treatments-xml/data/03/DD/33/03DD337DFFF0E37FFC681AA7FACD2A8F.xml
2024-06-21 12:22:17 +02:00

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<document id="309A30CA2F0E745980C89753981243B4" ID-DOI="10.1111/j.1096-3642.2007.00285.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5429616" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630421174910" checkinUser="felipe" docAuthor="Fröbisch, Jörg" docDate="2007" docId="03DD337DFFF0E37FFC681AA7FACD2A8F" docLanguage="en" docName="j.1096-3642.2007.00285.x.pdf" docOrigin="Zoological Journal of the Linnean Society 150 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2007.00285.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Kombuisia frerensis Hotton 1974" docType="treatment" docVersion="3" lastPageNumber="119" masterDocId="FFE44B05FFF1E37DFFFB1C10FF9D2B51" masterDocTitle="The cranial anatomy of Kombuisia frerensis Hotton (Synapsida, Dicynodontia) and a new phylogeny of anomodont therapsids" masterLastPageNumber="144" masterPageNumber="117" pageNumber="118" updateTime="1699264149696" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="45A47FD6D2D3EE06E9AFDF01BABFA7A6">The cranial anatomy of Kombuisia frerensis Hotton (Synapsida, Dicynodontia) and a new phylogeny of anomodont therapsids</mods:title>
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<mods:namePart id="0A3231EF950891E516AD95204262FA2D">Fröbisch, Jörg</mods:namePart>
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<mods:title id="48B7E26054399BB99ADED71832A8807A">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="551FDC266B515178EA85F2C31465D9A9">2007</mods:date>
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<mods:identifier id="D1C8AA5A0278F7771112C1929554A857" type="DOI">10.1111/j.1096-3642.2007.00285.x</mods:identifier>
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<subSubSection id="C36ED1E0FFF0E37CFC681AA7FAD42D9E" box="[915,1353,1719,1743]" pageId="1" pageNumber="118" type="nomenclature">
<paragraph id="8BCB826BFFF0E37CFC681AA7FAD42D9E" blockId="1.[915,1353,1719,1743]" box="[915,1353,1719,1743]" pageId="1" pageNumber="118">
<heading id="D0833507FFF0E37CFC681AA7FAD42D9E" box="[915,1353,1719,1743]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="118" reason="2">
<taxonomicName id="4C74F9E8FFF0E37CFC681AA7FAD42D9E" authority="HOTTON, 1974" authorityName="Hotton" authorityYear="1974" box="[915,1353,1719,1743]" class="Reptilia" family="Dicynodontidae" genus="Kombuisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="1" pageNumber="118" phylum="Chordata" rank="species" species="frerensis">
<emphasis id="B9005E79FFF0E37CFC681AA7FB042D9C" box="[915,1177,1719,1742]" italics="true" pageId="1" pageNumber="118">KOMBUISIA FRERENSIS</emphasis>
<bibRefCitation id="EFE5FF9AFFF0E37CFB5A1AA7FAD42D9E" author="Hotton N" box="[1185,1353,1719,1743]" pageId="1" pageNumber="118" pagination="157 - 165" refId="ref14630" refString="Hotton N. 1974. A new dicynodont (Reptilia, Therapsida) from Cynognathus Zone deposits of South Africa. Annals of the South African Museum 64: 157 - 165." type="journal article" year="1974">HOTTON, 1974</bibRefCitation>
</taxonomicName>
</heading>
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<subSubSection id="C36ED1E0FFF0E37FFCC11ACFFACD2A8F" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118" type="materials_examined">
<paragraph id="8BCB826BFFF0E37FFCC11ACFFF6B2A47" blockId="1.[826,1442,1759,1904]" lastBlockId="2.[142,758,195,278]" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118">
<materialsCitation id="3B1C8836FFF0E37FFCC11ACFFF6B2A47" collectionCode="USNM" county="National Museum of Natural History" lastPageId="2" lastPageNumber="119" location="Palaeontological Research in Johannesburg" municipality="Bernard Price Institute of Palaeontological Research" pageId="1" pageNumber="118" specimenCount="2" stateProvince="Washington" typeStatus="holotype">
<emphasis id="B9005E79FFF0E37CFCC11ACFFC382DA5" box="[826,933,1759,1780]" italics="true" pageId="1" pageNumber="118">
<typeStatus id="54CF3CC9FFF0E37CFCC11ACFFC022DA5" box="[826,927,1759,1780]" pageId="1" pageNumber="118" type="holotype">Holotype</typeStatus>
:
</emphasis>
BP/1/430, an almost complete small skull with lower jaws. The specimen was originally catalogued as
<collectionCode id="ED651AAEFFF0E37CFC4F1B0DFB9E2C63" box="[948,1027,1821,1842]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="118" type="Museum">USNM</collectionCode>
<quantity id="4C8C2F8EFFF0E37CFBF51B0DFBE92C62" box="[1038,1140,1821,1843]" metricMagnitude="2" metricUnit="m" metricValue="5.825744" pageId="1" pageNumber="118" unit="in" value="22936.0">22936 in</quantity>
the
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(Smithsonian Institution) in
<collectingRegion id="49B04C89FFF0E37CFAA01B2BFC182C21" country="United States of America" name="Washington" pageId="1" pageNumber="118">Washington</collectingRegion>
, DC, but was transferred to the
<collectingMunicipality id="6BAF1811FFF0E37FFB041B4AFDCC2B88" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118">
Bernard Price Institute of
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in Johannesburg after its designation as
<typeStatus id="54CF3CC9FFF3E37FFE161CF2FDD32BA9" box="[493,590,226,248]" pageId="2" pageNumber="119" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C74F9E8FFF3E37FFD801CF2FF722A47" authorityName="Hotton" authorityYear="1974" class="Reptilia" family="Dicynodontidae" genus="Kombuisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="2" pageNumber="119" phylum="Chordata" rank="species" species="frerensis">
<emphasis id="B9005E79FFF3E37FFD801CF2FF722A47" italics="true" pageId="2" pageNumber="119">Kombuisia frerensis</emphasis>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BCB826BFFF3E37FFF751D3FFE902909" blockId="2.[142,758,303,600]" pageId="2" pageNumber="119">
<emphasis id="B9005E79FFF3E37FFF751D3FFEB82A15" box="[142,293,303,324]" italics="true" pageId="2" pageNumber="119">Type locality:</emphasis>
BP/1/430 was collected about 1.6 km south of the farm Lady Frere,
<collectingRegion id="49B04C89FFF3E37FFDF21D5EFF232AD3" country="South Africa" name="Eastern Cape" pageId="2" pageNumber="119">Eastern Cape Province</collectingRegion>
,
<collectingCountry id="F363C2FBFFF3E37FFF301D7CFEC22AD3" box="[203,351,364,386]" name="South Africa" pageId="2" pageNumber="119">South Africa</collectingCountry>
. The deposits exposed at the locality belong to the Burgersdorp Formation (Beaufort Group) of the Karoo Basin. This formation is late Early to Middle Triassic in age and corresponds to the
<taxonomicName id="4C74F9E8FFF3E37FFF441DF7FDF72AAC" authority="Assemblage Zone (AZ)" box="[191,618,487,509]" class="Reptilia" family="Cynognathidae" genus="Cynognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="2" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B9005E79FFF3E37FFF441DF7FEC82AAD" box="[191,341,487,508]" italics="true" pageId="2" pageNumber="119">Cynognathus</emphasis>
Assemblage Zone (AZ)
</taxonomicName>
(
<bibRefCitation id="EFE5FF9AFFF3E37FFD851DF7FEDF294A" author="Hancox PJ &amp; Rubidge BS" pageId="2" pageNumber="119" pagination="563 - 577" refId="ref14515" refString="Hancox PJ, Rubidge BS. 2001. Breakthroughs in the biodiversity, biogeography, biostratigraphy, and basin analysis of the Beaufort Group. Journal of African Earth Sciences 33: 563 - 577." type="journal article" year="2001">Hancox &amp; Rubidge, 2001</bibRefCitation>
). More specifically, BP/1/430 was found in subzone B (early Anisian) of the
<taxonomicName id="4C74F9E8FFF3E37FFD5C1E34FE902909" authority="AZ." authorityName="Assemblage Zone (Hancox &amp; Rubidge" authorityYear="2001" class="Reptilia" family="Cynognathidae" genus="Cynognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="2" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B9005E79FFF3E37FFD5C1E34FF402909" italics="true" pageId="2" pageNumber="119">Cynognathus</emphasis>
AZ.
</taxonomicName>
</paragraph>
<paragraph id="8BCB826BFFF3E37FFF751E61FF4A29F4" blockId="2.[142,758,625,677]" pageId="2" pageNumber="119">
<emphasis id="B9005E79FFF3E37FFF751E61FEF229D7" box="[142,367,625,646]" italics="true" pageId="2" pageNumber="119">Referred specimens:</emphasis>
NM QR1835, a snout and dentaries.
</paragraph>
<paragraph id="8BCB826BFFF3E37FFF751EADFE1C281F" blockId="2.[142,758,701,846]" pageId="2" pageNumber="119">
<emphasis id="B9005E79FFF3E37FFF751EADFEBD2983" box="[142,288,701,722]" italics="true" pageId="2" pageNumber="119">Distribution:</emphasis>
The two only known specimens (BP/1/ 430 and NM QR1835) that have been referred to
<taxonomicName id="4C74F9E8FFF3E37FFD421ECDFEA32841" authorityName="Hotton" authorityYear="1974" class="Reptilia" family="Dicynodontidae" genus="Kombuisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="2" pageNumber="119" phylum="Chordata" rank="species" species="frerensis">
<emphasis id="B9005E79FFF3E37FFD421ECDFEA32841" italics="true" pageId="2" pageNumber="119">Kombuisia frerensis</emphasis>
</taxonomicName>
are from the Burgersdorp Formation, which outcrops only in the eastern part of the South African Karoo Basin.
</paragraph>
<paragraph id="8BCB826BFFF3E37FFF751F76FD6928CA" blockId="2.[142,757,870,923]" pageId="2" pageNumber="119">
<emphasis id="B9005E79FFF3E37FFF751F76FEEA282D" box="[142,375,870,892]" italics="true" pageId="2" pageNumber="119">Stratigraphic range:</emphasis>
BP/1/430 and NM QR1835 were both collected in the middle part of the Burgersdorp
</paragraph>
<paragraph id="8BCB826BFFF3E37FFCDD1CD3FB282BA6" blockId="2.[806,1422,195,248]" pageId="2" pageNumber="119">
Formation, which corresponds to the early Anisian subzone B of the
<taxonomicName id="4C74F9E8FFF3E37FFC141CF2FB282BA6" authority="AZ." authorityName="Assemblage Zone (Hancox &amp; Rubidge" authorityYear="2001" box="[1007,1205,226,247]" class="Reptilia" family="Cynognathidae" genus="Cynognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="2" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B9005E79FFF3E37FFC141CF2FB182BA6" box="[1007,1157,226,247]" italics="true" pageId="2" pageNumber="119">Cynognathus</emphasis>
AZ.
</taxonomicName>
</paragraph>
<paragraph id="8BCB826BFFF3E37FFCDD1D00FACD2A8F" blockId="2.[806,1422,272,478]" pageId="2" pageNumber="119">
<emphasis id="B9005E79FFF3E37FFCDD1D00FC652A74" box="[806,1016,272,293]" italics="true" pageId="2" pageNumber="119">Revised diagnosis:</emphasis>
Edentulous kingoriid dicynodont autapomorphic in its possession of the following characters: absence of a pineal foramen, an inverted triangular shape of the interparietal bone, lack of fusion of articular and prearticular bones, and presence of an elongate, slender parietal posterolateral process that extends onto the occipital edge of the skull roof.
</paragraph>
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</treatment>
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