256 lines
25 KiB
XML
256 lines
25 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.4.4.13" ID-GBIF-Dataset="6cd3d259-465d-4ab3-8a32-d57a812bb6e0" ID-ISSN="2624-2834" ID-Zenodo-Dep="5507925" ID-ZooBank="D1F5A853-32D1-470B-A609-D54F7B8C60CA" checkinTime="1631645857228" checkinUser="carolina" docAuthor="JIANG, RI-XIN, ZHANG, HONG-RUI, ELDREDGE, K. TARO, SONG, XIAO-BIN, LI, YAN-DA, TIHELKA, ERIK, HUANG, DI-YING, WANG, SHUO, ENGEL, MICHAEL S. & CAI, CHEN-YANG" docDate="2021" docId="03B28787AB480677FF0D8231FB3CFD76" docLanguage="en" docName="palaeoentomology.4.4.13.pdf" docOrigin="Palaeoentomology 4 (4)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.4.13" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Tanytermitidae Engel 2021, fam. nov." docType="treatment" docVersion="6" lastPageNumber="384" masterDocId="FF8BFFFFAB42067DFF858453FF89FB0A" masterDocTitle="Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)" masterLastPageNumber="389" masterPageNumber="374" pageNumber="384" updateTime="1661815541049" updateUser="admin">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Further evidence of Cretaceous termitophily: Description of new termite hosts of the trichopseniine Cretotrichopsenius (Coleoptera: Staphylinidae), with emendations to the classification of lower termites (Isoptera)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>JIANG, RI-XIN</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>ZHANG, HONG-RUI</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>ELDREDGE, K. TARO</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>SONG, XIAO-BIN</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>LI, YAN-DA</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>TIHELKA, ERIK</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>HUANG, DI-YING</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>WANG, SHUO</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>ENGEL, MICHAEL S.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>CAI, CHEN-YANG</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Palaeoentomology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-08-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>4</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>374</mods:start>
|
||
<mods:end>389</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/palaeoentomology.4.4.13</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/palaeoentomology.4.4.13</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">6cd3d259-465d-4ab3-8a32-d57a812bb6e0</mods:identifier>
|
||
<mods:identifier type="ISSN">2624-2834</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">5507925</mods:identifier>
|
||
<mods:identifier type="ZooBank">D1F5A853-32D1-470B-A609-D54F7B8C60CA</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5507953" ID-GBIF-Taxon="188191027" ID-Zenodo-Dep="5507953" LSID="urn:lsid:plazi:treatment:03B28787AB480677FF0D8231FB3CFD76" httpUri="http://treatment.plazi.org/id/03B28787AB480677FF0D8231FB3CFD76" lastPageNumber="384" pageId="10" pageNumber="384">
|
||
<subSubSection box="[136,519,1634,1660]" pageId="10" pageNumber="384" type="nomenclature">
|
||
<paragraph blockId="10.[136,519,1634,1660]" box="[136,519,1634,1660]" pageId="10" pageNumber="384">
|
||
<heading bold="true" box="[136,519,1634,1660]" fontSize="10" level="2" pageId="10" pageNumber="384" reason="3">
|
||
<emphasis bold="true" box="[136,519,1634,1660]" pageId="10" pageNumber="384">
|
||
<taxonomicName authority="Engel" authorityName="Engel" authorityYear="2021" box="[136,397,1634,1660]" class="Insecta" family="Tanytermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family" status="fam. nov.">Tanytermitidae Engel</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[410,519,1634,1660]" pageId="10" pageNumber="384" rank="family">fam. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="384" type="type_taxon">
|
||
<paragraph blockId="10.[136,778,1706,2093]" box="[136,625,1706,1733]" pageId="10" pageNumber="384">
|
||
<emphasis bold="true" box="[136,277,1706,1732]" pageId="10" pageNumber="384">
|
||
<typeStatus box="[136,195,1706,1732]" pageId="10" pageNumber="384">Type</typeStatus>
|
||
genus.
|
||
</emphasis>
|
||
<taxonomicName authority="Engel et al., 2007" authorityName="Engel" authorityYear="2007" box="[284,622,1706,1733]" class="Insecta" genus="Tanytermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[284,412,1707,1732]" italics="true" pageId="10" pageNumber="384">Tanytermes</emphasis>
|
||
<bibRefCitation author="Engel, M. S. & Grimaldi, D. A. & Krishna, K." box="[419,622,1706,1733]" pageId="10" pageNumber="384" pagination="1 - 32" refId="ref9937" refString="Engel, M. S., Grimaldi, D. A. & Krishna, K. (2007) Primitive termites from the Early Cretaceous of Asia (Isoptera). Stuttgarter Beitrage zur Naturkunde, Serie B, Geologie und Palaontologie, 371, 1 - 32." type="journal article" year="2007">
|
||
Engel
|
||
<emphasis box="[492,541,1706,1732]" italics="true" pageId="10" pageNumber="384">et al</emphasis>
|
||
., 2007
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="10.[136,778,1706,2093]" pageId="10" pageNumber="384">
|
||
<emphasis bold="true" box="[189,395,1742,1768]" pageId="10" pageNumber="384">Included genera.</emphasis>
|
||
Aside from the
|
||
<typeStatus box="[598,646,1743,1769]" pageId="10" pageNumber="384">type</typeStatus>
|
||
genus, the family includes the new genus
|
||
<taxonomicName authority="Engel & Cai" authorityName="Engel & Cai" authorityYear="2021" class="Insecta" family="Kalotermitidae" genus="Tanytermitalis" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[507,668,1778,1804]" italics="true" pageId="10" pageNumber="384">Tanytermitalis</emphasis>
|
||
Engel & Cai
|
||
</taxonomicName>
|
||
,
|
||
<emphasis bold="true" box="[189,294,1814,1840]" pageId="10" pageNumber="384">
|
||
<taxonomicNameLabel box="[189,294,1814,1840]" pageId="10" pageNumber="384" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<emphasis box="[310,418,1814,1840]" italics="true" pageId="10" pageNumber="384">vide infra</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="384" type="diagnosis">
|
||
<paragraph blockId="10.[136,778,1706,2093]" lastBlockId="10.[810,1452,158,1661]" pageId="10" pageNumber="384">
|
||
<emphasis bold="true" box="[189,312,1850,1876]" pageId="10" pageNumber="384">Diagnosis.</emphasis>
|
||
Alate with head elongate, much like the longer alate heads of some
|
||
<taxonomicName authority="Froggatt, 1897" authorityName="Froggatt" authorityYear="1897" class="Insecta" family="Rhinotermitidae" genus="Heterotermes" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[513,664,1887,1912]" italics="true" pageId="10" pageNumber="384">Heterotermes</emphasis>
|
||
<bibRefCitation author="Froggatt, W. W." pageId="10" pageNumber="384" pagination="510 - 552" refId="ref10182" refString="Froggatt, W. W. (1897) Australian Termitidae, Part II. Proceedings of the Linnean Society of New South Wales, 21 (4), 510 - 552, + pls. xxxv - xxxvi. https: // doi. org / 10.5962 / bhl. part. 8483" type="journal article" year="1897">Froggatt, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[209,221,1923,1948]" italics="true" pageId="10" pageNumber="384">e</emphasis>
|
||
.
|
||
<emphasis box="[229,243,1923,1948]" italics="true" pageId="10" pageNumber="384">g</emphasis>
|
||
.,
|
||
<taxonomicName authority="(Froggatt, 1898)" baseAuthorityName="Froggatt" baseAuthorityYear="1898" box="[265,738,1922,1948]" class="Insecta" family="Rhinotermitidae" genus="Heterotermes" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="species" species="paradoxus">
|
||
<emphasis box="[265,542,1922,1948]" italics="true" pageId="10" pageNumber="384">Heterotermes paradoxus</emphasis>
|
||
(
|
||
<bibRefCitation author="Froggatt, W. W." box="[558,729,1922,1948]" pageId="10" pageNumber="384" pagination="721 - 758" refId="ref10237" refString="Froggatt, W. W. (1898) Australian Termitidae, Part III. Proceedings of the Linnean Society of New South Wales, 22 (4), 721 - 758, + pls. xxxiv - xxxv. https: // doi. org / 10.5962 / bhl. part. 12741" type="journal article" year="1898">Froggatt, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hill, 1927" authorityName="Hill" authorityYear="1927" class="Insecta" family="Rhinotermitidae" genus="Heterotermes" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="species" species="occiduus">
|
||
<emphasis box="[750,770,1923,1948]" italics="true" pageId="10" pageNumber="384">H</emphasis>
|
||
.
|
||
<emphasis box="[136,236,1958,1984]" italics="true" pageId="10" pageNumber="384">occiduus</emphasis>
|
||
<bibRefCitation author="Hill, G. F." box="[247,368,1958,1984]" pageId="10" pageNumber="384" pagination="5 - 120" refId="ref10441" refString="Hill, G. F. (1927) Termites from the Australian region - Part I. Memoirs of the National Museum, Melbourne, 7 (1), 5 - 120, + 9 pls. https: // doi. org / 10.24199 / j. mmv. 1927.7.01" type="book chapter" year="1927">Hill, 1927</bibRefCitation>
|
||
</taxonomicName>
|
||
), sub-rectangular, fontanelle absent (present in Neoisoptera) (
|
||
<figureCitation box="[425,517,1994,2020]" captionStart="FIGURE 1" captionStartId="2.[136,235,1689,1713]" captionTargetBox="[183,1404,179,1665]" captionTargetId="figure-90@2.[183,1404,179,1665]" captionTargetPageId="2" captionText="FIGURE 1. Mid-Cretaceous termites with their putative rove beetle termitophiles. A, Arceotermes hospitis gen. et sp. nov. and Cretotrichopsenius burmiticus Cai et al., 2017a (QUST-INSECT-0015). B, Tanytermitalis philetaerus gen. et sp. nov. and C. burmiticus (NIGP174707). C, The entire amber piece of QUST-INSECT-0015. D, The entire amber piece of NIGP174707. Scale bars = 1 mm in A & C; 2 mm in B; 10 mm in D." figureDoi="http://doi.org/10.5281/zenodo.5507927" httpUri="https://zenodo.org/record/5507927/files/figure.png" pageId="10" pageNumber="384">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation box="[530,546,1995,2021]" captionStart="FIGURE 7" captionStartId="11.[136,235,1309,1333]" captionTargetBox="[151,1436,189,1286]" captionTargetId="figure-295@11.[151,1436,189,1286]" captionTargetPageId="11" captionText="FIGURE 7. Holotype alate of Tanytermitalis philetaerus Engel & Cai, gen. et sp. nov., alate (NIGP174707). A, Dorsal habitus. B, Detail of portion of right antenna as preserved. C, Dorsal view of head. D, Ventral view of head, green fluorescence light source. E, Right foreleg. F, Left midleg. G, Left hind leg. Scale bars = 1 mm in A–D; 400 μm in E–G." figureDoi="http://doi.org/10.5281/zenodo.5507943" httpUri="https://zenodo.org/record/5507943/files/figure.png" pageId="10" pageNumber="384">7</figureCitation>
|
||
), Y-shaped cleavage scar absent; compound eyes small, circular (
|
||
<figureCitation box="[669,764,2030,2056]" captionStart="FIGURE 7" captionStartId="11.[136,235,1309,1333]" captionTargetBox="[151,1436,189,1286]" captionTargetId="figure-295@11.[151,1436,189,1286]" captionTargetPageId="11" captionText="FIGURE 7. Holotype alate of Tanytermitalis philetaerus Engel & Cai, gen. et sp. nov., alate (NIGP174707). A, Dorsal habitus. B, Detail of portion of right antenna as preserved. C, Dorsal view of head. D, Ventral view of head, green fluorescence light source. E, Right foreleg. F, Left midleg. G, Left hind leg. Scale bars = 1 mm in A–D; 400 μm in E–G." figureDoi="http://doi.org/10.5281/zenodo.5507943" httpUri="https://zenodo.org/record/5507943/files/figure.png" pageId="10" pageNumber="384">Fig. 7C</figureCitation>
|
||
); ocelli absent (present in Icoisoptera); mandibles not extending beyond apex of labrum (
|
||
<figureCitation box="[1186,1273,158,185]" captionStart="FIGURE 7" captionStartId="11.[136,235,1309,1333]" captionTargetBox="[151,1436,189,1286]" captionTargetId="figure-295@11.[151,1436,189,1286]" captionTargetPageId="11" captionText="FIGURE 7. Holotype alate of Tanytermitalis philetaerus Engel & Cai, gen. et sp. nov., alate (NIGP174707). A, Dorsal habitus. B, Detail of portion of right antenna as preserved. C, Dorsal view of head. D, Ventral view of head, green fluorescence light source. E, Right foreleg. F, Left midleg. G, Left hind leg. Scale bars = 1 mm in A–D; 400 μm in E–G." figureDoi="http://doi.org/10.5281/zenodo.5507943" httpUri="https://zenodo.org/record/5507943/files/figure.png" pageId="10" pageNumber="384">Fig. 7D</figureCitation>
|
||
),right mandible with two sharp marginal teeth and elongate apical tooth, without subsidiary tooth (present in
|
||
<taxonomicName box="[1244,1451,230,256]" class="Insecta" family="Archotermopsidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Archotermopsidae</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Holmgren" authorityYear="1910" box="[880,1055,266,293]" class="Insecta" family="Stolotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Stolotermitidae</taxonomicName>
|
||
) [dentition known only for
|
||
<taxonomicName box="[810,938,303,328]" class="Insecta" genus="Tanytermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[810,938,303,328]" italics="true" pageId="10" pageNumber="384">Tanytermes</emphasis>
|
||
</taxonomicName>
|
||
, obscured in
|
||
<taxonomicName authorityName="Engel & Cai" authorityYear="2021" box="[1126,1287,302,328]" class="Insecta" family="Kalotermitidae" genus="Tanytermitalis" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1126,1287,302,328]" italics="true" pageId="10" pageNumber="384">Tanytermitalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis box="[1314,1434,302,328]" italics="true" pageId="10" pageNumber="384">vide infra</emphasis>
|
||
]; antenna with 14–18 articles (longer in
|
||
<taxonomicName authorityName="Desneux" authorityYear="1904" box="[1268,1452,338,364]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Mastotermitidae</taxonomicName>
|
||
and Teletisoptera, although some
|
||
<taxonomicName authorityName="Holmgren" authorityYear="1910" box="[1190,1363,374,400]" class="Insecta" family="Stolotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Stolotermitidae</taxonomicName>
|
||
with as few as 15 articles) (
|
||
<figureCitation box="[1026,1146,410,437]" captionStart="FIGURE 7" captionStartId="11.[136,235,1309,1333]" captionTargetBox="[151,1436,189,1286]" captionTargetId="figure-295@11.[151,1436,189,1286]" captionTargetPageId="11" captionText="FIGURE 7. Holotype alate of Tanytermitalis philetaerus Engel & Cai, gen. et sp. nov., alate (NIGP174707). A, Dorsal habitus. B, Detail of portion of right antenna as preserved. C, Dorsal view of head. D, Ventral view of head, green fluorescence light source. E, Right foreleg. F, Left midleg. G, Left hind leg. Scale bars = 1 mm in A–D; 400 μm in E–G." figureDoi="http://doi.org/10.5281/zenodo.5507943" httpUri="https://zenodo.org/record/5507943/files/figure.png" pageId="10" pageNumber="384">Fig. 7A, B</figureCitation>
|
||
). As in all termites with the exception of some basal groups like Melqartitermitidae and Mylacrotermitidae (
|
||
<emphasis box="[1112,1245,482,508]" italics="true" pageId="10" pageNumber="384">vide supra</emphasis>
|
||
), Tanytermitidae lack ventral cervical sclerites (observable only in
|
||
<taxonomicName authorityName="Engel & Cai" authorityYear="2021" box="[810,971,554,580]" class="Insecta" family="Kalotermitidae" genus="Tanytermitalis" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[810,971,554,580]" italics="true" pageId="10" pageNumber="384">Tanytermitalis</emphasis>
|
||
</taxonomicName>
|
||
). Pronotum flat (weakly saddle shaped in
|
||
<taxonomicName authorityName="Desneux" authorityYear="1904" box="[843,1024,590,617]" class="Insecta" family="Hodotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Hodotermitidae</taxonomicName>
|
||
), either slightly narrower or broader than head. Tibial spur formula 2–3–2 or 3–3–2 (3–3–
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="384" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName authorityName="Froggatt" authorityYear="1897" box="[844,1014,662,689]" class="Insecta" family="Kalotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Kalotermitidae</taxonomicName>
|
||
) (
|
||
<figureCitation box="[1042,1170,662,688]" captionStart="FIGURE 7" captionStartId="11.[136,235,1309,1333]" captionTargetBox="[151,1436,189,1286]" captionTargetId="figure-295@11.[151,1436,189,1286]" captionTargetPageId="11" captionText="FIGURE 7. Holotype alate of Tanytermitalis philetaerus Engel & Cai, gen. et sp. nov., alate (NIGP174707). A, Dorsal habitus. B, Detail of portion of right antenna as preserved. C, Dorsal view of head. D, Ventral view of head, green fluorescence light source. E, Right foreleg. F, Left midleg. G, Left hind leg. Scale bars = 1 mm in A–D; 400 μm in E–G." figureDoi="http://doi.org/10.5281/zenodo.5507943" httpUri="https://zenodo.org/record/5507943/files/figure.png" pageId="10" pageNumber="384">Fig. 7E–G</figureCitation>
|
||
); tarsi tetramerous (
|
||
<emphasis box="[1410,1418,663,688]" italics="true" pageId="10" pageNumber="384">i</emphasis>
|
||
.
|
||
<emphasis box="[1425,1437,663,688]" italics="true" pageId="10" pageNumber="384">e</emphasis>
|
||
., second tarsomere absent—second tarsomere becomes cryptic in some Teletisoptera and then wholly absent in other clades) (
|
||
<figureCitation box="[966,1088,770,796]" captionStart="FIGURE 7" captionStartId="11.[136,235,1309,1333]" captionTargetBox="[151,1436,189,1286]" captionTargetId="figure-295@11.[151,1436,189,1286]" captionTargetPageId="11" captionText="FIGURE 7. Holotype alate of Tanytermitalis philetaerus Engel & Cai, gen. et sp. nov., alate (NIGP174707). A, Dorsal habitus. B, Detail of portion of right antenna as preserved. C, Dorsal view of head. D, Ventral view of head, green fluorescence light source. E, Right foreleg. F, Left midleg. G, Left hind leg. Scale bars = 1 mm in A–D; 400 μm in E–G." figureDoi="http://doi.org/10.5281/zenodo.5507943" httpUri="https://zenodo.org/record/5507943/files/figure.png" pageId="10" pageNumber="384">Fig. 7E–G</figureCitation>
|
||
); pretarsal claws simple, arolium present or absent. Wing membranes not reticulate; Sc, R, and Rs heavily pigmented, remaining longitudinal sectors lightly pigmented; forewing scale large, overlapping hind wing scale (plesiomorphy relative to
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[1254,1381,914,941]" class="Insecta" family="Termitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Termitidae</taxonomicName>
|
||
); CuP (claval fissure) terminating on posterior wing margin prior to basal cleavage suture (
|
||
<emphasis box="[1153,1161,987,1012]" italics="true" pageId="10" pageNumber="384">i</emphasis>
|
||
.
|
||
<emphasis box="[1168,1180,987,1012]" italics="true" pageId="10" pageNumber="384">e</emphasis>
|
||
., inside of scale and not along suture—terminating on suture in many, but not all, basal Neoisoptera; reversed notably in
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[1273,1400,1058,1085]" class="Insecta" family="Termitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Termitidae</taxonomicName>
|
||
); Sc short, R1 simple, Rs running parallel to costal margin, Rs without apical parallel posterior inferiors (apical parallel posterior inferiors present in
|
||
<taxonomicName authorityName="Desneux" authorityYear="1904" box="[1132,1313,1166,1193]" class="Insecta" family="Hodotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Hodotermitidae</taxonomicName>
|
||
), radial field narrow, M running about halfway between Rs and CuA and sclerotized like CuA (rather than like Rs). Hind wing without anal lobe (present in
|
||
<taxonomicName authorityName="Desneux" authorityYear="1904" box="[1176,1365,1274,1301]" class="Insecta" family="Mastotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Mastotermitidae</taxonomicName>
|
||
). Cerci dimerous (observable only in
|
||
<taxonomicName authorityName="Engel & Cai" authorityYear="2021" box="[1167,1328,1310,1336]" class="Insecta" family="Kalotermitidae" genus="Tanytermitalis" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1167,1328,1310,1336]" italics="true" pageId="10" pageNumber="384">Tanytermitalis</emphasis>
|
||
</taxonomicName>
|
||
, unknown for
|
||
<taxonomicName box="[849,977,1347,1372]" class="Insecta" genus="Tanytermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[849,977,1347,1372]" italics="true" pageId="10" pageNumber="384">Tanytermes</emphasis>
|
||
</taxonomicName>
|
||
), as in
|
||
<taxonomicName authorityName="Froggatt" authorityYear="1897" box="[1060,1232,1346,1372]" class="Insecta" family="Kalotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Kalotermitidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="10.[810,1452,158,1661]" pageId="10" pageNumber="384">
|
||
Cerci and forewing venation somewhat similar to some
|
||
<taxonomicName authorityName="Froggatt" authorityYear="1897" box="[875,1046,1418,1444]" class="Insecta" family="Kalotermitidae" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="384" phylum="Arthropoda" rank="family">Kalotermitidae</taxonomicName>
|
||
, likely a reflection of the position of Tanytermitidae as closer to Icoisoptera than to more basal grade termites (
|
||
<bibRefCitation author="Engel, M. S. & Grimaldi, D. A. & Krishna, K." box="[985,1190,1490,1517]" pageId="10" pageNumber="384" pagination="1 - 27" refId="ref9988" refString="Engel, M. S., Grimaldi, D. A. & Krishna, K. (2009) Termites (Isoptera): their phylogeny, classification, and rise to ecological dominance. American Museum Novitates, 3650, 1 - 27. https: // doi. org / 10.1206 / 651.1" type="journal article" year="2009">
|
||
Engel
|
||
<emphasis box="[1059,1109,1490,1516]" italics="true" pageId="10" pageNumber="384">et al</emphasis>
|
||
., 2009
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Engel, M. S. & Barden, P. & Riccio, M. L. & Grimaldi, D. A." box="[1201,1261,1490,1517]" pageId="10" pageNumber="384" pagination="522 - 530" refId="ref10116" refString="Engel, M. S., Barden, P., Riccio, M. L. & Grimaldi, D. A. (2016) Morphologically specialized termite castes and advanced sociality in the Early Cretaceous. Current Biology, 26 (4), 522 - 530. https: // doi. org / 10.1016 / j. cub. 2015.12.061" type="journal article" year="2016">2016</bibRefCitation>
|
||
), but differing in the shorter CuA (subapical or apical termination in most Icoisoptera, terminating at apical third in Tanytermitidae). In addition, ocelli are present in imagoes of Icoisoptera, while lacking in the present family.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |