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<document id="723E703D52C42EEB5BD186D7ECD848C5" ID-CLB-Dataset="21285" ID-DOI="10.1080/00222933.2015.1104393" ID-GBIF-Dataset="6d59feb3-23d2-4911-88b2-0db9163acf76" ID-ISSN="1464-5262" ID-Zenodo-Dep="3990373" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597772314732" checkinUser="carolina" docAuthor="Yamada, Kazutaka, Yasunaga, Tomohide &amp; Artchawakom, Taksin" docDate="2015" docId="EA37063720460D00414334D4FC04FB81" docLanguage="en" docName="JNATHIST.50.17-18.1103-1157.pdf.imf" docOrigin="Journal of Natural History 50" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Orius (Trichorius) crassus Yamada and Yasunaga, sp. nov." docType="treatment" docVersion="6" lastPageNumber="1135" masterDocId="160E7E4F20580D2040773102FFDBFE7C" masterDocTitle="The flower bug genus Orius Wolff, 1811 (Hemiptera: Heteroptera: Anthocoridae: Oriini) of Thailand" masterLastPageNumber="1157" masterPageNumber="1103" pageNumber="1133" updateTime="1698844749935" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="968D3B966397CD5CC72BA3D734ABFF17" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A3A0508D6ADDADDEB22A68368726177F">
<mods:title id="AEF0FD9CF769BF6D9653C2F0AF16BEBE">The flower bug genus Orius Wolff, 1811 (Hemiptera: Heteroptera: Anthocoridae: Oriini) of Thailand</mods:title>
</mods:titleInfo>
<mods:name id="79E9C56AE88DEEBDE9ED0435820ED2DC" type="personal">
<mods:role id="EC4B747C8FAB1204F897FE2B31FCAFFD">
<mods:roleTerm id="F43CB77F30EC9DC49F59CB4E3F9AF768">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A89214ECBBD6B82DC76B477D95E66D95">Yamada, Kazutaka</mods:namePart>
<mods:affiliation id="D11203104D6DA0E9F61F42AA6429301B">Tokushima Prefectural Museum, Bunka-no-Mori Park, Tokushima, Japan;</mods:affiliation>
</mods:name>
<mods:name id="E7C131ADAF238B63EB8376B1C13CF0C6" type="personal">
<mods:role id="3D8304C88ED7F9D814C153E2000A972B">
<mods:roleTerm id="2B97AC4E9D885FCA8A4BCA34E288A689">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="22BE3AB01B9252A56D5B4F25859F09DA">Yasunaga, Tomohide</mods:namePart>
<mods:affiliation id="F80D9A959A90D7A5BB5082D3F5C27441">American Museum of Natural History, New York, NY, USA; &amp; Plant Protection Division, Myanmar Ministry of Agriculture &amp; Irrigation, c / o Japan International Cooperation Agency (JICA), Yangon, Myanmar;</mods:affiliation>
</mods:name>
<mods:name id="06BD4A46E6416E5118EE00C1D91D5855" type="personal">
<mods:role id="9D7F518217014F1DAA9C1B5581B894F3">
<mods:roleTerm id="B51FFCBAC9BAF609CA0F7B0F06D75A89">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1D8487075A5D79434D08DCF6BA887809">Artchawakom, Taksin</mods:namePart>
<mods:affiliation id="2546024CC7495C2F1C1875C56D253306">Sakaerat Environmental Research Station (SERS), Thailand Institute of Scientific and Technological Research (TISTR), Ministry of Science and Technology, Nakhon Ratchasima, Thailand</mods:affiliation>
</mods:name>
<mods:typeOfResource id="3F847B51952E87580B84D1CF762F0A90">text</mods:typeOfResource>
<mods:relatedItem id="529C8DD1A44DCE2443C663AB66DB48E2" type="host">
<mods:titleInfo id="29F85AC2692EF35B0A4A5619A02713FB">
<mods:title id="FC15195B64256DD4664D9FB4C356C1A0">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="1567498D044046D7F113C95C19228F73">
<mods:date id="7FDDE5065B754C0C1BDB527A9917841B">2015</mods:date>
<mods:detail id="EE0AC9EE2B3976FD5932207A2FCE2AA0" type="pubDate">
<mods:number id="484300E8D9BBDC8FE29AF1213FFF0D79">2015-09-30</mods:number>
</mods:detail>
<mods:detail id="F9376E15D186AEF446AC59A41FB852FB" type="volume">
<mods:number id="7ED2F324A63BDCF0871B769B8010A119">50</mods:number>
</mods:detail>
<mods:extent id="D36D06A31A9DA18051431334433B7AA5" unit="page">
<mods:start id="FA9C4B21CD3BE733C739B85141CC9C8F">1103</mods:start>
<mods:end id="5C1FBED8E249BAD7626FC662E5877DC1">1157</mods:end>
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<mods:classification id="9EC1B2E97A9B292DB47277304D21F89E">journal article</mods:classification>
<mods:identifier id="6431D76428C3CBD22D5366E7E1B9514D" type="CLB-Dataset">21285</mods:identifier>
<mods:identifier id="9E214FA8DEB8F9A99AB50E1AE3F44038" type="DOI">10.1080/00222933.2015.1104393</mods:identifier>
<mods:identifier id="342AF0D4FFA5074AC5A268817C0C648A" type="GBIF-Dataset">6d59feb3-23d2-4911-88b2-0db9163acf76</mods:identifier>
<mods:identifier id="594C824F5AA744597099DEBC45328681" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="3F1216AAD4AF6ACDFA746B5622175DDD" type="Zenodo-Dep">3990373</mods:identifier>
</mods:mods>
<treatment id="EA37063720460D00414334D4FC04FB81" ID-DOI="http://doi.org/10.5281/zenodo.4332767" ID-GBIF-Taxon="166254623" ID-Zenodo-Dep="4332767" LSID="urn:lsid:plazi:treatment:EA37063720460D00414334D4FC04FB81" httpUri="http://treatment.plazi.org/id/EA37063720460D00414334D4FC04FB81" lastPageId="32" lastPageNumber="1135" pageId="30" pageNumber="1133">
<subSubSection id="2A84E4AA20460D3E414334D4FC2BFB93" box="[308,1008,1494,1519]" pageId="30" pageNumber="1133" type="nomenclature">
<paragraph id="6221B72120460D3E414334D4FC2BFB93" blockId="30.[308,1008,1494,1553]" box="[308,1008,1494,1519]" pageId="30" pageNumber="1133">
<heading id="3969004D20460D3E414334D4FC2BFB93" box="[308,1008,1494,1519]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="1133" reason="2">
<taxonomicName id="A59ECCA220460D3E414334D4FC5DFB93" authority="Yamada and Yasunaga" authorityName="Yamada and Yasunaga" box="[308,902,1494,1519]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="1133" phylum="Arthropoda" rank="species" species="crassus" status="sp. nov." subGenus="Trichorius">
<emphasis id="50EA6B3320460D3E414334D4FDB0FB93" bold="true" box="[308,619,1494,1519]" inLineHeading="true" pageId="30" pageNumber="1133" reason="3">
<emphasis id="50EA6B3320460D3E414334D4FEAEFB93" bold="true" box="[308,373,1494,1519]" italics="true" pageId="30" pageNumber="1133">Orius</emphasis>
(
<emphasis id="50EA6B3320460D3E41F034D4FDDBFB93" bold="true" box="[391,512,1494,1519]" italics="true" pageId="30" pageNumber="1133">Trichorius</emphasis>
)
<emphasis id="50EA6B3320460D3E426534D4FDB0FB93" bold="true" box="[530,619,1494,1519]" italics="true" pageId="30" pageNumber="1133">crassus</emphasis>
</emphasis>
Yamada and Yasunaga
</taxonomicName>
,
<taxonomicNameLabel id="4BD9D64820460D3E43E434D4FC2BFB93" box="[915,1008,1494,1519]" pageId="30" pageNumber="1133" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20460D3E41AE34FAFC91F86D" box="[473,842,1528,1553]" pageId="30" pageNumber="1133" type="description">
<paragraph id="6221B72120460D3E41AE34FAFC91F86D" blockId="30.[308,1008,1494,1553]" box="[473,842,1528,1553]" pageId="30" pageNumber="1133">
(
<figureCitation id="FAA5ABA420460D3E419634FAFDABF86D" box="[481,624,1528,1553]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="30" pageNumber="1133">Figures 3K,L</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E420834FAFD41F86D" box="[639,666,1528,1553]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="30" pageNumber="1133">5L</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E42DD34FAFD03F86D" box="[682,728,1528,1553]" captionStart="Figure 10" captionStartId="13.[160,225,1109,1132]" captionTargetBox="[183,1138,143,1074]" captionTargetId="figure@13.[178,1138,142,1078]" captionTargetPageId="13" captionText="Figure 10. Head and pronotum, male, dorsal view. (A) O. crassus, holotype; (B) O. maxidentex; (C) O. tantillus; (D) O. dravidiensis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990393" httpUri="https://zenodo.org/record/3990393/files/figure.png" pageId="30" pageNumber="1133">10A</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E429034FAFCCDF86D" box="[743,790,1528,1553]" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="30" pageNumber="1133">12D</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E435234FAFC99F86D" box="[805,834,1528,1553]" captionStart="Figure 17" captionStartId="39.[160,225,809,832]" captionTargetBox="[177,1135,146,778]" captionTargetId="figure@39.[177,1138,142,778]" captionTargetPageId="39" captionText="Figure 17. Paramere, three different views. (A–C), O. crassus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990409" httpUri="https://zenodo.org/record/3990409/files/figure.png" pageId="30" pageNumber="1133">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20460D3E40D7373CFD7CF8C3" pageId="30" pageNumber="1133" type="materials_examined">
<paragraph id="6221B72120460D3E40D7373CFE9EF82B" blockId="30.[160,1155,1598,1727]" box="[160,325,1598,1623]" pageId="30" pageNumber="1133">
<heading id="3969004D20460D3E40D7373CFE9EF82B" box="[160,325,1598,1623]" fontSize="11" level="3" pageId="30" pageNumber="1133" reason="5">
<emphasis id="50EA6B3320460D3E40D7373CFE9EF82B" bold="true" box="[160,325,1598,1623]" inLineHeading="true" italics="true" pageId="30" pageNumber="1133" reason="3">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120460D3E40D73763FD7CF8C3" blockId="30.[160,1155,1598,1727]" pageId="30" pageNumber="1133">
<materialsCitation id="D2F6BD7C20460D3E40D73763FD7CF8C3" ID-GBIF-Occurrence="2831862388" collectingDate="2010-03-17" collectingDateMax="2010-03-20" collectingDateMin="2010-03-17" collectionCode="DOA" collectorName="Sakaerat Environmental Research Station, N &amp; T. Yasunaga &amp; K. Yamada" country="Thailand" elevation="407" location="Sakaerat Environmental Research Station" pageId="30" pageNumber="1133" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima" typeStatus="holotype">
<emphasis id="50EA6B3320460D3E40D73763FECAF806" bold="true" box="[160,273,1633,1658]" inLineHeading="true" italics="true" pageId="30" pageNumber="1133" reason="3">
<typeStatus id="BD25098320460D3E40D73763FED0F806" box="[160,267,1633,1658]" pageId="30" pageNumber="1133" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="74987CA820460D3E41563760FEEEF806" box="[289,309,1634,1658]" pageId="30" pageNumber="1133" type="male">♂</specimenCount>
(
<figureCitation id="FAA5ABA420460D3E41323760FE0DF806" box="[325,470,1634,1658]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="30" pageNumber="1133">Figures 3K, L</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E41933760FE24F806" box="[484,511,1634,1658]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="30" pageNumber="1133">5L</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E427A3760FDE3F806" box="[525,568,1634,1658]" captionStart="Figure 10" captionStartId="13.[160,225,1109,1132]" captionTargetBox="[183,1138,143,1074]" captionTargetId="figure@13.[178,1138,142,1078]" captionTargetPageId="13" captionText="Figure 10. Head and pronotum, male, dorsal view. (A) O. crassus, holotype; (B) O. maxidentex; (C) O. tantillus; (D) O. dravidiensis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990393" httpUri="https://zenodo.org/record/3990393/files/figure.png" pageId="30" pageNumber="1133">10A</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E42303760FDAFF806" box="[583,628,1634,1658]" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="30" pageNumber="1133">12D</figureCitation>
,
<figureCitation id="FAA5ABA420460D3E42F53760FD46F806" box="[642,669,1634,1658]" captionStart="Figure 17" captionStartId="39.[160,225,809,832]" captionTargetBox="[177,1135,146,778]" captionTargetId="figure@39.[177,1138,142,778]" captionTargetPageId="39" captionText="Figure 17. Paramere, three different views. (A–C), O. crassus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990409" httpUri="https://zenodo.org/record/3990409/files/figure.png" pageId="30" pageNumber="1133">17</figureCitation>
),
<collectingCountry id="1A89F7B120460D3E42C43760FCF0F806" box="[691,811,1634,1658]" name="Thailand" pageId="30" pageNumber="1133">THAILAND</collectingCountry>
,
<collectingRegion id="A05A79C320460D3E43403760FBC3F806" box="[823,1048,1634,1658]" country="Thailand" name="Nakhon Ratchasima" pageId="30" pageNumber="1133">Nakhon Ratchasima</collectingRegion>
,
<collectorName id="CF6BD2F720460D3E44533760FDFBF8E1" pageId="30" pageNumber="1133">
<location id="6741E1FA20460D3E44533760FDDCF8E1" LSID="urn:lsid:plazi:treatment:EA37063720460D00414334D4FC04FB81:6741E1FA20460D3E44533760FDDCF8E1" country="Thailand" name="Sakaerat Environmental Research Station" pageId="30" pageNumber="1133" stateProvince="Nakhon Ratchasima">Sakaerat Environmental Research Station</location>
, N
</collectorName>
14°30ʹ26.9ʺ, E101°55ʹ39.2ʺ,
<quantity id="A5661AC420460D3E43263787FC4CF8E1" box="[849,919,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="4.07" pageId="30" pageNumber="1133" unit="m" value="407.0">
<elevation id="E9B3501220460D3E43263787FC4CF8E1" box="[849,919,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="4.07" pageId="30" pageNumber="1133" unit="m" value="407.0">407 m</elevation>
</quantity>
alt.,
<collectingMethod id="BBDFCF3620460D3E43BC3787FBEEF8E1" box="[971,1077,1669,1693]" pageId="30" pageNumber="1133">light trap</collectingMethod>
,
<date id="162091E120460D3E44483787FEF3F8C3" pageId="30" pageNumber="1133" value="2010-03-17" valueMax="2010-03-20" valueMin="2010-03-17">
<collectingDate id="0664680920460D3E44483787FEF3F8C3" pageId="30" pageNumber="1133" value="2010-03-17" valueMax="2010-03-20" valueMin="2010-03-17">
17
<emphasis id="50EA6B3320460D3E442D3786FBBCF8E0" box="[1114,1127,1668,1692]" italics="true" pageId="30" pageNumber="1133">–</emphasis>
20 March 2010
</collectingDate>
</date>
,
<collectorName id="CF6BD2F720460D3E414337A5FE60F8C3" box="[308,443,1703,1727]" pageId="30" pageNumber="1133">T. Yasunaga</collectorName>
&amp;
<collectorName id="CF6BD2F720460D3E41AA37A5FD8FF8C3" box="[477,596,1703,1727]" pageId="30" pageNumber="1133">K. Yamada</collectorName>
(
<collectionCode id="048F2FE420460D3E421437A5FD46F8C3" box="[611,669,1703,1727]" country="Thailand" httpUri="http://grbio.org/cool/v7nk-crsc" name="Department Of Agriculture, Thailand" pageId="30" pageNumber="1133">DOA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20470D3F40D7318CFC3DFFE6" pageId="31" pageNumber="1134" type="diagnosis">
<paragraph id="6221B72120470D3F40D7318CFECFFEDB" blockId="31.[160,1156,142,410]" box="[160,276,142,167]" pageId="31" pageNumber="1134">
<heading id="3969004D20470D3F40D7318CFECFFEDB" box="[160,276,142,167]" fontSize="11" level="3" pageId="31" pageNumber="1134" reason="5">
<emphasis id="50EA6B3320470D3F40D7318CFECFFEDB" bold="true" box="[160,276,142,167]" inLineHeading="true" italics="true" pageId="31" pageNumber="1134" reason="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120470D3F40D731B0FC3DFFE6" blockId="31.[160,1156,142,410]" pageId="31" pageNumber="1134">
Recognised by the following characters: body generally greyish brown (
<figureCitation id="FAA5ABA420470D3F439D31B0FBADFEB6" box="[1002,1142,178,202]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3K, L</figureCitation>
); hemelytra greyish brown, paler than pronotum and scutellum, cuneus somewhat darker than remainder of corium (
<figureCitation id="FAA5ABA420470D3F418831F5FDAFFF73" box="[511,628,247,271]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3K</figureCitation>
); antennal segment II thickened, fusiform (
<figureCitation id="FAA5ABA420470D3F40D03018FEC3FF4E" box="[167,280,282,306]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="31" pageNumber="1134">Figure 5L</figureCitation>
); legs pale yellow, each femur dark brown excluding pale yellow apex (
<figureCitation id="FAA5ABA420470D3F40D0303EFEC8FF28" box="[167,275,316,340]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3L</figureCitation>
); cone thin, pointed at apex in dorsal view, widened in lateral view, slightly elevated dorsally with weak projection (
<figureCitation id="FAA5ABA420470D3F4202305DFD39FF0B" box="[629,738,351,375]" captionStart="Figure 17" captionStartId="39.[160,225,809,832]" captionTargetBox="[177,1135,146,778]" captionTargetId="figure@39.[177,1138,142,778]" captionTargetPageId="39" captionText="Figure 17. Paramere, three different views. (A–C), O. crassus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990409" httpUri="https://zenodo.org/record/3990409/files/figure.png" pageId="31" pageNumber="1134">Figure 17</figureCitation>
); flagellum much longer than maximum width of cone, distinctly remote at base from the cone (
<figureCitation id="FAA5ABA420470D3F43193080FC02FFE6" box="[878,985,386,410]" captionStart="Figure 17" captionStartId="39.[160,225,809,832]" captionTargetBox="[177,1135,146,778]" captionTargetId="figure@39.[177,1138,142,778]" captionTargetPageId="39" captionText="Figure 17. Paramere, three different views. (A–C), O. crassus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990409" httpUri="https://zenodo.org/record/3990409/files/figure.png" pageId="31" pageNumber="1134">Figure 17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20470D0040D730C4FB5FFC9A" lastPageId="32" lastPageNumber="1135" pageId="31" pageNumber="1134" type="description">
<paragraph id="6221B72120470D3F40D730C4FEFDFFA3" blockId="31.[160,1156,454,1727]" box="[160,294,454,479]" pageId="31" pageNumber="1134">
<heading id="3969004D20470D3F40D730C4FEFDFFA3" box="[160,294,454,479]" fontSize="11" level="3" pageId="31" pageNumber="1134" reason="5">
<emphasis id="50EA6B3320470D3F40D730C4FEFDFFA3" bold="true" box="[160,294,454,479]" inLineHeading="true" italics="true" pageId="31" pageNumber="1134" reason="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120470D0040D730EBFCDFFFE5" blockId="31.[160,1156,454,1727]" lastBlockId="32.[160,1156,142,409]" lastPageId="32" lastPageNumber="1135" pageId="31" pageNumber="1134">
<emphasis id="50EA6B3320470D3F40D730EBFEB9FC7E" bold="true" box="[160,354,489,514]" inLineHeading="true" italics="true" pageId="31" pageNumber="1134" reason="3">
Male (
<typeStatus id="BD25098320470D3F409B30EBFE89FC7E" box="[236,338,489,514]" pageId="31" pageNumber="1134" type="holotype">holotype</typeStatus>
).
</emphasis>
Colouration: body generally greyish brown (
<figureCitation id="FAA5ABA420470D3F431D30E8FC2AFC7E" box="[874,1009,490,514]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3K, L</figureCitation>
). Head greyish brown, with paler apex; venter of head pale brown; eyes reddish brown, area surrounding ocellus red to reddish brown (
<figureCitation id="FAA5ABA420470D3F42C3332DFCFEFC3B" box="[692,805,559,583]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="31" pageNumber="1134">Figure 5L</figureCitation>
). Antennae yellowish brown; segment I dark brown; segments III and IV with dark brown tinge. Pronotum and scutellum greyish brown to blackish brown (
<figureCitation id="FAA5ABA420470D3F42B23377FCEEFCF1" box="[709,821,629,653]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="31" pageNumber="1134">Figure 5L</figureCitation>
). Hemelytra greyish brown, paler than pronotum and scutellum; cuneus somewhat darker than remainder of corium; membrane uniformly somber dark brown (
<figureCitation id="FAA5ABA420470D3F42F433B8FD36FCAE" box="[643,749,698,722]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3K</figureCitation>
). Legs pale yellow; each femur dark brown excluding pale yellow apex (
<figureCitation id="FAA5ABA420470D3F423833DEFD66FC88" box="[591,701,732,756]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3L</figureCitation>
). Venter of thorax and abdomen dull greyish brown to dark brown (
<figureCitation id="FAA5ABA420470D3F418F33FDFDBAFD6B" box="[504,609,767,791]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3L</figureCitation>
). Structure: body oval (
<figureCitation id="FAA5ABA420470D3F431033FDFC0AFD6B" box="[871,977,767,791]" captionStart="Figure 3" captionStartId="4.[160,225,1402,1425]" captionTargetBox="[178,1130,160,1368]" captionTargetId="figure@4.[178,1138,142,1371]" captionTargetPageId="4" captionText="Figure 3. Habitus. (A, B) O. filiferus, holotype, male, dorsal and lateral views; (C, D) same, paratype, female, dorsal and lateral views; (E, F) O. machaerus, holotype, male, dorsal and lateral views; (G, H) same, paratype, female, dorsal and lateral views; (I, J) O. inthanonus, holotype, male, dorsal and lateral views; (K, L) O. crassus, holotype, male, dorsal and lateral views. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3990379" httpUri="https://zenodo.org/record/3990379/files/figure.png" pageId="31" pageNumber="1134">Figure 3K</figureCitation>
). Head smooth, sparsely covered with short setae, 0.63 times as long as width across eyes, and with a longer erect seta on each side of clypeus, near anteromedial margin of each eye, and between eye and ocellus; ante-ocular portion 0.58 times as long as length of eye in dorsal view; vertex 1.9 times as wide as eye in dorsal view; eye proximate to anterior margin of pronotum, 1.3 times as long as eye width in dorsal view; neck indistinct (
<figureCitation id="FAA5ABA420470D3F40DF32CDFEF8FD9B" box="[168,291,975,999]" captionStart="Figure 10" captionStartId="13.[160,225,1109,1132]" captionTargetBox="[183,1138,143,1074]" captionTargetId="figure@13.[178,1138,142,1078]" captionTargetPageId="13" captionText="Figure 10. Head and pronotum, male, dorsal view. (A) O. crassus, holotype; (B) O. maxidentex; (C) O. tantillus; (D) O. dravidiensis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990393" httpUri="https://zenodo.org/record/3990393/files/figure.png" pageId="31" pageNumber="1134">Figure 10A</figureCitation>
). Antennal segment I stout, exceeding apex of head, sparsely covered with short suberect setae; segment II thickened, fusiform, 0.63 times as long as head width across eyes, densely covered with short suberect setae which are as long as width of the segment; segments III and IV slightly narrower than maximum width of segment II, covered with long erect setae intermixed with short decumbent setae, longest seta longer than width of respective segment; segment III slightly shorter than segment IV (
<figureCitation id="FAA5ABA420470D3F40DF359DFEF9FACB" box="[168,290,1183,1207]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="31" pageNumber="1134">Figures 5L</figureCitation>
,
<figureCitation id="FAA5ABA420470D3F4140359DFEB9FACB" box="[311,354,1183,1207]" captionStart="Figure 10" captionStartId="13.[160,225,1109,1132]" captionTargetBox="[183,1138,143,1074]" captionTargetId="figure@13.[178,1138,142,1078]" captionTargetPageId="13" captionText="Figure 10. Head and pronotum, male, dorsal view. (A) O. crassus, holotype; (B) O. maxidentex; (C) O. tantillus; (D) O. dravidiensis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990393" httpUri="https://zenodo.org/record/3990393/files/figure.png" pageId="31" pageNumber="1134">10A</figureCitation>
). Labium extending to the procoxae, sparsely covered with short suberect setae. Pronotum smooth, shiny, sparsely covered with long decumbent setae; anterior margin nearly straight, width as wide as mesal length; lateral margin nearly straight, slightly angulate at anterior corner; lateral carinae expanded anteriorly; posterior margin shallowly concave, width 2.4 times as wide as anterior pronotal width; collar distinct, strigose, with transverse weak groove, demarcated by deep impression with short setae and tiny punctures from callus; callus strongly convex, impunctate, with scattered long setae, demarcated posteriorly by deep transverse impression with short setae and tiny punctures (
<figureCitation id="FAA5ABA420470D3F41BC34B6FD9BFBB0" box="[459,576,1460,1484]" captionStart="Figure 5" captionStartId="6.[160,225,1375,1398]" captionTargetBox="[183,1133,144,1344]" captionTargetId="figure@6.[178,1138,142,1344]" captionTargetPageId="6" captionText="Figure 5. Head and pronotum, dorsal view. (A, B) O. sakaerat; (C, D) O. taksini; (E, F) O. tomokunii; (G, H) O. filiferus; (I, J) O. machaerus; (K), O. inthanonus; (L) O. crassus; (M, N) O. maxidentex; (O, P) O. tantillus; (Q, R) O. dravidiensis." figureDoi="http://doi.org/10.5281/zenodo.3990383" httpUri="https://zenodo.org/record/3990383/files/figure.png" pageId="31" pageNumber="1134">Figures 5L</figureCitation>
,
<figureCitation id="FAA5ABA420470D3F423934B6FDA1FBB0" box="[590,634,1460,1484]" captionStart="Figure 10" captionStartId="13.[160,225,1109,1132]" captionTargetBox="[183,1138,143,1074]" captionTargetId="figure@13.[178,1138,142,1078]" captionTargetPageId="13" captionText="Figure 10. Head and pronotum, male, dorsal view. (A) O. crassus, holotype; (B) O. maxidentex; (C) O. tantillus; (D) O. dravidiensis. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3990393" httpUri="https://zenodo.org/record/3990393/files/figure.png" pageId="31" pageNumber="1134">10A</figureCitation>
). Scutellum nearly equilateral, slightly shorter than basal width, deeply depressed and covered with long setae through middle. Hemelytra overall sparsely covered with long decumbent setae and tiny punctures; costal margin rounded; maximum width of endocorium 1.87 times as wide as embolium; embolial margin rounded; cuneal margin slightly curved, about 0.57 times as long as embolial margin; membrane with two visible veins, one located near costal margin and one near posterior margin of the membrane. Legs densely covered with decumbent setae; male protibiae with a row of about 17 small fuscous teeth on ventral side. Male genitalia (
<figureCitation id="FAA5ABA420780D004166318CFE41FEDA" box="[273,410,142,166]" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="32" pageNumber="1135">Figures 12D</figureCitation>
,
<figureCitation id="FAA5ABA420780D0041DE318CFE1FFEDA" box="[425,452,142,166]" captionStart="Figure 17" captionStartId="39.[160,225,809,832]" captionTargetBox="[177,1135,146,778]" captionTargetId="figure@39.[177,1138,142,778]" captionTargetPageId="39" captionText="Figure 17. Paramere, three different views. (A–C), O. crassus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990409" httpUri="https://zenodo.org/record/3990409/files/figure.png" pageId="32" pageNumber="1135">17</figureCitation>
): pygophore ovoid shaped, posteroventrally covered with 10 long, stout setae, of which the longest setae are about as long as half length of pygophore (
<figureCitation id="FAA5ABA420780D00415931D6FE76FE90" box="[302,429,212,236]" captionStart="Figure 12" captionStartId="24.[160,225,1563,1586]" captionTargetBox="[178,1132,142,1529]" captionTargetId="figure@24.[178,1138,142,1532]" captionTargetPageId="24" captionText="Figure 12. Pygophore with paramere, dorsal view. (A) O. filiferus, holotype; (B) O. machaerus, holotype; (C) O. inthanonus, holotype; (D) O. crassus, holotype. Abbreviations: eb = ejaculatory bulb; prm = paramere; pyg = pygophore. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3990397" httpUri="https://zenodo.org/record/3990397/files/figure.png" pageId="32" pageNumber="1135">Figure 12D</figureCitation>
); mediodorsal surface densely distributed with short, suberect setae; cone thin and pointed at apex in dorsal view, widened in lateral view, slightly elevated dorsally with weak projection (
<figureCitation id="FAA5ABA420780D00421D301BFD0EFF4D" box="[618,725,281,305]" captionStart="Figure 17" captionStartId="39.[160,225,809,832]" captionTargetBox="[177,1135,146,778]" captionTargetId="figure@39.[177,1138,142,778]" captionTargetPageId="39" captionText="Figure 17. Paramere, three different views. (A–C), O. crassus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990409" httpUri="https://zenodo.org/record/3990409/files/figure.png" pageId="32" pageNumber="1135">Figure 17</figureCitation>
); flagellum with outer shorter branch and inner longer branch, shorter branch slightly bent at apex, longer branch nearly straight in dorsal view and much longer than maximum width of cone, basal portion of flagellum distinctly remote from the cone (
<figureCitation id="FAA5ABA420780D0042FB3083FD2CFFE5" box="[652,759,385,409]" captionStart="Figure 17" captionStartId="39.[160,225,809,832]" captionTargetBox="[177,1135,146,778]" captionTargetId="figure@39.[177,1138,142,778]" captionTargetPageId="39" captionText="Figure 17. Paramere, three different views. (A–C), O. crassus, holotype. Abbreviations: cn = cone; flg = flagellum. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3990409" httpUri="https://zenodo.org/record/3990409/files/figure.png" pageId="32" pageNumber="1135">Figure 17</figureCitation>
).
</paragraph>
<paragraph id="6221B72120780D0040D730D2FEA4FF95" blockId="32.[160,383,464,489]" box="[160,383,464,489]" pageId="32" pageNumber="1135">
<emphasis id="50EA6B3320780D0040D730D2FF27FF95" bold="true" box="[160,252,464,489]" inLineHeading="true" italics="true" pageId="32" pageNumber="1135" reason="3">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="6221B72120780D0040D7331DFE4FFC44" blockId="32.[160,1156,543,742]" box="[160,404,543,568]" pageId="32" pageNumber="1135">
<heading id="3969004D20780D0040D7331DFE4FFC44" box="[160,404,543,568]" fontSize="11" level="3" pageId="32" pageNumber="1135" reason="5">
<emphasis id="50EA6B3320780D0040D7331DFE4FFC44" bold="true" box="[160,404,543,568]" inLineHeading="true" italics="true" pageId="32" pageNumber="1135" reason="3">Measurements (mm)</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120780D0040D73341FB5FFC9A" blockId="32.[160,1156,543,742]" pageId="32" pageNumber="1135">
[
<specimenCount id="74987CA820780D0040D03341FF60FC27" box="[167,187,579,603]" pageId="32" pageNumber="1135" type="male">♂</specimenCount>
(n = 1)]. Body length 1.93; head length (excluding neck) 0.24; head width across eyes 0.38; vertex width 0.19; width between ocelli 0.13; lengths of antennal segments I
<emphasis id="50EA6B3320780D0043933367FC2AFC01" box="[996,1009,613,637]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
II: I
<emphasis id="50EA6B3320780D0044613367FBF8FC01" box="[1046,1059,613,637]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
0.09, II
<emphasis id="50EA6B3320780D0044013367FB58FC01" box="[1142,1155,613,637]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
0.24, III
<emphasis id="50EA6B3320780D004086338AFF25FCDC" box="[241,254,648,672]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
0.17, IV
<emphasis id="50EA6B3320780D00412F338AFEBEFCDC" box="[344,357,648,672]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
0.19; lengths of labial segments II
<emphasis id="50EA6B3320780D0042A1338AFD38FCDC" box="[726,739,648,672]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
IV: II
<emphasis id="50EA6B3320780D004362338AFCF9FCDC" box="[789,802,648,672]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
0.06, III
<emphasis id="50EA6B3320780D00430D338AFC5CFCDC" box="[890,903,648,672]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
0.24, IV
<emphasis id="50EA6B3320780D004396338AFC35FCDC" box="[993,1006,648,672]" italics="true" pageId="32" pageNumber="1135">–</emphasis>
0.19; anterior pronotal width 0.33; mesal pronotal length 0.33; basal pronotal width 0.78; length of embolial margin 0.63; length of cuneal margin 0.36; maximum width across hemelytra 0.89.
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20780D0040D7321EFED7FD07" pageId="32" pageNumber="1135" type="etymology">
<paragraph id="6221B72120780D0040D7321EFEC6FD49" blockId="32.[160,1156,796,891]" box="[160,285,796,821]" pageId="32" pageNumber="1135">
<heading id="3969004D20780D0040D7321EFEC6FD49" box="[160,285,796,821]" fontSize="11" level="3" pageId="32" pageNumber="1135" reason="5">
<emphasis id="50EA6B3320780D0040D7321EFEC6FD49" bold="true" box="[160,285,796,821]" inLineHeading="true" italics="true" pageId="32" pageNumber="1135" reason="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120780D0040D73242FED7FD07" blockId="32.[160,1156,796,891]" pageId="32" pageNumber="1135">
From Latin,
<taxonomicName id="A59ECCA220780D00415B3242FEA1FD24" box="[300,378,832,856]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="50EA6B3320780D00415B3242FEA1FD24" box="[300,378,832,856]" italics="true" pageId="32" pageNumber="1135">crassus</emphasis>
</taxonomicName>
(= thick, stout), referring to thickened, fusiform male antennae; an adjective.
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20780D0040D732B0FD41FD91" box="[160,666,946,1005]" pageId="32" pageNumber="1135" type="distribution">
<paragraph id="6221B72120780D0040D732B0FEF7FDB7" blockId="32.[160,666,946,1005]" box="[160,300,946,971]" pageId="32" pageNumber="1135">
<heading id="3969004D20780D0040D732B0FEF7FDB7" box="[160,300,946,971]" fontSize="11" level="3" pageId="32" pageNumber="1135" reason="5">
<emphasis id="50EA6B3320780D0040D732B0FEF7FDB7" bold="true" box="[160,300,946,971]" inLineHeading="true" italics="true" pageId="32" pageNumber="1135" reason="3">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120780D0040D732D7FD41FD91" blockId="32.[160,666,946,1005]" box="[160,666,981,1005]" pageId="32" pageNumber="1135">
Northeastern
<collectingCountry id="1A89F7B120780D00414B32D7FE47FD91" box="[316,412,981,1005]" name="Thailand" pageId="32" pageNumber="1135">Thailand</collectingCountry>
(
<collectingRegion id="A05A79C320780D0041DC32D7FD4BFD91" box="[427,656,981,1005]" country="Thailand" name="Nakhon Ratchasima" pageId="32" pageNumber="1135">Nakhon Ratchasima</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="2A84E4AA20780D0040D73526FC04FB81" pageId="32" pageNumber="1135" type="discussion">
<paragraph id="6221B72120780D0040D73526FEDDFA41" blockId="32.[160,1156,1060,1431]" box="[160,262,1060,1085]" pageId="32" pageNumber="1135">
<heading id="3969004D20780D0040D73526FEDDFA41" box="[160,262,1060,1085]" fontSize="11" level="3" pageId="32" pageNumber="1135" reason="5">
<emphasis id="50EA6B3320780D0040D73526FEDDFA41" bold="true" box="[160,262,1060,1085]" inLineHeading="true" italics="true" pageId="32" pageNumber="1135" reason="3">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120780D0040D73545FE07FBEB" blockId="32.[160,1156,1060,1431]" pageId="32" pageNumber="1135">
<taxonomicName id="A59ECCA220780D0040D73545FEF5FA23" box="[160,302,1095,1119]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="50EA6B3320780D0040D73545FEF5FA23" box="[160,302,1095,1119]" italics="true" pageId="32" pageNumber="1135">Orius crassus</emphasis>
</taxonomicName>
evidently differs from
<taxonomicName id="A59ECCA220780D0042473545FC8EFA23" authority="Yasunaga, 1997" authorityName="Yasunaga" authorityYear="1997" box="[560,853,1095,1119]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D0042473545FD46FA23" box="[560,669,1095,1119]" italics="true" pageId="32" pageNumber="1135">O. atratus</emphasis>
Yasunaga, 1997
</taxonomicName>
from
<collectingCountry id="1A89F7B120780D0043ED3545FC07FA23" box="[922,988,1095,1119]" name="Japan" pageId="32" pageNumber="1135">Japan</collectingCountry>
and
<collectingCountry id="1A89F7B120780D0044613545FBBDFA23" box="[1046,1126,1095,1119]" name="Taiwan" pageId="32" pageNumber="1135">Taiwan</collectingCountry>
in having a generally greyish brown body (in
<taxonomicName id="A59ECCA220780D0042FA3568FD01FAFE" authorityName="Yasunaga" authorityYear="1997" box="[653,730,1130,1154]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D0042FA3568FD01FAFE" box="[653,730,1130,1154]" italics="true" pageId="32" pageNumber="1135">atratus</emphasis>
</taxonomicName>
, shiny blackish), the hemelytra being paler than pronotum and scutellum with somewhat darker cuneus (in
<taxonomicName id="A59ECCA220780D0043C5358FFC24FAD9" authorityName="Yasunaga" authorityYear="1997" box="[946,1023,1165,1189]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D0043C5358FFC24FAD9" box="[946,1023,1165,1189]" italics="true" pageId="32" pageNumber="1135">atratus</emphasis>
</taxonomicName>
, shiny dark, chestnut brown to more blackish), the much thicker male antennae (in
<taxonomicName id="A59ECCA220780D00438935ADFB90FABB" authorityName="Yasunaga" authorityYear="1997" box="[1022,1099,1199,1223]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D00438935ADFB90FABB" box="[1022,1099,1199,1223]" italics="true" pageId="32" pageNumber="1135">atratus</emphasis>
</taxonomicName>
, not strongly thickened in male), and the labium being extending to the procoxae (in
<taxonomicName id="A59ECCA220780D00444635D0FBA5FA96" authorityName="Yasunaga" authorityYear="1997" box="[1073,1150,1234,1258]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D00444635D0FBA5FA96" box="[1073,1150,1234,1258]" italics="true" pageId="32" pageNumber="1135">atratus</emphasis>
</taxonomicName>
, reaching the mesocoxae). In the male genitalia, however, there are few differences between
<taxonomicName id="A59ECCA220780D0041793415FE87FB53" box="[270,348,1303,1327]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="50EA6B3320780D0041793415FE87FB53" box="[270,348,1303,1327]" italics="true" pageId="32" pageNumber="1135">crassus</emphasis>
</taxonomicName>
and
<taxonomicName id="A59ECCA220780D0041EB3415FE32FB53" authorityName="Yasunaga" authorityYear="1997" box="[412,489,1303,1327]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D0041EB3415FE32FB53" box="[412,489,1303,1327]" italics="true" pageId="32" pageNumber="1135">atratus</emphasis>
</taxonomicName>
. The former can be distinguished from the latter by the flagellum with nearly straight inner longer branch (in
<taxonomicName id="A59ECCA220780D0043573438FCB6FB2E" authorityName="Yasunaga" authorityYear="1997" box="[800,877,1338,1362]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D0043573438FCB6FB2E" box="[800,877,1338,1362]" italics="true" pageId="32" pageNumber="1135">atratus</emphasis>
</taxonomicName>
, sinuate) and the basal portion of flagellum being distinctly remote at base from the cone (in
<taxonomicName id="A59ECCA220780D004394345FFBEBFB09" authorityName="Yasunaga" authorityYear="1997" box="[995,1072,1373,1397]" class="Insecta" family="Anthocoridae" genus="Orius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="1135" phylum="Arthropoda" rank="species" species="atratus">
<emphasis id="50EA6B3320780D004394345FFBEBFB09" box="[995,1072,1373,1397]" italics="true" pageId="32" pageNumber="1135">atratus</emphasis>
</taxonomicName>
, rather proximate to base of cone).
</paragraph>
<paragraph id="6221B72120780D0040D734C3FF22FBA6" blockId="32.[160,991,1473,1533]" box="[160,249,1473,1498]" pageId="32" pageNumber="1135">
<heading id="3969004D20780D0040D734C3FF22FBA6" box="[160,249,1473,1498]" fontSize="11" level="3" pageId="32" pageNumber="1135" reason="5">
<emphasis id="50EA6B3320780D0040D734C3FF22FBA6" bold="true" box="[160,249,1473,1498]" inLineHeading="true" italics="true" pageId="32" pageNumber="1135" reason="3">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="6221B72120780D0040D734E7FC04FB81" blockId="32.[160,991,1473,1533]" box="[160,991,1509,1533]" pageId="32" pageNumber="1135">Unknown. A single specimen was collected with an ultraviolet light trap.</paragraph>
</subSubSection>
</treatment>
</document>
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