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<document id="5B172EFE822693CCA9A309C63A6E6FAD" ID-DOI="10.11646/zootaxa.1478.1.6" ID-ISSN="1175­5334" ID-Zenodo-Dep="10088153" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625847271708" checkinUser="felipe" docAuthor="TISHECHKIN, ALEXEY K." docDate="2007" docId="03DA8783FF90FFB1FF1DFA9FFEB573EE" docLanguage="en" docName="zt01478p064.pdf.imf" docOrigin="Zootaxa 1478" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Ptomaphagus (Appadelopsis) merritti TISHECHKIN, 2007, sp. n." docType="treatment" docVersion="3" lastPageNumber="62" masterDocId="FFE3FFFBFF90FFB0FF8AFFCFFFE57407" masterDocTitle="A new species of Ptomaphagus (Appadelopsis) (Coleoptera: Leiodidae) from Great Smoky Mountains National Park, U. S. A." masterLastPageNumber="64" masterPageNumber="61" pageNumber="61" updateTime="1699509735627" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D23BF52724A5828AF26B125B4B93C994" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5D66233361312B2B36AC49BD5A9E6937">
<mods:title id="54B6F66E589C8DCA7E440FB6A06117E7">A new species of Ptomaphagus (Appadelopsis) (Coleoptera: Leiodidae) from Great Smoky Mountains National Park, U. S. A.</mods:title>
</mods:titleInfo>
<mods:name id="1AB7BE7CD26A22F74401A2C8BDA271E5" type="personal">
<mods:role id="2D1824754D0451A004EC91E688C545C4">
<mods:roleTerm id="C9D454236A93303DEE99897D0AD76144">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AE8946FB2B01AB648C6EDB0053A21E5B">TISHECHKIN, ALEXEY K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="AE0909BC2400872CADF40B7450C5D282">text</mods:typeOfResource>
<mods:relatedItem id="966A56BF7024341A22940ABDA005B98F" type="host">
<mods:titleInfo id="280548505E601B6551767292693F14A1">
<mods:title id="AC3937BE037CD9DE26495170A8B2FCEB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="10EB16E0AF53DBC4462692E7F695D96B">
<mods:date id="4C06B341EFEDEF4701EF0C11C52A7DCB">2007</mods:date>
<mods:detail id="307F5B3E3CC3AE9905CA902542BCB80C" type="pubDate">
<mods:number id="CC9A4EB99CC715B75792365A894279AF">2007-05-21</mods:number>
</mods:detail>
<mods:detail id="DF7C3E3599F9335AD07127A5898BD86C" type="volume">
<mods:number id="F23DDC779DA1A7ADDFF9DFF72C3AD9D0">1478</mods:number>
</mods:detail>
<mods:extent id="270D300A607E8EC5115BCC0676EB5C31" unit="page">
<mods:start id="DF1FE335727C87F97DA991CF204F76DF">61</mods:start>
<mods:end id="EE654A4EFDC0C67A10411F59E2145A80">64</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="32633971D1267AF444BED05C03E211FD">journal article</mods:classification>
<mods:identifier id="07840C066E7570FFD01098E70ED0836C" type="DOI">10.11646/zootaxa.1478.1.6</mods:identifier>
<mods:identifier id="46D871A050E56EA378B028195512B7E3" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="2C17AD22E0C7BF493BF46389FDBCC01A" type="Zenodo-Dep">10088153</mods:identifier>
<mods:identifier id="C24AF42F916056D0C0FD710390E58344" type="ZooBank">E96669B9-0038-4C00-8457-54D8B6281FF5</mods:identifier>
</mods:mods>
<treatment id="03DA8783FF90FFB1FF1DFA9FFEB573EE" LSID="urn:lsid:plazi:treatment:03DA8783FF90FFB1FF1DFA9FFEB573EE" httpUri="http://treatment.plazi.org/id/03DA8783FF90FFB1FF1DFA9FFEB573EE" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61">
<subSubSection id="C369651EFF90FFB0FF1DFA9FFD147160" box="[151,753,1359,1384]" pageId="0" pageNumber="61" type="nomenclature">
<paragraph id="8BCC3695FF90FFB0FF1DFA9FFD147160" blockId="0.[151,753,1359,1384]" box="[151,753,1359,1384]" pageId="0" pageNumber="61">
<heading id="D08481F9FF90FFB0FF1DFA9FFD147160" bold="true" box="[151,753,1359,1384]" fontSize="10" level="2" pageId="0" pageNumber="61" reason="2">
<emphasis id="B907EA87FF90FFB0FF1DFA9FFD147160" bold="true" box="[151,753,1359,1384]" pageId="0" pageNumber="61">
<taxonomicName id="4C734D16FF90FFB0FF1DFA9FFDD4716F" ID-CoL="78LPV" authorityName="Tishechkin" authorityYear="2007" box="[151,561,1359,1384]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="merritti" status="sp. nov." subGenus="Appadelopsis">
<emphasis id="B907EA87FF90FFB0FF1DFA9FFECB716F" bold="true" box="[151,302,1360,1384]" italics="true" pageId="0" pageNumber="61">Ptomaphagus</emphasis>
(
<emphasis id="B907EA87FF90FFB0FEB7FA9FFE2B716F" bold="true" box="[317,462,1360,1384]" italics="true" pageId="0" pageNumber="61">Appadelopsis</emphasis>
)
<emphasis id="B907EA87FF90FFB0FE54FA9FFDD4716F" bold="true" box="[478,561,1360,1384]" italics="true" pageId="0" pageNumber="61">merritti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457FCFF90FFB0FDB2FA80FD967160" box="[568,627,1359,1383]" pageId="0" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="13482A10FF90FFB0FD08FA80FD0C7160" box="[642,745,1359,1383]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2–6" captionStartId-0="1.[151,255,1080,1104]" captionStartId-1="1.[151,269,1724,1748]" captionTargetBox-0="[264,1323,386,1073]" captionTargetBox-1="[324,1245,1193,1701]" captionTargetId-0="figure-192@1.[264,1323,366,1090]" captionTargetId-1="figure-198@1.[701,1267,1193,1707]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="FIGURE 1. Ptomaphagus merritti sp. n. Dorsal habitus of the holotype." captionText-1="FIGURES 2–6. Genitalia of Ptomaphagus merritti sp. n. 2—aedeagus, lateral view; 3—apex of median lobe, frontal view; 4—apex of median lobe, dorso-lateral view; 5—spermatheca, dorsal view; 6—spermatheca, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.10088155" figureDoi-1="http://doi.org/10.5281/zenodo.10088157" httpUri-0="https://zenodo.org/record/10088155/files/figure.png" httpUri-1="https://zenodo.org/record/10088157/files/figure.png" pageId="0" pageNumber="61">Figs. 1–6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369651EFF90FFB0FF1DFA59FDF872AC" pageId="0" pageNumber="61" type="materials_examined">
<paragraph id="8BCC3695FF90FFB0FF1DFA59FDF872AC" blockId="0.[151,1436,1428,2020]" pageId="0" pageNumber="61">
Type material.
<materialsCitation id="3B1B3CC8FF90FFB0FEDBFA59FC8471F3" collectingDate="2006-04-14" collectionCode="FMNH" collectorName="A. Tishechkin" elevation="1370" latitude="35.526833" longLatPrecision="11" longitude="-83.819664" pageId="0" pageNumber="61" specimenCode="LSAM 0109735" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B907EA87FF90FFB0FEDBFA59FE2771A9" bold="true" box="[337,450,1430,1454]" pageId="0" pageNumber="61">
<typeStatus id="54C88837FF90FFB0FEDBFA59FE5871A9" box="[337,445,1430,1454]" pageId="0" pageNumber="61" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D75FD1CFF90FFB0FE45FA5BFE0671AF" box="[463,483,1428,1448]" pageId="0" pageNumber="61" type="male">♂</specimenCount>
, labeled: “N
<taxonomicName id="4C734D16FF90FFB0FD0CFA58FB2A71A9" authority=": Swain Co., GSMNP, Appalachian Tr." authorityName="Swain Co., GSMNP, Appalachian Tr." box="[646,1231,1431,1454]" class="Insecta" family="Scarabaeidae" genus="Carolina" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">CAROLINA: Swain Co., GSMNP, Appalachian Tr.</taxonomicName>
nr. Doe Knob at
<geoCoordinate id="EE475052FF90FFB0FF1DFA75FEF771D6" box="[151,274,1466,1489]" degrees="35" direction="north" minutes="31.61" orientation="latitude" pageId="0" pageNumber="61" precision="9" value="35.526833">35º31.61’N</geoCoordinate>
<geoCoordinate id="EE475052FF90FFB0FE92FA75FE7F71D6" box="[280,410,1466,1489]" degrees="83" direction="west" minutes="49.18" orientation="longitude" pageId="0" pageNumber="61" precision="9" value="-83.819664">83º49.18’W</geoCoordinate>
.
<quantity id="4C8B9B70FF90FFB0FE2EFA75FE1D71D6" box="[420,504,1466,1489]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="0" pageNumber="61" unit="m" value="1370.0">
<elevation id="005ED1A6FF90FFB0FE2EFA75FE1D71D6" box="[420,504,1466,1489]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="0" pageNumber="61" unit="m" value="1370.0">1370 m</elevation>
</quantity>
. Forest litter.
<date id="FFCD1055FF90FFB0FD00FA75FD1971D6" box="[650,764,1466,1489]" pageId="0" pageNumber="61" value="2006-04-14">
<collectingDate id="EF89E9BDFF90FFB0FD00FA75FD1971D6" box="[650,764,1466,1489]" pageId="0" pageNumber="61" value="2006-04-14">14.iv.2006</collectingDate>
</date>
,
<collectorName id="26865343FF90FFB0FC82FA75FC7871D6" box="[776,925,1466,1489]" pageId="0" pageNumber="61">A. Tishechkin</collectorName>
/
<specimenCode id="DBD59EEEFF90FFB0FC39FA75FB8471D6" box="[947,1121,1466,1489]" collectionCode="LSAM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="0" pageNumber="61">LSAM 0109735</specimenCode>
/
<typeStatus id="54C88837FF90FFB0FBFFFA75FAE771D6" box="[1141,1282,1466,1489]" pageId="0" pageNumber="61" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C734D16FF90FFB0FA83FA75FE7771F3" authorityName="Tishechkin" authorityYear="2007" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="merritti" status="sp. nov." subGenus="Appadelopsis">
<emphasis id="B907EA87FF90FFB0FA83FA75FA7971D6" box="[1289,1436,1466,1489]" italics="true" pageId="0" pageNumber="61">Ptomaphagus</emphasis>
(
<emphasis id="B907EA87FF90FFB0FF2AFA12FECA71F3" box="[160,303,1501,1524]" italics="true" pageId="0" pageNumber="61">Appadelopsis</emphasis>
)
<emphasis id="B907EA87FF90FFB0FECBFA12FE7771F3" box="[321,402,1501,1524]" italics="true" pageId="0" pageNumber="61">merritti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457FCFF90FFB0FE16FA12FE3271F3" box="[412,471,1501,1524]" pageId="0" pageNumber="61" rank="species">sp. n.</taxonomicNameLabel>
<collectorName id="26865343FF90FFB0FE6AFA12FD9D71F3" box="[480,632,1501,1524]" pageId="0" pageNumber="61">A. Tishechkin</collectorName>
des. 2007” (
<collectionCode id="ED62AE50FF90FFB0FC8DFA12FCBF71F3" box="[775,858,1501,1524]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="0" pageNumber="61" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1B3CC8FF90FFB0FCE4FA13FD95721F" collectingDate="2006-04-14" collectionCode="LSAM, GSMNP" collectorName="A. Tishechkin" country="United States of America" elevation="1370" latitude="35.526833" longLatPrecision="11" longitude="-83.819664" pageId="0" pageNumber="61" specimenCount="1" specimenCount-female="1" stateProvince="North Carolina" typeStatus="paratype">
<emphasis id="B907EA87FF90FFB0FCE4FA13FC0571F3" bold="true" box="[878,992,1500,1524]" pageId="0" pageNumber="61">
<typeStatus id="54C88837FF90FFB0FCE4FA13FC0571F3" box="[878,992,1500,1524]" pageId="0" pageNumber="61" type="paratype">Paratypes</typeStatus>
</emphasis>
(5):
<emphasis id="B907EA87FF90FFB0FB92FA13FB4171F3" bold="true" box="[1048,1188,1500,1524]" pageId="0" pageNumber="61">
<collectingCountry id="F3647605FF90FFB0FB92FA13FBB471F3" box="[1048,1105,1500,1524]" name="United States of America" pageId="0" pageNumber="61">USA</collectingCountry>
:
<collectingRegion id="49B7F877FF90FFB0FBEAFA13FB4171F3" box="[1120,1188,1500,1524]" country="United States of America" name="North Carolina" pageId="0" pageNumber="61">North</collectingRegion>
</emphasis>
<taxonomicName id="4C734D16FF90FFB0FB27FA13FA7D71F3" authority=": Swain Co." authorityName="Swain Co." box="[1197,1432,1500,1524]" class="Insecta" family="Scarabaeidae" genus="Carolina" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B907EA87FF90FFB0FB27FA13FAFF71F3" bold="true" box="[1197,1306,1500,1524]" pageId="0" pageNumber="61">Carolina:</emphasis>
Swain Co.
</taxonomicName>
: same data as holotype (
<specimenCount id="9D75FD1CFF90FFB0FE18FA30FE437214" box="[402,422,1535,1555]" pageId="0" pageNumber="61" type="female">♀</specimenCount>
,
<collectionCode id="ED62AE50FF90FFB0FE39F9CEFDE7721F" box="[435,514,1537,1560]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="0" pageNumber="61">LSAM</collectionCode>
);
<collectionCode id="ED62AE50FF90FFB0FD9EF9CEFD95721F" box="[532,624,1537,1560]" pageId="0" pageNumber="61">GSMNP</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B1B3CC8FF90FFB0FDF1F9CEFCD57239" collectingDate="2005-10-02" collectionCode="LSAM, GSMNP" collectorName="W. D. Merritt" country="United States of America" elevation="1495" latitude="35.630333" location="Hyatt Ridge Trs." longLatPrecision="11" longitude="-83.2345" municipality="Beech Gap" pageId="0" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
jct. of
<collectingMunicipality id="6BA8ACEFFF90FFB0FD4AF9CEFCD0721F" box="[704,821,1537,1560]" pageId="0" pageNumber="61">Beech Gap</collectingMunicipality>
and
<location id="8EAC604EFF90FFB0FCE0F9CEFBF9721F" LSID="urn:lsid:plazi:treatment:03DA8783FF90FFB1FF1DFA9FFEB573EE:8EAC604EFF90FFB0FCE0F9CEFBF9721F" box="[874,1052,1537,1560]" country="United States of America" latitude="35.630333" longLatPrecision="11" longitude="-83.2345" municipality="Beech Gap" name="Hyatt Ridge Trs." pageId="0" pageNumber="61" stateProvince="North Carolina">Hyatt Ridge Trs.</location>
<geoCoordinate id="EE475052FF90FFB0FBA9F9CEFB78721F" box="[1059,1181,1537,1560]" degrees="35" direction="north" minutes="37.82" orientation="latitude" pageId="0" pageNumber="61" precision="9" value="35.630333">35º37.82’N</geoCoordinate>
<geoCoordinate id="EE475052FF90FFB0FB2EF9CEFAC0721F" box="[1188,1317,1537,1560]" degrees="83" direction="west" minutes="14.07" orientation="longitude" pageId="0" pageNumber="61" precision="9" value="-83.2345">83º14.07’W</geoCoordinate>
,
<quantity id="4C8B9B70FF90FFB0FABAF9CEFA66721F" box="[1328,1411,1537,1560]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" pageId="0" pageNumber="61" unit="m" value="1495.0">
<elevation id="005ED1A6FF90FFB0FABAF9CEFA66721F" box="[1328,1411,1537,1560]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" pageId="0" pageNumber="61" unit="m" value="1495.0">1495 m</elevation>
</quantity>
,
<date id="FFCD1055FF90FFB0FA04F9CEFECE7239" pageId="0" pageNumber="61" value="2005-10-02">
<collectingDate id="EF89E9BDFF90FFB0FA04F9CEFECE7239" pageId="0" pageNumber="61" value="2005-10-02">2 October 2005</collectingDate>
</date>
,
<collectorName id="26865343FF90FFB0FEBCF9E8FE277239" box="[310,450,1575,1598]" pageId="0" pageNumber="61">W.D. Merritt</collectorName>
, forest litter (
<specimenCount id="9D75FD1CFF90FFB0FDD8F9EBFD83723F" box="[594,614,1572,1592]" pageId="0" pageNumber="61" type="male">♂</specimenCount>
,
<collectionCode id="ED62AE50FF90FFB0FDF9F9E8FD277239" box="[627,706,1575,1598]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="0" pageNumber="61">LSAM</collectionCode>
);
<collectionCode id="ED62AE50FF90FFB0FD5EF9E8FCD57239" box="[724,816,1575,1598]" pageId="0" pageNumber="61">GSMNP</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B1B3CC8FF90FFB0FCB0F9E8FCF37264" collectingDate="2002-07-22" collectionCode="GSMNP" collectorName="C. E. Carlton &amp; Berlese" country="United States of America" latitude="35.5995" location="upper Deep Creek Tr." longLatPrecision="11" longitude="-83.4245" pageId="0" pageNumber="61" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="North Carolina" typeStatus="paratype">
<location id="8EAC604EFF90FFB0FCB0F9E8FBC47239" LSID="urn:lsid:plazi:treatment:03DA8783FF90FFB1FF1DFA9FFEB573EE:8EAC604EFF90FFB0FCB0F9E8FBC47239" box="[826,1057,1575,1598]" country="United States of America" latitude="35.5995" longLatPrecision="11" longitude="-83.4245" name="upper Deep Creek Tr." pageId="0" pageNumber="61" stateProvince="North Carolina">upper Deep Creek Tr.</location>
at
<geoCoordinate id="EE475052FF90FFB0FBCBF9E8FB5E7239" box="[1089,1211,1575,1598]" degrees="35" direction="north" minutes="35.97" orientation="latitude" pageId="0" pageNumber="61" precision="9" value="35.5995">35º35.97’N</geoCoordinate>
<geoCoordinate id="EE475052FF90FFB0FB48F9E8FAA67239" box="[1218,1347,1575,1598]" degrees="83" direction="west" minutes="25.47" orientation="longitude" pageId="0" pageNumber="61" precision="9" value="-83.4245">83º25.47’W</geoCoordinate>
,
<date id="FFCD1055FF90FFB0FAC4F9E8FF2A7264" pageId="0" pageNumber="61" value="2002-07-22">
<collectingDate id="EF89E9BDFF90FFB0FAC4F9E8FF2A7264" pageId="0" pageNumber="61" value="2002-07-22">22 July 2002</collectingDate>
</date>
,
<collectorName id="26865343FF90FFB0FF50F983FE897264" box="[218,364,1612,1635]" pageId="0" pageNumber="61">C. E. Carlton</collectorName>
, moist upland
<collectorName id="26865343FF90FFB0FD80F983FDBF7264" box="[522,602,1612,1635]" pageId="0" pageNumber="61">Berlese</collectorName>
(
<specimenCount id="9D75FD1CFF90FFB0FDE1F985FD9A7259" box="[619,639,1610,1630]" pageId="0" pageNumber="61" type="male">♂</specimenCount>
,
<specimenCount id="9D75FD1CFF90FFB0FD07F985FD457259" box="[653,672,1610,1630]" pageId="0" pageNumber="61" type="female">♀</specimenCount>
,
<collectionCode id="ED62AE50FF90FFB0FD24F983FCEA7264" box="[686,783,1612,1635]" pageId="0" pageNumber="61">GSMNP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1B3CC8FF90FFB0FCA8F983FB657264" box="[802,1152,1612,1636]" collectionCode="GSMNP" country="United States of America" county="Sevier Co." location="Sevier Co." pageId="0" pageNumber="61" specimenCount="1" stateProvince="Tennessee" typeStatus="paratype">
<emphasis id="B907EA87FF90FFB0FCA8F983FC797263" bold="true" box="[802,924,1612,1636]" pageId="0" pageNumber="61">
<collectingRegion id="49B7F877FF90FFB0FCA8F983FC727263" box="[802,919,1612,1636]" country="United States of America" name="Tennessee" pageId="0" pageNumber="61">Tennessee</collectingRegion>
:
</emphasis>
<collectingCounty id="62AD4E19FF90FFB0FC2EF983FBFC7264" box="[932,1049,1612,1635]" pageId="0" pageNumber="61">Sevier Co.</collectingCounty>
:
<collectionCode id="ED62AE50FF90FFB0FBAEF983FB657264" box="[1060,1152,1612,1635]" pageId="0" pageNumber="61">GSMNP</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B1B3CC8FF90FFB0FB01F983FDFC72AC" collectingDate="2001-06-28" collectionCode="LSAM" collectorName="Moseley &amp; C. E. Carlton &amp; A. K. Tishechkin &amp; Berlese" country="United States of America" elevation="1750" latitude="35.61" location="Beech Gap on Clingmans Dome Rd." longLatPrecision="92" longitude="-83.44383" municipality="Appalachian Tr." pageId="0" pageNumber="61" specimenCount="1" specimenCount-male="1" stateProvince="Tennessee" typeStatus="paratype">
<collectingMunicipality id="6BA8ACEFFF90FFB0FB01F983FADD7264" box="[1163,1336,1612,1635]" pageId="0" pageNumber="61">Appalachian Tr.</collectingMunicipality>
at
<location id="8EAC604EFF90FFB0FAD3F983FE377280" LSID="urn:lsid:plazi:treatment:03DA8783FF90FFB1FF1DFA9FFEB573EE:8EAC604EFF90FFB0FAD3F983FE377280" country="United States of America" latitude="35.61" longLatPrecision="92" longitude="-83.44383" municipality="Appalachian Tr." name="Beech Gap on Clingmans Dome Rd." pageId="0" pageNumber="61" stateProvince="Tennessee">Beech Gap on Clingmans Dome Rd.</location>
<geoCoordinate id="EE475052FF90FFB0FE52F9BFFDA07280" box="[472,581,1648,1671]" degrees="35" direction="north" minutes="36.6" orientation="latitude" pageId="0" pageNumber="61" precision="92" value="35.61">35º36.6’N</geoCoordinate>
<geoCoordinate id="EE475052FF90FFB0FDC1F9BFFD297280" box="[587,716,1648,1671]" degrees="83" direction="west" minutes="26.63" orientation="longitude" pageId="0" pageNumber="61" precision="9" value="-83.44383">83º26.63’W</geoCoordinate>
,
<quantity id="4C8B9B70FF90FFB0FD5DF9BFFCCC7280" box="[727,809,1648,1671]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="0" pageNumber="61" unit="m" value="1750.0">
<elevation id="005ED1A6FF90FFB0FD5DF9BFFCCC7280" box="[727,809,1648,1671]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="0" pageNumber="61" unit="m" value="1750.0">1750 m</elevation>
</quantity>
,
<date id="FFCD1055FF90FFB0FCBEF9BFFC267280" box="[820,963,1648,1671]" pageId="0" pageNumber="61" value="2001-06-28">
<collectingDate id="EF89E9BDFF90FFB0FCBEF9BFFC267280" box="[820,963,1648,1671]" pageId="0" pageNumber="61" value="2001-06-28">28 June 2001</collectingDate>
</date>
, V.
<collectorName id="26865343FF90FFB0FC61F9BFFBAD7280" box="[1003,1096,1648,1671]" pageId="0" pageNumber="61">Moseley</collectorName>
,
<collectorName id="26865343FF90FFB0FBD8F9BFFB3B7280" box="[1106,1246,1648,1671]" pageId="0" pageNumber="61">C. E. Carlton</collectorName>
&amp;
<collectorName id="26865343FF90FFB0FB75F9BFFF5972AC" pageId="0" pageNumber="61">A. K. Tishechkin</collectorName>
, forest litter
<collectorName id="26865343FF90FFB0FEC9F95BFE7672AC" box="[323,403,1684,1707]" pageId="0" pageNumber="61">Berlese</collectorName>
(
<specimenCount id="9D75FD1CFF90FFB0FE28F95DFE5372A1" box="[418,438,1682,1702]" pageId="0" pageNumber="61" type="male">♂</specimenCount>
,
<collectionCode id="ED62AE50FF90FFB0FE49F95BFDF772AC" box="[451,530,1684,1707]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="0" pageNumber="61">LSAM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C369651EFF90FFB0FF4CF978FD3A737B" pageId="0" pageNumber="61" type="description">
<paragraph id="8BCC3695FF90FFB0FF4CF978FD3A737B" blockId="0.[151,1436,1428,2020]" pageId="0" pageNumber="61">
<emphasis id="B907EA87FF90FFB0FF4CF978FEB472C8" bold="true" box="[198,337,1719,1743]" inLineHeading="true" pageId="0" pageNumber="61" reason="1">Description.</emphasis>
A typical member of
<taxonomicName id="4C734D16FF90FFB0FDCBF977FC9872C8" authority="" authorityName="Gnaspini" authorityYear="1996" box="[577,893,1720,1743]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Appadelopsis">
<emphasis id="B907EA87FF90FFB0FDCBF977FD3672C8" box="[577,723,1720,1743]" italics="true" pageId="0" pageNumber="61">Ptomaphagus</emphasis>
(
<emphasis id="B907EA87FF90FFB0FD6EF977FC9672C8" box="[740,883,1720,1743]" italics="true" pageId="0" pageNumber="61">Appadelopsis</emphasis>
)
</taxonomicName>
(
<figureCitation id="13482A10FF90FFB0FC07F977FC2972C8" box="[909,972,1720,1743]" captionStart="FIGURE 1" captionStartId="1.[151,255,1080,1104]" captionTargetBox="[264,1323,386,1073]" captionTargetId="figure-192@1.[264,1323,366,1090]" captionTargetPageId="1" captionText="FIGURE 1. Ptomaphagus merritti sp. n. Dorsal habitus of the holotype." figureDoi="http://doi.org/10.5281/zenodo.10088155" httpUri="https://zenodo.org/record/10088155/files/figure.png" pageId="0" pageNumber="61">Fig. 1</figureCitation>
). Body length
<quantity id="4C8B9B70FF90FFB0FBFBF977FAEA72C8" box="[1137,1295,1720,1743]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.85" metricValueMin="2.75" pageId="0" pageNumber="61" unit="mm" value="2.8" valueMax="2.85" valueMin="2.75">2.75–2.85 mm</quantity>
, width
<quantity id="4C8B9B70FF90FFB0FAEAF977FF1972F6" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="0" pageNumber="61" unit="mm" value="1.375" valueMax="1.4" valueMin="1.35">1.35– 1.40 mm</quantity>
. Pronotum 1.72–1.78 times wider then long, 2.09–2.14 times shorter then elytra. Elytra together 1.23–1.29 times longer then wide. Aedeagus apex is similar to that of
<taxonomicName id="4C734D16FF90FFB0FC99F932FC5C7313" authority="(Peck)" baseAuthorityName="Peck" baseAuthorityYear="1979" box="[787,953,1789,1812]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="suteri">
<emphasis id="B907EA87FF90FFB0FC99F932FC8F7313" box="[787,874,1789,1812]" italics="true" pageId="0" pageNumber="61">P.suteri</emphasis>
(Peck)
</taxonomicName>
, with dorsal section upturned into long, wide knob with a distinct depression in the middle (
<figureCitation id="13482A10FF90FFB0FD0AF8EFFD027330" box="[640,743,1824,1847]" captionStart="FIGURES 2–6" captionStartId="1.[151,269,1724,1748]" captionTargetBox="[324,1245,1193,1701]" captionTargetId="figure-198@1.[701,1267,1193,1707]" captionTargetPageId="1" captionText="FIGURES 2–6. Genitalia of Ptomaphagus merritti sp. n. 2—aedeagus, lateral view; 3—apex of median lobe, frontal view; 4—apex of median lobe, dorso-lateral view; 5—spermatheca, dorsal view; 6—spermatheca, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10088157" httpUri="https://zenodo.org/record/10088157/files/figure.png" pageId="0" pageNumber="61">Figs. 2–4</figureCitation>
). Edges of both dorsal and ventral sections in frontal view deeply asymmetrically sinuate (
<figureCitation id="13482A10FF90FFB0FE17F88DFE3B735E" box="[413,478,1858,1881]" captionStart="FIGURES 2–6" captionStartId="1.[151,269,1724,1748]" captionTargetBox="[324,1245,1193,1701]" captionTargetId="figure-198@1.[701,1267,1193,1707]" captionTargetPageId="1" captionText="FIGURES 2–6. Genitalia of Ptomaphagus merritti sp. n. 2—aedeagus, lateral view; 3—apex of median lobe, frontal view; 4—apex of median lobe, dorso-lateral view; 5—spermatheca, dorsal view; 6—spermatheca, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10088157" httpUri="https://zenodo.org/record/10088157/files/figure.png" pageId="0" pageNumber="61">Fig. 3</figureCitation>
). Spermatheca (
<figureCitation id="13482A10FF90FFB0FD01F88DFD0A735E" box="[651,751,1858,1881]" captionStart="FIGURES 2–6" captionStartId="1.[151,269,1724,1748]" captionTargetBox="[324,1245,1193,1701]" captionTargetId="figure-198@1.[701,1267,1193,1707]" captionTargetPageId="1" captionText="FIGURES 2–6. Genitalia of Ptomaphagus merritti sp. n. 2—aedeagus, lateral view; 3—apex of median lobe, frontal view; 4—apex of median lobe, dorso-lateral view; 5—spermatheca, dorsal view; 6—spermatheca, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10088157" httpUri="https://zenodo.org/record/10088157/files/figure.png" pageId="0" pageNumber="61">Figs. 5–6</figureCitation>
) with sharp bend in posterior end, distinctly sinuate central section and anterior crest perpendicular to middle section.
</paragraph>
</subSubSection>
<subSubSection id="C369651EFF90FFB1FF4CF848FDF6753E" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61" type="diagnosis">
<paragraph id="8BCC3695FF90FFB1FF4CF848FDF6753E" blockId="0.[151,1436,1428,2020]" lastBlockId="1.[151,1436,152,313]" lastPageId="1" lastPageNumber="62" pageId="0" pageNumber="61">
<emphasis id="B907EA87FF90FFB0FF4CF848FEDE7398" bold="true" box="[198,315,1927,1951]" inLineHeading="true" pageId="0" pageNumber="61" reason="1">Diagnosis.</emphasis>
The genitalia, primarily the aedeagus, of
<taxonomicName id="4C734D16FF90FFB0FC9EF847FC997398" box="[788,892,1928,1951]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="merriti">
<emphasis id="B907EA87FF90FFB0FC9EF847FC997398" box="[788,892,1928,1951]" italics="true" pageId="0" pageNumber="61">P. merriti</emphasis>
</taxonomicName>
is similar to the genitalia of only one described
<taxonomicName id="4C734D16FF90FFB0FF1DF865FEC273C6" authorityName="Gnaspini" authorityYear="1996" box="[151,295,1962,1985]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="subGenus" subGenus="Appadelopsis">
<emphasis id="B907EA87FF90FFB0FF1DF865FEC273C6" box="[151,295,1962,1985]" italics="true" pageId="0" pageNumber="61">Appadelopsis</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C734D16FF90FFB0FE02F865FD4773C6" authority="(Peck, 1978)" baseAuthorityName="Peck" baseAuthorityYear="1979" box="[392,674,1962,1985]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="suteri" subGenus="Appadelopsis">
<emphasis id="B907EA87FF90FFB0FE02F865FE4573C6" box="[392,416,1962,1985]" italics="true" pageId="0" pageNumber="61">P.</emphasis>
(
<emphasis id="B907EA87FF90FFB0FE26F865FE2673C6" box="[428,451,1962,1985]" italics="true" pageId="0" pageNumber="61">A.</emphasis>
)
<emphasis id="B907EA87FF90FFB0FE59F865FDF573C6" box="[467,528,1962,1985]" italics="true" pageId="0" pageNumber="61">suteri</emphasis>
(
<bibRefCitation id="EFE24B64FF90FFB0FD94F865FD7F73C6" author="Peck, S. B." box="[542,666,1962,1985]" pageId="0" pageNumber="61" pagination="355 - 382" refId="ref2231" refString="Peck, S. B. (1978) Systematics and evolution of forest litter Adelopsis in the southern Appalachians (Coleoptera: Leiodidae: Catopinae). Psyche, 85 (4), 355 - 382." type="journal article" year="1978">Peck, 1978</bibRefCitation>
)
</taxonomicName>
. The sinuate margins of the dorsal and ventral sections of the aedeagus are unique to the new species, in
<taxonomicName id="4C734D16FF90FFB0FD85F802FD8C73E3" baseAuthorityName="Peck" baseAuthorityYear="1979" box="[527,617,1997,2020]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="suteri">
<emphasis id="B907EA87FF90FFB0FD85F802FD8C73E3" box="[527,617,1997,2020]" italics="true" pageId="0" pageNumber="61">P. suteri</emphasis>
</taxonomicName>
they are almost straight. The shape of the apical knob of the aedeagus is slightly different in these species. Spermatheca of
<taxonomicName id="4C734D16FF91FFB1FD3FFF57FCC474A8" authorityName="Tishechkin" authorityYear="2007" box="[693,801,152,175]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="merritti">
<emphasis id="B907EA87FF91FFB1FD3FFF57FCC474A8" box="[693,801,152,175]" italics="true" pageId="1" pageNumber="62">P. merritti</emphasis>
</taxonomicName>
has a distinctly sinuate central section and anterior section almost rectangularly attached to it, whereas in
<taxonomicName id="4C734D16FF91FFB1FD2BFF75FD1E74D6" baseAuthorityName="Peck" baseAuthorityYear="1979" box="[673,763,186,209]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="suteri">
<emphasis id="B907EA87FF91FFB1FD2BFF75FD1E74D6" box="[673,763,186,209]" italics="true" pageId="1" pageNumber="62">P. suteri</emphasis>
</taxonomicName>
the spermatheca has a straight middle section and the angle between it and apical section is acute. Also, these species have widely allopatric distributions, with
<taxonomicName id="4C734D16FF91FFB1FB37FF12FAF174F3" baseAuthorityName="Peck" baseAuthorityYear="1979" box="[1213,1300,221,244]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="suteri">
<emphasis id="B907EA87FF91FFB1FB37FF12FAF174F3" box="[1213,1300,221,244]" italics="true" pageId="1" pageNumber="62">P. suteri</emphasis>
</taxonomicName>
found in the southern Nantahala Mountains south of GSMNP, an area known for numerous narrow endemics among specialized litterdwelling beetles (
<bibRefCitation id="EFE24B64FF91FFB1FED9FEEDFE27753E" author="Barr, T. C." box="[339,450,290,313]" pageId="1" pageNumber="62" pagination="1 - 51" refId="ref1970" refString="Barr, T. C. (1974) The eyeless beetles of the genus Arianops Brendel (Coleoptera, Pselaphidae). Bulletin of the American Museum of Natural History, 154 (1), 1 - 51." type="journal article" year="1974">Barr 1974</bibRefCitation>
,
<bibRefCitation id="EFE24B64FF91FFB1FE47FEEDFDE2753E" author="Barr, T. C." box="[461,519,290,313]" pageId="1" pageNumber="62" pagination="29 - 75" refId="ref2011" refString="Barr, T. C. (1979) Revision of Appalachian Trechus (Coleoptera: Carabidae). Brimleyana, 2, 29 - 75." type="journal article" year="1979">1979</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0C661DFF91FFB1FF1DFBF7FC7E7057" ID-DOI="http://doi.org/10.5281/zenodo.10088155" ID-Zenodo-Dep="10088155" box="[151,923,1080,1104]" httpUri="https://zenodo.org/record/10088155/files/figure.png" pageId="1" pageNumber="62" startId="1.[151,255,1080,1104]" targetBox="[264,1323,386,1073]" targetPageId="1">
<paragraph id="8BCC3695FF91FFB1FF1DFBF7FC7E7057" blockId="1.[151,923,1080,1104]" box="[151,923,1080,1104]" pageId="1" pageNumber="62">
<emphasis id="B907EA87FF91FFB1FF1DFBF7FEFC7057" bold="true" box="[151,281,1080,1104]" pageId="1" pageNumber="62">FIGURE 1.</emphasis>
<taxonomicName id="4C734D16FF91FFB1FEAAFBF6FDEF7057" authorityName="Tishechkin" authorityYear="2007" box="[288,522,1081,1104]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="merritti" status="sp. nov.">
<emphasis id="B907EA87FF91FFB1FEAAFBF6FDEF7057" box="[288,522,1081,1104]" italics="true" pageId="1" pageNumber="62">Ptomaphagus merritti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457FCFF91FFB1FD9BFBF6FDAE7057" box="[529,587,1081,1104]" pageId="1" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
Dorsal habitus of the holotype.
</paragraph>
</caption>
<caption id="DF0C661DFF91FFB1FF1DF973FADD72F0" ID-DOI="http://doi.org/10.5281/zenodo.10088157" ID-Zenodo-Dep="10088157" httpUri="https://zenodo.org/record/10088157/files/figure.png" pageId="1" pageNumber="62" startId="1.[151,269,1724,1748]" targetBox="[324,1245,1193,1701]" targetPageId="1">
<paragraph id="8BCC3695FF91FFB1FF1DF973FADD72F0" blockId="1.[151,1436,1724,1783]" pageId="1" pageNumber="62">
<emphasis id="B907EA87FF91FFB1FF1DF973FEA072D3" bold="true" box="[151,325,1724,1748]" pageId="1" pageNumber="62">FIGURES 2–6.</emphasis>
Genitalia of
<taxonomicName id="4C734D16FF91FFB1FE52F972FD2072D3" authorityName="Tishechkin" authorityYear="2007" box="[472,709,1725,1748]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="merritti" status="sp. nov.">
<emphasis id="B907EA87FF91FFB1FE52F972FD2072D3" box="[472,709,1725,1748]" italics="true" pageId="1" pageNumber="62">Ptomaphagus merritti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23457FCFF91FFB1FD45F972FCEF72D3" box="[719,778,1725,1748]" pageId="1" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
2—aedeagus, lateral view; 3—apex of median lobe, frontal view; 4—apex of median lobe, dorso-lateral view; 5—spermatheca, dorsal view; 6—spermatheca, lateral view.
</paragraph>
</caption>
<subSubSection id="C369651EFF91FFB1FF4CF888FEB573EE" pageId="1" pageNumber="62" type="etymology">
<paragraph id="8BCC3695FF91FFB1FF4CF888FEB573EE" blockId="1.[151,1436,1863,2025]" pageId="1" pageNumber="62">
<emphasis id="B907EA87FF91FFB1FF4CF888FEA27358" bold="true" box="[198,327,1863,1887]" inLineHeading="true" pageId="1" pageNumber="62" reason="1">Etymology.</emphasis>
I dedicate this species to William D. Merritt, collector of one of the
<typeStatus id="54C88837FF91FFB1FB92F887FB647358" box="[1048,1153,1864,1887]" pageId="1" pageNumber="62" type="paratype">paratypes</typeStatus>
, in appreciation of his outstanding volunteer contribution to the GSMNP ATBI.
<emphasis id="B907EA87FF91FFB1FF4CF843FEC173A3" bold="true" box="[198,292,1932,1956]" inLineHeading="true" pageId="1" pageNumber="62" reason="1">Habitat.</emphasis>
Found in deep forest litter at
<quantity id="4C8B9B70FF91FFB1FDEBF842FD1173A3" box="[609,756,1933,1956]" metricMagnitude="3" metricUnit="m" metricValue="1.575" metricValueMax="1.75" metricValueMin="1.4" pageId="1" pageNumber="62" unit="m" value="1575.0" valueMax="1750.0" valueMin="1400.0">1400–1750 m</quantity>
of altitude.
<emphasis id="B907EA87FF91FFB1FF4CF860FEBD73C0" bold="true" box="[198,344,1967,1991]" inLineHeading="true" pageId="1" pageNumber="62" reason="1">Distribution.</emphasis>
Apparent endemic of the Great Smoky Mountains, known from scattered localities at high altitudes of GSMNP (
<figureCitation id="13482A10FF91FFB1FE8BF81DFEA673EE" box="[257,323,2002,2025]" captionStart="FIGURE 7" captionStartId="2.[151,255,687,711]" captionTargetBox="[264,1323,151,663]" captionTargetId="figure-577@2.[264,1323,147,663]" captionTargetPageId="2" captionText="FIGURE 7. Known distribution of Ptomaphagus merritti sp. n. (squares) and P.fumosus(Peck, 1978) (circles, filled—for historic records, open—for recent ATBI records) in the Great Smoky Mountains National Park." figureDoi="http://doi.org/10.5281/zenodo.10088159" httpUri="https://zenodo.org/record/10088159/files/figure.png" pageId="1" pageNumber="62">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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