440 lines
56 KiB
XML
440 lines
56 KiB
XML
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<mods:title id="24BF87CDB1F8C8FDBBEC46DA7620ACC1">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
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<mods:namePart id="83C6186350C2046F6F4128E18F5B08A4">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="6AA8557265E6E223C94145E2F664AFC0">Email: kinkhavilay @ gmail. com & urn: lsid: zoobank. org: author: 2 E 620 B 9 C- 48 BA- 48 E 6 - A 017 - FE 8 ED 2 AB 0 E 59</mods:affiliation>
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<mods:nameIdentifier id="43A7D543F86C646B2142E6AC645C5171" type="email">kinkhavilay@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2AADEF6EC547578E7819131EEB9F7A6B">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="CB98D3D8B92FE8250127CE84366149D3">urn: lsid: zoobank. org: author: C 2 E 2 FA 6 B-A 3 F 9 - 4 F 33 - B 447 - B 59 B 1 BD 322 D 4 & Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:nameIdentifier id="AE6AA248457CDFCCA47D5721FF68375D" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="40D6D6441717274E1D27D2C469B633F3">Panha, Somsak</mods:namePart>
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<mods:affiliation id="9440F9CD8DC6B132F31DF40AC63ACDDD">urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 & Corresponding authors: jirasak 4 @ yahoo. com; somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:nameIdentifier id="E5D19D205FE1DA9903AF8F395B48FAC9" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:title id="6ED4D3DA4AFC392BFDDC8FD0557D2A06">European Journal of Taxonomy</mods:title>
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<mods:date id="2D9D415ADC65C4BE0E6D1A60B4FEC699">2017</mods:date>
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<mods:number id="3B6E7A14351B79B76BDB9DA5909FA33D">2017-06-13</mods:number>
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<treatment id="03DAD35F6935EF1FFE5CAC66FB8E2660" ID-DOI="http://doi.org/10.5281/zenodo.3848197" ID-GBIF-Taxon="164161060" ID-Zenodo-Dep="3848197" LSID="urn:lsid:plazi:treatment:03DAD35F6935EF1FFE5CAC66FB8E2660" httpUri="http://treatment.plazi.org/id/03DAD35F6935EF1FFE5CAC66FB8E2660" lastPageId="23" lastPageNumber="24" pageId="19" pageNumber="20">
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<subSubSection id="C36931C26935EF1BFE5CAC66FB942610" box="[471,1116,856,883]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8BCC62496935EF1BFE5CAC66FB942610" blockId="19.[471,1116,856,919]" box="[471,1116,856,883]" pageId="19" pageNumber="20">
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<heading id="D084D5256935EF1BFE5CAC66FB942610" box="[471,1116,856,883]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
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<taxonomicName id="4C7319CA6935EF1BFE5CAC66FB942610" ID-CoL="D2K6" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[471,1116,856,883]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="areolatus" subGenus="Syndromus">
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<emphasis id="B907BE5B6935EF1BFE5CAC66FC572610" bold="true" box="[471,927,856,883]" pageId="19" pageNumber="20">
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<emphasis id="B907BE5B6935EF1BFE5CAC66FD4C2610" bold="true" box="[471,644,856,883]" italics="true" pageId="19" pageNumber="20">Amphidromus</emphasis>
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(
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<emphasis id="B907BE5B6935EF1BFD1EAC66FCD62610" bold="true" box="[661,798,856,883]" italics="true" pageId="19" pageNumber="20">Syndromus</emphasis>
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)
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<emphasis id="B907BE5B6935EF1BFCA4AC66FC572610" bold="true" box="[815,927,856,883]" italics="true" pageId="19" pageNumber="20">areolatus</emphasis>
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</emphasis>
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(Pfeiffer, 1861)
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36931C26935EF1BFD94AC42FBDD26F4" box="[543,1045,892,919]" pageId="19" pageNumber="20" type="description">
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<paragraph id="8BCC62496935EF1BFD94AC42FBDD26F4" blockId="19.[471,1116,856,919]" box="[543,1045,892,919]" pageId="19" pageNumber="20">
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<figureCitation id="13487ECC6935EF1BFD94AC42FDA026F4" box="[543,616,892,919]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="19" pageNumber="20">Figs 1</figureCitation>
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,
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<figureCitation id="13487ECC6935EF1BFDFDAC43FD4C26F4" box="[630,644,893,919]" captionStart="Fig" captionStartId="12.[189,232,1823,1849]" captionTargetBox="[196,1393,710,1775]" captionTargetId="figure@12.[189,1399,704,1781]" captionTargetPageId="12" captionText="Fig. 7. SEM images of the radula. A–C. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos (CUMZ 7012). D–F. Amphidromus syndromoideus sp. nov., holotype (CUMZ 7019). G–I. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapue, Laos (CUMZ 7023). A, D, G = central tooth with the first to fifth to eighth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829758" httpUri="https://zenodo.org/record/3829758/files/figure.png" pageId="19" pageNumber="20">7</figureCitation>
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G−I, 9A, 10A−C, 11A–B;
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<tableCitation id="C6F157F26935EF1BFC35AC42FBDD26F4" box="[958,1045,892,919]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetId="graphics@3.[189,1398,1154,1193]" captionTargetPageId="3" captionText="Table 1 (continued on next page). Shell measurements of nine species of Amphidromus from Laos. Specimen catalogue numbers are indicated in parentheses." httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="19" pageNumber="20" tableUuid="DF0C32C16925EF0BFF36AE32FCA1242B">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36931C26935EF1BFF36AC81FA9F2100" pageId="19" pageNumber="20" type="reference_group">
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<paragraph id="8BCC62496935EF1BFF36AC81FEFA269D" blockId="19.[189,1398,959,1022]" pageId="19" pageNumber="20">
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<treatmentCitationGroup id="AB6345676935EF1BFF36AC81FEFA269D" pageId="19" pageNumber="20">
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<treatmentCitation id="0AD244586935EF1BFF36AC81FD4E26BA" author="Pfeiffer L." box="[189,646,959,985]" page="194" pageId="19" pageNumber="20" year="1861">
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<taxonomicName id="4C7319CA6935EF1BFF36AC81FD4E26BA" authority="Pfeiffer, 1861 a: 194" authorityName="Pfeiffer" authorityPageNumber="194" authorityYear="1861" box="[189,646,959,985]" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="areolatus">
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<emphasis id="B907BE5B6935EF1BFF36AC81FE5A26BA" box="[189,402,959,985]" italics="true" pageId="19" pageNumber="20">Bulimus areolatus</emphasis>
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<bibRefCitation id="EFE21FB86935EF1BFE12AC81FD4E26BA" author="Pfeiffer L." box="[409,646,959,985]" pageId="19" pageNumber="20" pagination="190 - 196" refId="ref19251" refString="Pfeiffer L. 1861 a. Descriptions of new land-shells, in the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London 29: 190 - 196. (Published in parts, dates follow Duncan 1937.) Available from http: // biodiversitylibrary. org / page / 28672714 [accessed 9 Sept. 2013]." type="journal article" year="1861">Pfeiffer, 1861a: 194</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="54C8DCEB6935EF1BFD1AACFEFD0326B9" box="[657,715,960,986]" pageId="19" pageNumber="20">Type</typeStatus>
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locality:
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<collectingCountry id="F36422D96935EF1BFCB1AC81FCBE26BA" box="[826,886,959,985]" name="Thailand" pageId="19" pageNumber="20">Siam</collectingCountry>
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||
[
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<collectingCountry id="F36422D96935EF1BFC0FAC81FC3E26B9" box="[900,1014,959,986]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
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].
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<treatmentCitation id="0AD244586935EF1BFB8CAC81FB2726BA" author="Pfeiffer L." box="[1031,1263,959,985]" page="172" pageId="19" pageNumber="20" year="1861">
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<bibRefCitation id="EFE21FB86935EF1BFB8CAC81FB2726BA" author="Pfeiffer L." box="[1031,1263,959,985]" pageId="19" pageNumber="20" pagination="161 - 178" refId="ref19316" refString="Pfeiffer L. 1861 b (1860 - 1866). Novitates Conchologicae. Series Prima Mollusca Extramarina, Vol. 2: 139 - 303, pls 37 - 72 (1861: 161 - 178, pls 43 - 48). Theodor Fischer Verlag, Kassel. (Published in parts, dates follow Johnson 1969.) Available from http: // biodiversitylibrary. org / item / 122926 [accessed 8 Jun. 2016]." type="journal article" year="1861">Pfeiffer 1861b: 172</bibRefCitation>
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</treatmentCitation>
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, pl. 46, figs 11–12.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCC62496935EF1BFF36AB1BFA9F2100" blockId="19.[189,1399,1061,1124]" pageId="19" pageNumber="20">
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<treatmentCitationGroup id="AB6345676935EF1BFF36AB1BFA9F2100" pageId="19" pageNumber="20">
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<taxonomicName id="4C7319CA6935EF1BFF36AB1BFE1E215C" ID-CoL="D2K6" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[189,470,1061,1087]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="areolatus">
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<emphasis id="B907BE5B6935EF1BFF36AB1BFE1E215C" box="[189,470,1061,1087]" italics="true" pageId="19" pageNumber="20">Amphidromus areolatus</emphasis>
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</taxonomicName>
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||
–
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<treatmentCitation id="0AD244586935EF1BFE79AB1BFD78215C" author="Fulton H. C." box="[498,688,1061,1087]" page="81" pageId="19" pageNumber="20" year="1896">
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<bibRefCitation id="EFE21FB86935EF1BFE79AB1BFD78215C" author="Fulton H. C." box="[498,688,1061,1087]" pageId="19" pageNumber="20" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton 1896: 81</bibRefCitation>
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</treatmentCitation>
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||
. —
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||
<treatmentCitation id="0AD244586935EF1BFD6BAB1BFC26215C" author="Pilsbry H. A." box="[736,1006,1061,1087]" page="198" pageId="19" pageNumber="20" year="1900">
|
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<bibRefCitation id="EFE21FB86935EF1BFD6BAB1BFC26215C" author="Pilsbry H. A." box="[736,1006,1061,1087]" pageId="19" pageNumber="20" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 198–199</bibRefCitation>
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||
</treatmentCitation>
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||
, pl. 63, figs 85–86. —
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<treatmentCitation id="0AD244586935EF1BFB73AB1BFE672107" author="Laidlaw F. F. & Solem A." page="564" pageId="19" pageNumber="20" year="1961">
|
||
<bibRefCitation id="EFE21FB86935EF1BFB73AB1BFE672107" author="Laidlaw F. F. & Solem A." pageId="19" pageNumber="20" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. & Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">
|
||
Laidlaw & Solem
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||
<date id="FFCD44896935EF1BFEBEAB77FEBC2100" box="[309,372,1097,1123]" pageId="19" pageNumber="20" value="1961">1961</date>
|
||
: 564
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||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AD244586935EF1BFE54AB77FD2C2107" author="Solem A." box="[479,740,1097,1124]" page="624" pageId="19" pageNumber="20" year="1965">
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||
<bibRefCitation id="EFE21FB86935EF1BFE54AB77FD2C2107" author="Solem A." box="[479,740,1097,1124]" pageId="19" pageNumber="20" pagination="615 - 628" refId="ref19790" refString="Solem A. 1965. Land snails of the genus Amphidromus from Thailand (Mollusca: Pumonata: Camaenidae). Proceedings of the United States National Museum 117: 615 - 628. Available from http: // biodiversitylibrary. org / page / 7759057 [accessed 8 Jun. 2016]." type="journal article" year="1965">Solem 1965: 624–625</bibRefCitation>
|
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</treatmentCitation>
|
||
, pl. 1, figs 4–7. —
|
||
<treatmentCitation id="0AD244586935EF1BFC4FAB77FB2A2107" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[964,1250,1097,1124]" httpUri="http://treatment.plazi.org/id/506E173437547B027D9D2A8FF8DBA4DC" page="58" pageId="19" pageNumber="20" year="2015">
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<bibRefCitation id="EFE21FB86935EF1BFC4FAB77FB2A2107" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[964,1250,1097,1124]" pageId="19" pageNumber="20" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
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||
Sutcharit
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<emphasis id="B907BE5B6935EF1BFBBEAB74FBA12100" box="[1077,1129,1097,1123]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
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||
. 2015: 58
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||
</bibRefCitation>
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</treatmentCitation>
|
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, fig. 3j–k.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36931C26935EF1BFF36ABB4FEA720AF" pageId="19" pageNumber="20" type="materials_examined">
|
||
<paragraph id="8BCC62496935EF1BFF36ABB4FE6221C7" blockId="19.[189,426,1162,1188]" box="[189,426,1162,1188]" pageId="19" pageNumber="20">
|
||
<heading id="D084D5256935EF1BFF36ABB4FE6221C7" bold="true" box="[189,426,1162,1188]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B907BE5B6935EF1BFF36ABB4FE6221C7" bold="true" box="[189,426,1162,1188]" pageId="19" pageNumber="20">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496935EF1BFF61AB85FE5121B7" blockId="19.[189,1399,1210,1310]" box="[234,409,1210,1237]" pageId="19" pageNumber="20">
|
||
<emphasis id="B907BE5B6935EF1BFF61AB85FE5121B7" bold="true" box="[234,409,1210,1237]" pageId="19" pageNumber="20">
|
||
<typeStatus id="54C8DCEB6935EF1BFF61AB85FEEF21B6" box="[234,295,1211,1237]" pageId="19" pageNumber="20">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496935EF1BFF36ABE1FE6A207D" blockId="19.[189,1399,1210,1310]" pageId="19" pageNumber="20">
|
||
<collectingCountry id="F36422D96935EF1BFF36ABE1FE91219A" box="[189,345,1247,1273]" name="Thailand" pageId="19" pageNumber="20">THAILAND</collectingCountry>
|
||
:
|
||
<typeStatus id="54C8DCEB6935EF1BFEE3ABE1FE15219A" box="[360,477,1247,1273]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
|
||
<collectionCode id="ED62FA8C6935EF1BFE63ABE1FD9F219A" box="[488,599,1247,1273]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="20" type="Museum">NHMUK</collectionCode>
|
||
19601430 (
|
||
<bibRefCitation id="EFE21FB86935EF1BFD61ABE1FC222199" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[746,1002,1247,1274]" pageId="19" pageNumber="20" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
|
||
Sutcharit
|
||
<emphasis id="B907BE5B6935EF1BFCE9ABDEFC51219A" box="[866,921,1247,1273]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 3j);
|
||
<typeStatus id="54C8DCEB6935EF1BFBD5ABE1FB34219A" box="[1118,1276,1247,1273]" pageId="19" pageNumber="20" type="paralectotype">paralectotype</typeStatus>
|
||
<collectionCode id="ED62FA8C6935EF1BFA83ABE1FABF219A" box="[1288,1399,1247,1273]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="19" pageNumber="20" type="Museum">NHMUK</collectionCode>
|
||
19601431 (1 shell).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496935EF1BFF61AA7AFE6E203D" blockId="19.[189,1399,1348,1484]" box="[234,422,1348,1374]" pageId="19" pageNumber="20">
|
||
<emphasis id="B907BE5B6935EF1BFF61AA7AFE6E203D" bold="true" box="[234,422,1348,1374]" pageId="19" pageNumber="20">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496935EF1BFF36AA57FEA720AF" blockId="19.[189,1399,1348,1484]" pageId="19" pageNumber="20">
|
||
<collectingCountry id="F36422D96935EF1BFF36AA57FEC620E0" box="[189,270,1385,1411]" name="Laos" pageId="19" pageNumber="20">LAOS</collectingCountry>
|
||
: 20 shells (
|
||
<figureCitation id="13487ECC6935EF1BFE1AAA57FE2C20E0" box="[401,484,1385,1411]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
|
||
A−C,
|
||
<collectionCode id="ED62FA8C6935EF1BFDA5AA57FD4B20E0" box="[558,643,1385,1411]" country="0" name="Cameroon University, Museum of Zoology" pageId="19" pageNumber="20">CUMZ</collectionCode>
|
||
7022),
|
||
<specimenCount id="9D75A9C06935EF1BFD55AA57FCB420E0" box="[734,892,1385,1411]" pageId="19" pageNumber="20" type="generic">17 specimens</specimenCount>
|
||
in freezer (
|
||
<figureCitation id="13487ECC6935EF1BFB8FAA57FB8520E0" box="[1028,1101,1385,1411]" captionStart="Fig" captionStartId="12.[189,232,1823,1849]" captionTargetBox="[196,1393,710,1775]" captionTargetId="figure@12.[189,1399,704,1781]" captionTargetPageId="12" captionText="Fig. 7. SEM images of the radula. A–C. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos (CUMZ 7012). D–F. Amphidromus syndromoideus sp. nov., holotype (CUMZ 7019). G–I. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapue, Laos (CUMZ 7023). A, D, G = central tooth with the first to fifth to eighth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829758" httpUri="https://zenodo.org/record/3829758/files/figure.png" pageId="19" pageNumber="20">Figs 7</figureCitation>
|
||
G−I, 9
|
||
<collectionCode id="ED62FA8C6935EF1BFB10AA57FB7B20E0" box="[1179,1203,1385,1411]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="19" pageNumber="20" type="Herbarium">A</collectionCode>
|
||
, 11A–B,
|
||
<collectionCode id="ED62FA8C6935EF1BFAA9AA57FABF20E0" box="[1314,1399,1385,1411]" country="0" name="Cameroon University, Museum of Zoology" pageId="19" pageNumber="20">CUMZ</collectionCode>
|
||
7023), Thad Fek, Sammakeexay District,
|
||
<collectingRegion id="49B7ACAB6935EF1BFD10AAB0FD3420CB" box="[667,764,1422,1448]" country="Laos" name="Attapu" pageId="19" pageNumber="20">Attapeu</collectingRegion>
|
||
; 8 shells, Thad Phasoam, Paksong District,
|
||
<collectingRegion id="49B7ACAB6935EF1BFB65AAB3FABF20C4" box="[1262,1399,1421,1447]" country="Laos" name="Champasak" pageId="19" pageNumber="20">Champasak</collectingRegion>
|
||
(
|
||
<collectionCode id="ED62FA8C6935EF1BFF4EAA8CFED420AF" box="[197,284,1458,1484]" country="0" name="Cameroon University, Museum of Zoology" pageId="19" pageNumber="20">CUMZ</collectionCode>
|
||
7024).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C26935EF1FFF36AACDFAFE242A" lastPageId="23" lastPageNumber="24" pageId="19" pageNumber="20" type="description">
|
||
<paragraph id="8BCC62496935EF1BFF36AACDFE86236E" blockId="19.[189,334,1523,1549]" box="[189,334,1523,1549]" pageId="19" pageNumber="20">
|
||
<heading id="D084D5256935EF1BFF36AACDFE86236E" bold="true" box="[189,334,1523,1549]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B907BE5B6935EF1BFF36AACDFE86236E" bold="true" box="[189,334,1523,1549]" pageId="19" pageNumber="20">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496935EF1BFF36A91DFBAE2390" blockId="19.[189,1399,1571,1780]" pageId="19" pageNumber="20">Shell sinistral, elongate conical, rather small, thin and glossy. Spire conical; apex acute, with black spot on tip. Whorls 5 to 6 slightly convex; suture depressed; last whorl rounded. Periostracum transparent to thin corneous. Last whorl process, with yellow to absent Band 1; Bands 2 to 4 (sometimes Bands 2 to 5) usually merge and become brownish slanted blotches with flame shape; Band 6 yellowish or roseate. Parietal callus thin and transparent. Aperture auriform; peristome expanded to weakly expanded; lip white to transparent. Columella straight and white. Umbilicus narrowly opened.</paragraph>
|
||
<paragraph id="8BCC62496935EF1BFF61A825FE8D2256" blockId="19.[189,1399,1819,2027]" box="[234,325,1819,1845]" pageId="19" pageNumber="20">
|
||
<emphasis id="B907BE5B6935EF1BFF61A825FE8D2256" bold="true" box="[234,325,1819,1845]" pageId="19" pageNumber="20">Radula</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496935EF1CFF36A801FDA6242A" blockId="19.[189,1399,1819,2027]" lastBlockId="20.[189,1398,267,329]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
Teeth arranged in V-shaped rows, each row contains about 126 (62-(13-9)-1-(9-13)-63) teeth. Central tooth tricuspid, with triangular shaped mesocone, ectocones very small, with pointed cusp. Lateral teeth bicuspid, endocone small, with curved cusp; ectocone large, with truncate or curved cusps, gradually transformed to tricuspid marginal teeth. Tricuspid marginal teeth start around tooth number 9 to 13 outwards; endocone curve shaped; mesocone of largest size, with long and obtuse cusp; ectocone smallest, with pointed cusp. Outermost teeth with small serrated endocone and extocone; mesocone large, with curved cusps (
|
||
<figureCitation id="13487ECC6932EF1CFE61AE10FDE6242A" box="[490,558,302,329]" captionStart="Fig" captionStartId="12.[189,232,1823,1849]" captionTargetBox="[196,1393,710,1775]" captionTargetId="figure@12.[189,1399,704,1781]" captionTargetPageId="12" captionText="Fig. 7. SEM images of the radula. A–C. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos (CUMZ 7012). D–F. Amphidromus syndromoideus sp. nov., holotype (CUMZ 7019). G–I. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapue, Laos (CUMZ 7023). A, D, G = central tooth with the first to fifth to eighth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829758" httpUri="https://zenodo.org/record/3829758/files/figure.png" pageId="20" pageNumber="21">Fig. 7</figureCitation>
|
||
G–I).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496932EF1CFF61AE50FE6C24EA" blockId="20.[189,1399,366,533]" box="[234,420,366,393]" pageId="20" pageNumber="21">
|
||
<emphasis id="B907BE5B6932EF1CFF61AE50FE6C24EA" bold="true" box="[234,420,366,393]" pageId="20" pageNumber="21">Genital organs</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496932EF1CFF36AEACFE562776" blockId="20.[189,1399,366,533]" pageId="20" pageNumber="21">
|
||
Atrium (at) rather long (n = 10). Penis (p) long, cylindrical and enlarged. Epiphallus (e) short, of about same length as penis; flagellum (fl) short and terminated in pointed tip. Appendix absent. Penial retractor muscle (pr) thin and relatively long. Vas deferens (vd) narrow tube connecting epiphallus and free oviduct (
|
||
<figureCitation id="13487ECC6932EF1CFEAEAEC5FE592776" box="[293,401,507,533]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="20" pageNumber="21">Fig. 11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0C32C16932EF1CFF36A821FBE1228A" ID-DOI="http://doi.org/10.5281/zenodo.3829762" ID-Zenodo-Dep="3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1823,1849]" targetBox="[189,1399,656,1782]" targetPageId="20">
|
||
<paragraph id="8BCC62496932EF1CFF36A821FBE1228A" blockId="20.[189,1399,1823,2026]" pageId="20" pageNumber="21">
|
||
<emphasis id="B907BE5B6932EF1CFF36A821FEC6225A" bold="true" box="[189,270,1823,1849]" pageId="20" pageNumber="21">Fig. 9.</emphasis>
|
||
Living snails of
|
||
<taxonomicName id="4C7319CA6932EF1CFE6FA821FCE02259" authority="" authorityName="Pilsbry" authorityYear="1900" box="[484,808,1823,1850]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="subGenus" subGenus="Syndromus">
|
||
<emphasis id="B907BE5B6932EF1CFE6FA821FD40225A" box="[484,648,1823,1849]" italics="true" pageId="20" pageNumber="21">Amphidromus</emphasis>
|
||
(
|
||
<emphasis id="B907BE5B6932EF1CFD16A821FCD6225A" box="[669,798,1823,1849]" italics="true" pageId="20" pageNumber="21">Syndromus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B907BE5B6932EF1CFCE5A821FC4B225A" bold="true" box="[878,899,1823,1849]" pageId="20" pageNumber="21">A</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6932EF1CFC1DA821FABF2259" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[918,1399,1823,1850]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="areolatus">
|
||
<emphasis id="B907BE5B6932EF1CFC1DA821FB7B225A" box="[918,1203,1823,1849]" italics="true" pageId="20" pageNumber="21">Amphidromus areolatus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
from Thad Fek, Attapeu, Laos (CUMZ 7023).
|
||
<emphasis id="B907BE5B6932EF1CFD78A87DFCCF223E" bold="true" box="[755,775,1859,1885]" pageId="20" pageNumber="21">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6932EF1CFC92A87CFB05223E" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[793,1229,1858,1885]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B6932EF1CFC92A87CFBC2223F" box="[793,1034,1858,1884]" italics="true" pageId="20" pageNumber="21">Amphidromus flavus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027).
|
||
<emphasis id="B907BE5B6932EF1CFD68A85BFCD4221C" bold="true" box="[739,796,1893,1919]" pageId="20" pageNumber="21">C–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6932EF1CFCA0A858FB3A22E3" authority="Morlet, 1891" authorityName="Morlet" authorityYear="1891" box="[811,1266,1894,1920]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="xiengensis">
|
||
<emphasis id="B907BE5B6932EF1CFCA0A858FB98221C" box="[811,1104,1894,1920]" italics="true" pageId="20" pageNumber="21">Amphidromus xiengensis</emphasis>
|
||
Morlet, 1891
|
||
</taxonomicName>
|
||
from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “
|
||
<emphasis id="B907BE5B6932EF1CFBA6A8B7FB0E22C0" box="[1069,1222,1929,1955]" italics="true" pageId="20" pageNumber="21">multifasciata</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB86932EF1CFB51A8B7FAB922C0" author="Fulton H. C." box="[1242,1393,1929,1955]" pageId="20" pageNumber="21" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
|
||
, respectively.
|
||
<emphasis id="B907BE5B6932EF1CFEDDA892FE4222A5" bold="true" box="[342,394,1964,1990]" pageId="20" pageNumber="21">E–F</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6932EF1CFE13A892FC6A22A5" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[408,930,1964,1990]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="fuscolabris">
|
||
<emphasis id="B907BE5B6932EF1CFE13A892FD0C22A5" box="[408,708,1964,1990]" italics="true" pageId="20" pageNumber="21">Amphidromus fuscolabris</emphasis>
|
||
<bibRefCitation id="EFE21FB86932EF1CFD41A892FC6A22A5" author="Mollendorff O. F. von" box="[714,930,1964,1990]" pageId="20" pageNumber="21" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
from Ban Phone, Sekong, Laos (CUMZ 7041–7042).
|
||
<emphasis id="B907BE5B6932EF1CFED2A8F1FEA5228A" bold="true" box="[345,365,1999,2025]" pageId="20" pageNumber="21">E</emphasis>
|
||
. Typical color form.
|
||
<emphasis id="B907BE5B6932EF1CFDEEA8F1FDBF228A" bold="true" box="[613,631,1999,2025]" pageId="20" pageNumber="21">F</emphasis>
|
||
. Monochrome yellowish color form.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC62496933EF1DFF36AE35FBAF242A" blockId="21.[189,1398,267,329]" pageId="21" pageNumber="22">
|
||
Internal penial wall nearly smooth and corrugated into thin penial pilasters, which form fringe around penial verge. Penial verge rather small, conical, with smooth surface (
|
||
<figureCitation id="13487ECC6933EF1DFC64AE10FB91242A" box="[1007,1113,302,329]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="21" pageNumber="22">Fig. 11B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496933EF1DFF36AE51FA9D24AC" blockId="21.[189,1399,367,464]" pageId="21" pageNumber="22">
|
||
Vagina cylindrical, of about same length as penis. Gametolytic duct long, proximal to genital orifice, of larger diameter than vagina and coiled; distal to genital orifice tapering to small tube of about same length as proximal part and connected to gametolytic sac. Oviduct and albumin gland very small (
|
||
<figureCitation id="13487ECC6933EF1DFB56AE8BFA8024AC" box="[1245,1352,437,463]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="21" pageNumber="22">Fig. 11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF0C32C16933EF1DFF36A9E6FD7C228A" ID-DOI="http://doi.org/10.5281/zenodo.3829764" ID-Zenodo-Dep="3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1752,1778]" targetBox="[189,1399,539,1707]" targetPageId="21">
|
||
<paragraph id="8BCC62496933EF1DFF36A9E6FD7C228A" blockId="21.[189,1399,1752,2026]" pageId="21" pageNumber="22">
|
||
<emphasis id="B907BE5B6933EF1DFF36A9E6FED22390" bold="true" box="[189,282,1752,1779]" pageId="21" pageNumber="22">Fig. 10.</emphasis>
|
||
Shells of
|
||
<taxonomicName id="4C7319CA6933EF1DFE18A9E7FD1B2390" authority="" authorityName="Pilsbry" authorityYear="1900" box="[403,723,1753,1779]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="subGenus" subGenus="Syndromus">
|
||
<emphasis id="B907BE5B6933EF1DFE18A9E7FDFF2390" box="[403,567,1753,1779]" italics="true" pageId="21" pageNumber="22">Amphidromus</emphasis>
|
||
(
|
||
<emphasis id="B907BE5B6933EF1DFDC3A9E7FD012390" box="[584,713,1753,1779]" italics="true" pageId="21" pageNumber="22">Syndromus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B907BE5B6933EF1DFC98A9E6FC862391" bold="true" box="[787,846,1752,1778]" pageId="21" pageNumber="22">A−C</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6933EF1DFCD5A9E7FAFE2397" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[862,1334,1753,1780]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="areolatus">
|
||
<emphasis id="B907BE5B6933EF1DFCD5A9E7FBB02390" box="[862,1144,1753,1779]" italics="true" pageId="21" pageNumber="22">Amphidromus areolatus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
from Thad Fek, Attapeu, Laos (CUMZ 7022).
|
||
<emphasis id="B907BE5B6933EF1DFD2BA9C2FD7D2275" bold="true" box="[672,693,1788,1814]" pageId="21" pageNumber="22">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6933EF1DFD4FA9C2FBA72274" authority="(Morlet, 1886)" baseAuthorityName="Morlet" baseAuthorityYear="1886" box="[708,1135,1788,1815]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="begini">
|
||
<emphasis id="B907BE5B6933EF1DFD4FA9C2FC702275" box="[708,952,1788,1814]" italics="true" pageId="21" pageNumber="22">Amphidromus begini</emphasis>
|
||
(Morlet, 1886)
|
||
</taxonomicName>
|
||
, syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832).
|
||
<emphasis id="B907BE5B6933EF1DFD42A821FD35225A" bold="true" box="[713,765,1823,1849]" pageId="21" pageNumber="22">E-G</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6933EF1DFC85A821FB092259" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[782,1217,1823,1850]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B6933EF1DFC85A821FC37225A" box="[782,1023,1823,1849]" italics="true" pageId="21" pageNumber="22">Amphidromus flavus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
from Tam Pou Kham, Vientiane, Laos (CUMZ 7029).
|
||
<emphasis id="B907BE5B6933EF1DFDF4A87DFD5E223E" bold="true" box="[639,662,1859,1885]" pageId="21" pageNumber="22">H</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6933EF1DFD28A87CFC47223F" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[675,911,1858,1884]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B6933EF1DFD28A87CFC47223F" box="[675,911,1858,1884]" italics="true" pageId="21" pageNumber="22">Amphidromus flavus</emphasis>
|
||
</taxonomicName>
|
||
var. “
|
||
<emphasis id="B907BE5B6933EF1DFC59A87CFB91223F" box="[978,1113,1858,1884]" italics="true" pageId="21" pageNumber="22">indistinctus</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB86933EF1DFBE0A87CFACE223F" author="Pilsbry H. A." box="[1131,1286,1858,1884]" pageId="21" pageNumber="22" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
|
||
, holotype (ANSP 31486).
|
||
<emphasis id="B907BE5B6933EF1DFE08A858FE7C22E3" bold="true" box="[387,436,1894,1920]" pageId="21" pageNumber="22">I–K</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6933EF1DFE4CA85BFD73221C" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[455,699,1893,1919]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B6933EF1DFE4CA85BFD73221C" box="[455,699,1893,1919]" italics="true" pageId="21" pageNumber="22">Amphidromus flavus</emphasis>
|
||
</taxonomicName>
|
||
var. “
|
||
<emphasis id="B907BE5B6933EF1DFC80A858FC99221C" box="[779,849,1894,1919]" italics="true" pageId="21" pageNumber="22">tryoni</emphasis>
|
||
”
|
||
<bibRefCitation id="EFE21FB86933EF1DFCE1A85BFBC522E3" author="Pilsbry H. A." box="[874,1037,1893,1920]" pageId="21" pageNumber="22" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry, 1900</bibRefCitation>
|
||
.
|
||
<emphasis id="B907BE5B6933EF1DFB97A858FBEF22E3" bold="true" box="[1052,1063,1894,1920]" pageId="21" pageNumber="22">I</emphasis>
|
||
. Lectotype (ANSP 31488).
|
||
<emphasis id="B907BE5B6933EF1DFF36A8B7FF0422C0" bold="true" box="[189,204,1929,1955]" pageId="21" pageNumber="22">J</emphasis>
|
||
. Paralectotype (ANSP 252745).
|
||
<emphasis id="B907BE5B6933EF1DFDDCA8B7FDA622C0" bold="true" box="[599,622,1929,1955]" pageId="21" pageNumber="22">K</emphasis>
|
||
. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026).
|
||
<emphasis id="B907BE5B6933EF1DFE9DA892FE9C22A5" bold="true" box="[278,340,1964,1990]" pageId="21" pageNumber="22">L–M</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6933EF1DFEE2A892FC4722A5" authority="(Morlet, 1884)" baseAuthorityName="Morlet" baseAuthorityYear="1884" box="[361,911,1964,1990]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="semitessellatus">
|
||
<emphasis id="B907BE5B6933EF1DFEE2A892FD0222A5" box="[361,714,1964,1990]" italics="true" pageId="21" pageNumber="22">Amphidromus semitessellatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE21FB86933EF1DFD6BA892FC4E22A5" author="Morlet L." box="[736,902,1964,1990]" pageId="21" pageNumber="22" pagination="386 - 403" refId="ref18917" refString="Morlet L. 1884. Description d'especes nouvelles de coquilles recueillies par M. Pavie au Cambodge. Journal de Conchyliologie 32: 386 - 403. Available from http: // biodiversitylibrary. org / page / 16072179 [accessed 9 Sept. 2013]." type="journal article" year="1884">Morlet, 1884</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B907BE5B6933EF1DFC2BA892FC7C22A5" bold="true" box="[928,948,1964,1990]" pageId="21" pageNumber="22">L</emphasis>
|
||
. Lectotype (MNHN-IM-2000-1985).
|
||
<emphasis id="B907BE5B6933EF1DFF36A8F1FF11228A" bold="true" box="[189,217,1999,2025]" pageId="21" pageNumber="22">M</emphasis>
|
||
. Specimen from Cambodia (NHMUK).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C32C16930EF1EFF36A80AFCE622B8" ID-DOI="http://doi.org/10.5281/zenodo.3829766" ID-Zenodo-Dep="3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1844,1870]" targetBox="[189,1399,298,1799]" targetPageId="22">
|
||
<paragraph id="8BCC62496930EF1EFF36A80AFCE622B8" blockId="22.[189,1399,1844,2011]" pageId="22" pageNumber="23">
|
||
<emphasis id="B907BE5B6930EF1EFF36A80AFED4222D" bold="true" box="[189,284,1844,1870]" pageId="22" pageNumber="23">Fig. 11.</emphasis>
|
||
Genitalia of
|
||
<taxonomicName id="4C7319CA6930EF1EFE4AA80AFCCD222C" authority="" authorityName="Pilsbry" authorityYear="1900" box="[449,773,1844,1871]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="subGenus" subGenus="Syndromus">
|
||
<emphasis id="B907BE5B6930EF1EFE4AA80AFDAD222D" box="[449,613,1844,1870]" italics="true" pageId="22" pageNumber="23">Amphidromus</emphasis>
|
||
(
|
||
<emphasis id="B907BE5B6930EF1EFDF1A80AFD33222D" box="[634,763,1844,1870]" italics="true" pageId="22" pageNumber="23">Syndromus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="B907BE5B6930EF1EFCC7A80AFC4B222D" bold="true" box="[844,899,1844,1870]" pageId="22" pageNumber="23">A–B</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6930EF1EFC1DA80AFABF222C" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[918,1399,1844,1871]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="areolatus">
|
||
<emphasis id="B907BE5B6930EF1EFC1DA80AFB7B222D" box="[918,1203,1844,1870]" italics="true" pageId="22" pageNumber="23">Amphidromus areolatus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023).
|
||
<emphasis id="B907BE5B6930EF1EFDB6A844FDBE22F7" bold="true" box="[573,630,1914,1940]" pageId="22" pageNumber="23">C–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C7319CA6930EF1EFD0FA844FBE622F6" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[644,1070,1914,1941]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="flavus">
|
||
<emphasis id="B907BE5B6930EF1EFD0FA844FCBA22F7" box="[644,882,1914,1940]" italics="true" pageId="22" pageNumber="23">Amphidromus flavus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC62496931EF1FFF36AE35FAFE242A" blockId="23.[189,1398,267,329]" pageId="23" pageNumber="24">
|
||
Internal wall of vagina shows longitudinal vaginal pilasters (vp); proximal to genital orifice with nearly smooth surface for about half of its length, distally pilasters become corrugated ridges (
|
||
<figureCitation id="13487ECC6931EF1FFB35AE11FAE1242A" box="[1214,1321,303,329]" captionStart="Fig" captionStartId="22.[189,232,1844,1870]" captionTargetBox="[189,1399,298,1799]" captionTargetId="figure@22.[189,1399,298,1799]" captionTargetPageId="22" captionText="Fig. 11. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7023). C–D. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vagina chamber (CUMZ 7027)." figureDoi="http://doi.org/10.5281/zenodo.3829766" httpUri="https://zenodo.org/record/3829766/files/figure.png" pageId="23" pageNumber="24">Fig. 11B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C26931EF1FFF36AE4FFAC624BD" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="8BCC62496931EF1FFF36AE4FFE9E24E8" blockId="23.[189,342,369,395]" box="[189,342,369,395]" pageId="23" pageNumber="24">
|
||
<heading id="D084D5256931EF1FFF36AE4FFE9E24E8" bold="true" box="[189,342,369,395]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
|
||
<emphasis id="B907BE5B6931EF1FFF36AE4FFE9E24E8" bold="true" box="[189,342,369,395]" pageId="23" pageNumber="24">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36AE9EFAC624BD" blockId="23.[189,1398,416,479]" pageId="23" pageNumber="24">
|
||
The species is known from southern
|
||
<collectingCountry id="F36422D96931EF1FFDD2AE9FFD5D24D8" box="[601,661,417,443]" name="Laos" pageId="23" pageNumber="24">Laos</collectingCountry>
|
||
, where the specimens were collected at Thad Fek, Sammakeexay District, Attapue and Thad Phasoam, Paksong District,
|
||
<collectingRegion id="49B7ACAB6931EF1FFCCEAEFAFC0624BD" box="[837,974,452,478]" country="Laos" name="Champasak" pageId="23" pageNumber="24">Champasak</collectingRegion>
|
||
in a dry dipterocarp forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C26931EF1FFF36AD38FB8E2660" pageId="23" pageNumber="24" type="discussion">
|
||
<paragraph id="8BCC62496931EF1FFF36AD38FEE72743" blockId="23.[189,303,518,544]" box="[189,303,518,544]" pageId="23" pageNumber="24">
|
||
<heading id="D084D5256931EF1FFF36AD38FEE72743" bold="true" box="[189,303,518,544]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
|
||
<emphasis id="B907BE5B6931EF1FFF36AD38FEE72743" bold="true" box="[189,303,518,544]" pageId="23" pageNumber="24">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496931EF1FFF36AD0BFB8E2660" blockId="23.[189,1399,565,771]" pageId="23" pageNumber="24">
|
||
This species can be distinguished from
|
||
<taxonomicName id="4C7319CA6931EF1FFD1BAD08FC1B2733" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[656,979,565,592]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="zebrinus">
|
||
<emphasis id="B907BE5B6931EF1FFD1BAD08FCDD272C" box="[656,789,565,592]" italics="true" pageId="23" pageNumber="24">A. zebrinus</emphasis>
|
||
(Pfeiffer, 1861)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7319CA6931EF1FFB86AD08FAE52733" authority="(Morlet, 1886)" baseAuthorityName="Morlet" baseAuthorityYear="1886" box="[1037,1325,565,592]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="begini">
|
||
<emphasis id="B907BE5B6931EF1FFB86AD08FBBF272C" box="[1037,1143,565,592]" italics="true" pageId="23" pageNumber="24">A. begini</emphasis>
|
||
(Morlet, 1886)
|
||
</taxonomicName>
|
||
by its having a smooth shell surface and brownish spiral Bands 4 and 5 present, with white lip. In contrast,
|
||
<taxonomicName id="4C7319CA6931EF1FFF36AD43FEF727F4" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[189,319,637,663]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="zebrinus">
|
||
<emphasis id="B907BE5B6931EF1FFF36AD43FEF727F4" box="[189,319,637,663]" italics="true" pageId="23" pageNumber="24">A. zebrinus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFE21FB86931EF1FFEF0AD43FDA127FB" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[379,617,637,664]" pageId="23" pageNumber="24" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
|
||
Sutcharit
|
||
<emphasis id="B907BE5B6931EF1FFE60AD40FDD527F4" box="[491,541,637,663]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 15k) has an elongate conical shell shape, Band 1 absent, Bands 2 to 5 merged and developing slanted radial streaks, as well as a reddish band between Bands 5 and 6.
|
||
<taxonomicName id="4C7319CA6931EF1FFF36ADFBFE6727BC" baseAuthorityName="Morlet" baseAuthorityYear="1886" box="[189,431,709,735]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="begini">
|
||
<emphasis id="B907BE5B6931EF1FFF36ADFBFE6727BC" box="[189,431,709,735]" italics="true" pageId="23" pageNumber="24">Amphidromus begini</emphasis>
|
||
</taxonomicName>
|
||
exhibits strong radial ridges, Bands 2 to 6 merged and developing slanted brownish radial streaks, and brownish ribs (
|
||
<figureCitation id="13487ECC6931EF1FFDC2ADD6FD7D2660" box="[585,693,744,771]" captionStart="Fig" captionStartId="21.[189,232,1752,1778]" captionTargetBox="[189,1399,539,1707]" captionTargetId="figure@21.[189,1399,539,1707]" captionTargetPageId="21" captionText="Fig. 10. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7022). D. Amphidromus begini (Morlet, 1886), syntype from Strung- Trang, Cambodia (MNHN-IM-2000-1832). E-G. Amphidromus flavus (Pfeiffer, 1861) from Tam Pou Kham, Vientiane, Laos (CUMZ 7029). H. Amphidromus flavus var. “indistinctus” Pilsbry, 1900, holotype (ANSP 31486). I–K. Amphidromus flavus var. “tryoni” Pilsbry, 1900. I. Lectotype (ANSP 31488). J. Paralectotype (ANSP 252745). K. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7026). L–M. Amphidromus semitessellatus (Morlet, 1884). L. Lectotype (MNHN-IM-2000-1985). M. Specimen from Cambodia (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3829764" httpUri="https://zenodo.org/record/3829764/files/figure.png" pageId="23" pageNumber="24">Fig. 10D</figureCitation>
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,
|
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<typeStatus id="54C8DCEB6931EF1FFD49ADD7FCD52660" box="[706,797,745,771]" pageId="23" pageNumber="24" type="syntype">syntype</typeStatus>
|
||
MNHN-IM-2000-1832).
|
||
</paragraph>
|
||
</subSubSection>
|
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</treatment>
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</document> |