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<document id="5DB01D4EE735FE0CE02E19BF59E1E039" ID-DOI="10.5281/zenodo.191147" ID-GBIF-Dataset="e46b99bc-4894-4e3b-877e-2c06869dad00" ID-ISSN="1175-5326" ID-Zenodo-Dep="191147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461124342701" checkinUser="plazi" docAuthor="Blank, Miriam" docDate="2009" docId="03A8E167FFA10D6FFF51FB96F2C559D4" docLanguage="en" docName="zt02278p025.pdf" docOrigin="Zootaxa 2278" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Spiophanes Grube 1860" docType="treatment" docVersion="10" lastPageNumber="6" masterDocId="FF91991FFFA40D69FFC6FFF5F2155856" masterDocTitle="Spiophanes norrisi sp. nov. (Polychaeta: Spionidae) — a new species from the NE Pacific coast, separated from the Spiophanes bombyx complex based on both morphological and genetic studies" masterLastPageNumber="25" masterPageNumber="1" pageNumber="6" updateTime="1698596450614" updateUser="plazi">
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<mods:title id="24ACF541EF2CC042B799AA638B70D689">Spiophanes norrisi sp. nov. (Polychaeta: Spionidae) — a new species from the NE Pacific coast, separated from the Spiophanes bombyx complex based on both morphological and genetic studies</mods:title>
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<mods:date id="308CAE7C42D700B2CBC3D2AA29CE684E">2009</mods:date>
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<paragraph id="8BBE5071FFA10D6CFF51FB96F00E5C2B" blockId="5.[151,539,1123,1150]" box="[151,539,1123,1150]" pageId="5" pageNumber="6">
<heading id="D0F6E71DFFA10D6CFF51FB96F00E5C2B" bold="true" box="[151,539,1123,1150]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B9758C63FFA10D6CFF51FB96F00E5C2B" bold="true" box="[151,539,1123,1150]" pageId="5" pageNumber="6">
Genus
<taxonomicName id="4C012BF2FFA10D6CFF28FB91F00E5C2B" ID-CoL="84TRY" authority="Grube, 1860" authorityName="Grube" authorityYear="1860" box="[238,539,1123,1150]" class="Polychaeta" family="Spionidae" genus="Spiophanes" kingdom="Animalia" order="Spionida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="B9758C63FFA10D6CFF28FB91F36F5C28" bold="true" box="[238,378,1124,1150]" italics="true" pageId="5" pageNumber="6">Spiophanes</emphasis>
<bibRefCitation id="EF902D80FFA10D6CFE47FB96F00E5C2B" author="Grube" box="[385,539,1123,1149]" pageId="5" pageNumber="24" refString="Grube, A. E. (1860) Beschreibung neuer oder wenig bekannter Anneliden. Archiv der Naturgeschichte, 26, 71 - 118." type="journal article" year="1860">Grube, 1860</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31B03FAFFA10D6CFF51FB59F7585D5D" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BBE5071FFA10D6CFF51FB59F1315CB1" blockId="5.[151,1436,1196,1291]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB11775FFFA10D6CFF51FB59F1315CB1" infered="true" pageId="5" pageNumber="6">
<taxonomicName id="4C012BF2FFA10D6CFF51FB59F3D35C95" ID-CoL="84TRY" authority="Grube, 1860: 88" authorityName="Grube" authorityPageNumber="88" authorityYear="1860" box="[151,454,1196,1219]" class="Polychaeta" family="Spionidae" genus="Spiophanes" kingdom="Animalia" order="Spionida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="B9758C63FFA10D6CFF51FB59F3075C95" box="[151,274,1196,1219]" italics="true" pageId="5" pageNumber="6">Spiophanes</emphasis>
<treatmentCitation id="0AA07660FFA10D6CFEDEFB59F3D35C95" author="Grube" box="[280,454,1196,1219]" page="88" pageId="5" pageNumber="6" year="1860">
<bibRefCitation id="EF902D80FFA10D6CFEDEFB59F3B45C95" author="Grube" box="[280,417,1196,1219]" pageId="5" pageNumber="24" refString="Grube, A. E. (1860) Beschreibung neuer oder wenig bekannter Anneliden. Archiv der Naturgeschichte, 26, 71 - 118." type="journal article" year="1860">Grube, 1860</bibRefCitation>
: 88
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</taxonomicName>
89, pl. 5, fig. 1.
<typeStatus id="54BAEED3FFA10D6CFDBAFB59F0A45C95" box="[636,689,1196,1219]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="4C012BF2FFA10D6CFCD5FB59F67A5C95" ID-CoL="4Z69T" authority="Grube, 1860" authorityName="Grube" authorityYear="1860" box="[787,1135,1196,1219]" class="Polychaeta" family="Spionidae" genus="Spiophanes" kingdom="Animalia" order="Spionida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="kroyeri">
<emphasis id="B9758C63FFA10D6CFCD5FB59F63F5C95" box="[787,1066,1196,1219]" italics="true" pageId="5" pageNumber="6">
Spiophanes kroyeri
<bibRefCitation id="EF902D80FFA10D6CFC20FB59F63F5C95" author="Grube" box="[998,1066,1196,1219]" pageId="5" pageNumber="24" refString="Grube, A. E. (1860) Beschreibung neuer oder wenig bekannter Anneliden. Archiv der Naturgeschichte, 26, 71 - 118." type="journal article" year="1860">Grube</bibRefCitation>
</emphasis>
, 1860
</taxonomicName>
, by monotypy. Meißner &amp; Hutchings, 2003: 118120, figs. 12. Meißner, 2005: 6.
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</paragraph>
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<treatmentCitationGroup id="AB11775FFFA10D6CFF51FB01F7585D5D" box="[151,1357,1268,1291]" pageId="5" pageNumber="6">
<taxonomicName id="4C012BF2FFA10D6CFF51FB01F3F65D5D" ID-CoL="8dbdb866-d8b8-48e2-b4ed-16f697e732ef" authority="Chamberlin, 1919: 17" authorityName="Chamberlin" authorityPageNumber="17" authorityYear="1919" box="[151,483,1268,1291]" class="Polychaeta" family="Spionidae" genus="Morants" kingdom="Animalia" order="Spionida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="B9758C63FFA10D6CFF51FB01F2E45D5D" box="[151,241,1268,1291]" italics="true" pageId="5" pageNumber="6">Morants</emphasis>
<treatmentCitation id="0AA07660FFA10D6CFF3EFB01F3F65D5D" author="Chamberlin" box="[248,483,1268,1291]" page="17" pageId="5" pageNumber="6" year="1919">
<bibRefCitation id="EF902D80FFA10D6CFF3EFB01F3A95D5D" author="Chamberlin" box="[248,444,1268,1291]" pageId="5" pageNumber="24" refString="Chamberlin, R. V. (1919) New polychaetous annelids from Laguna Beach, California. Pomona College Journal of Entomology and Zoology, 11, 1 - 23." type="journal article" year="1919">Chamberlin, 1919</bibRefCitation>
: 17
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</taxonomicName>
.
<typeStatus id="54BAEED3FFA10D6CFE28FB01F0365D5D" box="[494,547,1268,1291]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="4C012BF2FFA10D6CFD43FB01F1E35D5D" ID-CoL="853JW" authority="Chamberlin, 1919" authorityName="Chamberlin" authorityYear="1919" box="[645,1014,1268,1291]" class="Polychaeta" family="Spionidae" genus="Morants" kingdom="Animalia" order="Spionida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="duplex">
<emphasis id="B9758C63FFA10D6CFD43FB01F1385D5D" box="[645,813,1268,1291]" italics="true" pageId="5" pageNumber="6">Morants duplex</emphasis>
<bibRefCitation id="EF902D80FFA10D6CFCF5FB01F1E35D5D" author="Chamberlin" box="[819,1014,1268,1291]" pageId="5" pageNumber="24" refString="Chamberlin, R. V. (1919) New polychaetous annelids from Laguna Beach, California. Pomona College Journal of Entomology and Zoology, 11, 1 - 23." type="journal article" year="1919">Chamberlin, 1919</bibRefCitation>
</taxonomicName>
, by monotypy. Junior synonym.
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</paragraph>
</subSubSection>
<subSubSection id="C31B03FAFFA10D6FFF51FACEF2C559D4" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BBE5071FFA10D6CFF51FACEF3F05E1E" blockId="5.[151,1437,1339,2024]" pageId="5" pageNumber="6">
<emphasis id="B9758C63FFA10D6CFF51FACEF3025D03" bold="true" box="[151,279,1339,1365]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Prostomium broad anteriorly; subtriangular, bell-shaped or rarely rounded; anterior margin often laterally extended, forming horns of different length; with or without occipital antenna. Eyes present or absent. Dorsal ciliated organs of different
<typeStatus id="54BAEED3FFA10D6CFD62FA74F0F05DCD" box="[676,741,1409,1435]" pageId="5" pageNumber="6">types</typeStatus>
present: (1) starting posterior to the prostomium as two continuous ciliated bands, differing in both length and also slightly in shape but maximally extending to chaetiger 17; (2) starting posterior to the prostomium as two short ciliated bands extending to the end of chaetiger 2, often followed by segmental ciliated patches of different shape; (3) starting posterior to the prostomium as pair of dorsal loops not extending beyond chaetiger 6; or (4) as ciliated grooves/short bands posterior to the prostomium.
</paragraph>
<paragraph id="8BBE5071FFA10D6CFF00F9A4F3E25ED8" blockId="5.[151,1437,1339,2024]" pageId="5" pageNumber="6">Peristomium moderately to well developed, forming lateral bulges or wings. Branchiae absent. Dorsal ciliated crests usually present.</paragraph>
<paragraph id="8BBE5071FFA10D6CFF00F963F6EE5F08" blockId="5.[151,1437,1339,2024]" pageId="5" pageNumber="6">Body divided into three distinct regions: (1) anterior region extending up to chaetiger 4, with parapodia 1 4 usually exhibiting well-developed neuro- and notopodial postchaetal lamellae compared to those of subsequent chaetigers, often positioned dorsally or dorsolaterally; (2) middle region extending from chaetiger 5 to last chaetiger bearing capillary chaetae rather than hooks in neuropodia, chaetigers usually with parapodial glandular organs (see Meißner &amp; Hutchings 2003); (3) posterior region indicated by presence of neuropodial hooks. Ventrolateral intersegmental pouches present or absent between neuropodia.</paragraph>
<paragraph id="8BBE5071FFA10D6FFF00F893F2C559D4" blockId="5.[151,1437,1339,2024]" lastBlockId="6.[151,1437,152,386]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">Notochaetae capillaries, often limbate or hirsute, in middle body region usually arranged in three rows. Neuropodia of chaetiger 1 with 12 conspicuous crook-like chaetae, other neurochaetae of chaetigers 14 simple, hirsute or limbate capillaries arranged in two rows. From chaetiger 5 neurochaetae short, broad, often sheathed and distally pointed; arranged in 12 indistinct rows. Neuropodial hooks first present from chaetiger 13 (in juvenile specimens of certain species), 14, 15, or 16; quadridentate, with main fang surmounted by single unpaired tooth and pair of smaller distal teeth, additional small teeth rarely present; reduced hood absent or present. Bacillary chaetae may be exposed from chaetigers 58. One or two ventral sabre chaetae often present from chaetiger 4, rarely from chaetigers 5, 9, 10, or 12, or sometimes not present until neuropodial hooks start; sabre chaetae accompanying hooks often with cryptic ridge (easy to observe under SEM). Stout, curved notochaetae often present in far posterior parapodia. Pygidium with two or more anal cirri.</paragraph>
</subSubSection>
</treatment>
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