, 1000X, Indiana, tributary to West Fork Mosquito Creek. sp = specillum, sa = subanal lobe." captionText-6="Figs. 2833. Leuctra tenuis. 28, paraprocts, lateral view, 500X, Alabama, Hillabee Creek; 29, paraprocts, lateral view, 500X, Massachusetts, Stillwater River; 30, paraprocts, lateral view, 500X, Maryland, Middle Fork; 31, paraprocts, lateral view, 2000X, Maryland, Middle Fork; 32, paraprocts, lateral view, 350X, Tennessee, Cosby Creek; 33, paraprocts, lateral view, 2000X, Tennessee, Cosby Creek. sp = specillum, sa = subanal lobe." captionText-7="Figs. 3438. Leuctra triloba. 34, 7th abdominal process, dorsal view, 350X, Alabama, tributary to West Fork Hatchett Creek; 35, 7th abdominal process, dorsal view, 350X, North Carolina, Wine Spring Creek; 36, L. triloba, 7th abdominal process, dorsal view, 500X, Tennessee, tributary to South Suck Creek; 37, paraprocts, caudal view, 750X, North Carolina, Wine Spring Creek; 38, L. triloba, paraprocts, caudal view, 1500X, North Carolina, Wine Spring Creek. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752780" figureDoi-1="http://doi.org/10.5281/zenodo.4752770" figureDoi-2="http://doi.org/10.5281/zenodo.4752774" figureDoi-3="http://doi.org/10.5281/zenodo.4752776" figureDoi-4="http://doi.org/10.5281/zenodo.4752778" figureDoi-5="http://doi.org/10.5281/zenodo.4752784" figureDoi-6="http://doi.org/10.5281/zenodo.4752786" figureDoi-7="http://doi.org/10.5281/zenodo.4752788" httpUri-0="https://zenodo.org/record/4752780/files/figure.png" httpUri-1="https://zenodo.org/record/4752770/files/figure.png" httpUri-2="https://zenodo.org/record/4752774/files/figure.png" httpUri-3="https://zenodo.org/record/4752776/files/figure.png" httpUri-4="https://zenodo.org/record/4752778/files/figure.png" httpUri-5="https://zenodo.org/record/4752784/files/figure.png" httpUri-6="https://zenodo.org/record/4752786/files/figure.png" httpUri-7="https://zenodo.org/record/4752788/files/figure.png" pageId="1" pageNumber="148">19, 3436</figureCitation>
) ..…………………………………………… 5
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<paragraph blockId="1.[835,1444,372,1908]" pageId="1" pageNumber="148">
<keyLead pageId="1" pageNumber="148">
4 7th abdominal dorsal process bearing three subtruncate lobes, lateral lobes small, medial lobe ca. 4X wider than lateral lobes (
<figureCitation box="[1343,1430,1235,1260]" captionStart="Figs" captionStartId="15.[189,241,1594,1618]" captionTargetBox="[249,1383,254,1548]" captionTargetId="figure-163@15.[249,1383,254,1548]" captionTargetPageId="15" captionText="Figs. 4347. Leuctra usdi, Tennessee, spring into Pinewood Branch. 43, 7th abdominal process, dorsal view, 350X; 44, paraprocts, caudal view, 500X; 45, paraprocts, caudal view, 2000X; 46, paraprocts, lateral view, 350X; 47, L. usdi, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi="http://doi.org/10.5281/zenodo.4752792" httpUri="https://zenodo.org/record/4752792/files/figure.png" pageId="1" pageNumber="148">Fig. 43</figureCitation>
); specillum bearing tubercles distally and along the outer distal margin (
<figureCitation box="[1161,1292,1307,1332]" captionStart-0="Figs" captionStart-1="Figs 49" captionStartId-0="15.[189,241,1594,1618]" captionStartId-1="3.[189,238,1552,1576]" captionTargetBox-0="[249,1383,254,1548]" captionTargetBox-1="[249,1383,218,1505]" captionTargetId-0="figure-163@15.[249,1383,254,1548]" captionTargetId-1="figure-210@3.[249,1383,218,1505]" captionTargetPageId-0="15" captionTargetPageId-1="3" captionText-0="Figs. 4347. Leuctra usdi, Tennessee, spring into Pinewood Branch. 43, 7th abdominal process, dorsal view, 350X; 44, paraprocts, caudal view, 500X; 45, paraprocts, caudal view, 2000X; 46, paraprocts, lateral view, 350X; 47, L. usdi, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." captionText-1="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752792" figureDoi-1="http://doi.org/10.5281/zenodo.4752774" httpUri-0="https://zenodo.org/record/4752792/files/figure.png" httpUri-1="https://zenodo.org/record/4752774/files/figure.png" pageId="1" pageNumber="148">Figs. 4447</figureCitation>
); late winter and spring-emergent species ......
<taxonomicName authority="Grubbs" authorityName="Grubbs" authorityYear="2010" box="[1265,1439,1343,1368]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="usdi">
<emphasis bold="true" box="[1265,1345,1343,1367]" italics="true" pageId="1" pageNumber="148">L. usdi</emphasis>
Grubbs
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</paragraph>
<paragraph blockId="1.[835,1444,372,1908]" pageId="1" pageNumber="148">
<keyLead pageId="1" pageNumber="148">
4 7th abdominal dorsal process lacking distinct lateral lobes, essentially straight along posterior margin (
<figureCitation box="[1106,1193,1451,1476]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="1" pageNumber="148">Fig. 48</figureCitation>
); specillum lacking tubercles along distal outer margin and at the tips (
<figureCitation box="[945,1088,1523,1548]" captionStart="Figs" captionStartId="16.[189,241,1664,1688]" captionTargetBox="[249,1383,290,1581]" captionTargetId="figure-100@16.[249,1383,290,1581]" captionTargetPageId="16" captionText="Figs. 4852. Leuctra variabilis, Tennessee, Twin Springs. 48, 7th abdominal process, dorsal view, 350X; 49, paraprocts, caudal view, 350X; 50, paraprocts, caudal view, 1500X; 51, paraprocts, lateral view, 350X; 52, L. variabilis, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752794" httpUri="https://zenodo.org/record/4752794/files/figure.png" pageId="1" pageNumber="148">Figs. 4952</figureCitation>
); autumn-emergent species ……………………………...
<taxonomicName authority="Hanson" authorityName="Hanson" authorityYear="1941" box="[1193,1438,1559,1583]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis bold="true" box="[1193,1339,1559,1583]" italics="true" pageId="1" pageNumber="148">L. variabilis</emphasis>
Hanson
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="1" pageNumber="148">
<paragraph blockId="1.[835,1444,372,1908]" pageId="1" pageNumber="148">
<keyLead pageId="1" pageNumber="148">
5 7th abdominal dorsal process bearing three rounded lobes, medial lobe narrowly-rounded posteriorly and nipple-like (
<figureCitation box="[1275,1428,1667,1691]" captionStart="Figs" captionStartId="13.[189,241,1706,1730]" captionTargetBox="[249,1383,362,1658]" captionTargetId="figure-160@13.[249,1383,362,1659]" captionTargetPageId="13" captionText="Figs. 3438. Leuctra triloba. 34, 7th abdominal process, dorsal view, 350X, Alabama, tributary to West Fork Hatchett Creek; 35, 7th abdominal process, dorsal view, 350X, North Carolina, Wine Spring Creek; 36, L. triloba, 7th abdominal process, dorsal view, 500X, Tennessee, tributary to South Suck Creek; 37, paraprocts, caudal view, 750X, North Carolina, Wine Spring Creek; 38, L. triloba, paraprocts, caudal view, 1500X, North Carolina, Wine Spring Creek. sp = specillum, sa = subanal lobe, ll = lateral lobe, ml = medial lobe." figureDoi="http://doi.org/10.5281/zenodo.4752788" httpUri="https://zenodo.org/record/4752788/files/figure.png" pageId="1" pageNumber="148">Figs. 3436</figureCitation>
); autumn- and winter-emergent species.……………………………..…
<taxonomicName authority="Claassen" authorityName=": Needham and Claassen" authorityYear="1925" box="[1219,1442,1739,1763]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="triloba">
<emphasis bold="true" box="[1219,1331,1739,1763]" italics="true" pageId="1" pageNumber="148">L. triloba</emphasis>
Claassen
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="1.[835,1444,372,1908]" pageId="1" pageNumber="148">
<keyLead pageId="1" pageNumber="148">
5 7th abdominal dorsal process lacking distinct lobes, medial posterior portion either broadlyrounded or subtriangular (
<figureCitation box="[1213,1323,1847,1871]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="1" pageNumber="148">Figs. 13</figureCitation>
,
<figureCitation box="[1344,1371,1847,1871]" captionStart="Figs" captionStartId="8.[189,241,1152,1176]" captionTargetBox="[249,1383,254,1105]" captionTargetId="figure-432@8.[249,1383,254,1105]" captionTargetPageId="8" captionText="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752780" httpUri="https://zenodo.org/record/4752780/files/figure.png" pageId="1" pageNumber="148">19</figureCitation>
); late spring- and summer-emergent species ….....… 6
</keyLead>
</paragraph>
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<paragraph blockId="2.[189,798,228,396]" pageId="2" pageNumber="149">
<keyLead pageId="2" pageNumber="149">
6 7
<superScript attach="left" box="[240,255,228,242]" fontSize="6" pageId="2" pageNumber="149">th</superScript>
abdominal dorsal lobe broadly rounded posteriorly (
<figureCitation box="[377,483,264,288]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="2" pageNumber="149">Figs. 13</figureCitation>
); specillum subtriangular distally (
<figureCitation box="[336,484,300,324]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="2" pageNumber="149">Figs. 4, 7, 9</figureCitation>
), subanal lobes recurved anteriorly, but not at tip (
<figureCitation box="[608,788,336,360]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="2" pageNumber="149">Figs. 4, 6, 9</figureCitation>
) …………………...……..
<taxonomicName authority="Claassen" authorityName=": Needham and Claassen" authorityYear="1925" box="[507,791,372,396]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis bold="true" box="[507,680,372,396]" italics="true" pageId="2" pageNumber="149">L. carolinensis</emphasis>
Claassen
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<paragraph blockId="2.[835,1444,228,396]" pageId="2" pageNumber="149">
<keyLead pageId="2" pageNumber="149">
6 7
<superScript attach="left" box="[886,901,228,242]" fontSize="6" pageId="2" pageNumber="149">th</superScript>
abdominal dorsal process broadly subtriangular posteriorly (
<figureCitation box="[1207,1296,264,288]" captionStart="Figs" captionStartId="8.[189,241,1152,1176]" captionTargetBox="[249,1383,254,1105]" captionTargetId="figure-432@8.[249,1383,254,1105]" captionTargetPageId="8" captionText="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752780" httpUri="https://zenodo.org/record/4752780/files/figure.png" pageId="2" pageNumber="149">Fig. 19</figureCitation>
); specillum truncate distally (
<figureCitation box="[1082,1163,300,324]" captionStart="Figs" captionStartId="8.[189,241,1152,1176]" captionTargetBox="[249,1383,254,1105]" captionTargetId="figure-432@8.[249,1383,254,1105]" captionTargetPageId="8" captionText="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752780" httpUri="https://zenodo.org/record/4752780/files/figure.png" pageId="2" pageNumber="149">Fig. 22</figureCitation>
), subanal lobes sharply recurved anteriorly at tips (
<figureCitation box="[1283,1436,336,360]" captionStart="Figs" captionStartId="8.[189,241,1152,1176]" captionTargetBox="[249,1383,254,1105]" captionTargetId="figure-432@8.[249,1383,254,1105]" captionTargetPageId="8" captionText="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752780" httpUri="https://zenodo.org/record/4752780/files/figure.png" pageId="2" pageNumber="149">Figs. 2122</figureCitation>
).………………………………..
<taxonomicName authority="Claassen" authorityName="Claassen" box="[1220,1443,372,396]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="tenella">
<emphasis bold="true" box="[1220,1331,372,396]" italics="true" pageId="2" pageNumber="149">L. tenella</emphasis>
Claassen
</taxonomicName>
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</paragraph>
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</subSubSection>
</treatment>
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