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<mods:title id="6F7A7FD4DE81C007DC31DAEA5EA4E7AB">Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex</mods:title>
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<treatment id="D47D1EAAA795C36E636E91F944D0EE3E" ID-GBIF-Taxon="154365975" LSID="urn:lsid:plazi:treatment:D47D1EAAA795C36E636E91F944D0EE3E" httpUri="http://treatment.plazi.org/id/D47D1EAAA795C36E636E91F944D0EE3E" lastPageNumber="177" pageId="0" pageNumber="177">
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<paragraph id="498B91CB7BC484D7121A022AD7F0A1D6" pageId="0" pageNumber="177">
Genus
<taxonomicName id="31B6306DDFC564D928D60AB38994B0F8" ID-CoL="8KWY8" authority="Sharp" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax Sharp</taxonomicName>
</paragraph>
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<subSubSection id="3D7D0EFD89F7DC6E34AB0DB50F01D174" pageId="0" pageNumber="177" type="reference_group">
<paragraph id="E03DB2219FD3C30345734CF101D0725D" pageId="0" pageNumber="177">
<taxonomicName id="0FD82D14262FD0C8C57EC69BE6A556F3" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
Sharp, 1903: 243 (type species
<taxonomicName id="A16F81213BEE0ADB923A3CA54B15ADF6" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax montivagus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="montivagus">Cyclothorax montivagus</taxonomicName>
Blackburn by
<bibRefCitation id="518268D222E6EE8438A40A6C0C904AF0" author="Andrewes, HE" journalOrPublisher=")" pageId="0" pageNumber="177" pagination="128 - 139" title="Papers on Oriental Carabidae. - XXXV. On the types of some Indian genera. Annals and Magazine of Natural History (11 th ser." volume="3" year="1939">Andrewes 1939</bibRefCitation>
).
</paragraph>
<paragraph id="F044E3FF0A91FFACEEEE46793CFD1C4A" pageId="0" pageNumber="177">
<taxonomicName id="D88EEF459367D6732274B276553A11F2" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Cyclothorax</taxonomicName>
MacLeay, 1871: 104 (not
<taxonomicName id="B47091658C562AE9CA92990C1BC5FBF8" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Cyclothorax</taxonomicName>
Frauenfeld, 1868; type species
<taxonomicName id="9F36CB87561CBD78A1F64A8E777848EF" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax punctipennis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="punctipennis">Cyclothorax punctipennis</taxonomicName>
MacLeay by monotypy; synonymy
<bibRefCitation id="78BBF51ECB401E965982DEB2144EE64B" author="Sloane, TG" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="177" pagination="566 - 642" title="Studies in Australian Entomology. no. XII. New Carabidae (Panageini, Bembidiini, Pogonini, Platysmatini, Platynini, Lebiini, with revisional lists of genera and species, some notes on synonymy, &amp; c.)." volume="28" year="1903">Sloane 1903</bibRefCitation>
).
</paragraph>
<paragraph id="9BA377224E459AFE6F5A7C24CCDEBAF1" pageId="0" pageNumber="177">
<taxonomicName id="5B6F7C6A9125FB1ECABE81D16A0FBDA2" class="Insecta" family="Carabidae" genus="Thriscothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thriscothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Thriscothorax</taxonomicName>
Sharp, 1903: 257 (type species
<taxonomicName id="E07814CC6E99776F049019BE37DA93BE" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax unctus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="unctus">Cyclothorax unctus</taxonomicName>
Blackburn by original designation; synonymy
<bibRefCitation id="3343E87BF3F3ABDE9A28CAD8D1DE267A" author="Britton, EB" journalOrPublisher="Bishop Museum" pageId="0" pageNumber="177" pagination="107 - 166" title="A revision of the Hawaiian species of Mecyclothorax (Coleoptera: Carabidae). Occasional Papers of the Bernice P." volume="19" year="1948">Britton 1948</bibRefCitation>
).
</paragraph>
<paragraph id="75D1928C5FBAA6AC1F9EE37AAC02F911" pageId="0" pageNumber="177">
<taxonomicName id="AC5E915F84BBE684571BB6DD2933DCA3" class="Insecta" family="Carabidae" genus="Atelothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Atelothorax</taxonomicName>
Sharp, 1903: 269 (type species
<taxonomicName id="6949D1E377AD642114C0BEB2D6922A95" class="Insecta" family="Carabidae" genus="Atelothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atelothorax optatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="optatus">Atelothorax optatus</taxonomicName>
Sharp by monotypy; synonymy
<bibRefCitation id="F62860EFFACF1063EE98053772A7FFBB" author="Britton, EB" journalOrPublisher="Bishop Museum" pageId="0" pageNumber="177" pagination="107 - 166" title="A revision of the Hawaiian species of Mecyclothorax (Coleoptera: Carabidae). Occasional Papers of the Bernice P." volume="19" year="1948">Britton 1948</bibRefCitation>
).
</paragraph>
<paragraph id="502644BA7DE1A2085D22E0FD43E224DC" pageId="0" pageNumber="177">
<taxonomicName id="138AB1AF260949E28809F99C5BFEFD2C" class="Insecta" family="Carabidae" genus="Metrothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Metrothorax</taxonomicName>
Sharp, 1903: 269 (type species
<taxonomicName id="4F74E161B5868AA1CFDF329DF52F1710" class="Insecta" family="Carabidae" genus="Metrothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrothorax molops" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="molops">Metrothorax molops</taxonomicName>
Sharp by
<bibRefCitation id="E86469438BBEF854AC8C09E7570CBF86" author="Lorenz, W" journalOrPublisher="Published by the author, Tutzing" pageId="0" pageNumber="177" title="Nomina Carabidarum, A Directory of the Scientific Names of Ground Beetles." year="1998">Lorenz 1998</bibRefCitation>
; synonymy
<bibRefCitation id="06838E039BC1ED16151128E52F3B73A2" author="Britton, EB" journalOrPublisher="Bishop Museum" pageId="0" pageNumber="177" pagination="107 - 166" title="A revision of the Hawaiian species of Mecyclothorax (Coleoptera: Carabidae). Occasional Papers of the Bernice P." volume="19" year="1948">Britton 1948</bibRefCitation>
).
</paragraph>
<paragraph id="6E0D70DC6E27E641C25D7954C2C3CE47" pageId="0" pageNumber="177">
<taxonomicName id="72FE951F2E8A26A1DD220599A4F7A220" class="Insecta" family="Carabidae" genus="Antagonaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antagonaspis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Antagonaspis</taxonomicName>
Enderlein, 1909: 488 (type species
<taxonomicName id="0FE2F3D80C1E6CA64AB5809CBD4438C6" class="Insecta" family="Carabidae" genus="Antagonaspis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antagonaspis sculptopunctata" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="sculptopunctata">Antagonaspis sculptopunctata</taxonomicName>
Enderlein by original designation; synonymy
<bibRefCitation id="90F23457BB1A0F7223ABB05649E28640" author="Jeannel, R" journalOrPublisher="Revue Francaise dʼEntomologie" pageId="0" pageNumber="177" pagination="1 - 55" title="Les Migadopides (ColeopteraAdephaga), une lignee subantarctique." volume="5" year="1938">Jeannel 1938</bibRefCitation>
).
</paragraph>
<paragraph id="0AD08F1381C2255C54E8BCADEBB1F6A4" pageId="0" pageNumber="177">
<taxonomicName id="D9D130D98B7CE0A766A8B7EB67418D24" class="Insecta" family="Carabidae" genus="Loeffleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loeffleria" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Loeffleria</taxonomicName>
Mandl, 1969: 54 (type species
<taxonomicName id="E47C53F7F6C68B70ADE25C854E8E1A24" class="Insecta" family="Carabidae" genus="Loeffleria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loeffleria globicollis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="globicollis">Loeffleria globicollis</taxonomicName>
Mandl by monotypy; synonymy
<bibRefCitation id="407792D00E189A4FDE8F84F8B2591C77" author="Baehr, M" journalOrPublisher="Spixiana" pageId="0" pageNumber="177" pagination="263 - 267" title="A reevaluation of Loeffleriaglobicollis Mandl from Borneo (Insecta, Coleoptera, Carabidae, Psydrinae)." volume="22" year="1999">Baehr and Lorenz 1999</bibRefCitation>
).
</paragraph>
<paragraph id="6D1CA171695E3308F6173B66907A00D2" pageId="0" pageNumber="177">
<taxonomicName id="1B4A621323830DA790FB50CBE67204AF" family="Carabidae" genus="Amblytelina" higherTaxonomySource="treatment-meta" infraspecific-rank="subgenus" kingdom="Animalia" lsidName="Amblytelina (Phacothorax)" order="Coleoptera" pageId="0" pageNumber="177" rank="subGenus" subGenus="Phacothorax">subgenus Phacothorax</taxonomicName>
Jeannel, 1944: 84 (type species
<taxonomicName id="EA3A6858515462A1F251322EAEFB1E9B" class="Insecta" family="Carabidae" genus="Phacothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phacothorax fleutiauxi" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="fleutiauxi">Phacothorax fleutiauxi</taxonomicName>
Jeannel by original designation; synonymy
<bibRefCitation id="EBCAFAF85B672E939C9D1B4B6C4BE9F7" author="Liebherr, JK" journalOrPublisher="New Zealand Entomologist" pageId="0" pageNumber="177" pagination="5 - 22" title="Revision of the New Zealand species of Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae, Mecyclothoracini) and the consequent removal of several species to Meonochilus gen. n. (Psydrinae: Meonini)." url="https://doi.org/10.1080/00779962.2009.9722172" volume="32" year="2009">Liebherr and Marris 2009</bibRefCitation>
).
</paragraph>
<paragraph id="182079234ED39E7D5861AC1FB4856B50" pageId="0" pageNumber="177">
<taxonomicName id="6340A02BA80994DF8DBFC32E44BD0EA6" family="Carabidae" genus="Amblytelina" higherTaxonomySource="treatment-meta" infraspecific-rank="subgenus" kingdom="Animalia" lsidName="Amblytelina (Meonochilus)" order="Coleoptera" pageId="0" pageNumber="177" rank="subGenus" subGenus="Meonochilus">subgenus Meonochilus</taxonomicName>
Liebherr &amp; Marris, 2009: 10 (type species
<taxonomicName id="9337DE6D3AF85BF1A91FB7C895E6A383" class="Insecta" family="Carabidae" genus="Tarastethus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tarastethus amplipennis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="amplipennis">Tarastethus amplipennis</taxonomicName>
Broun by original designation; synonymy
<bibRefCitation id="F42B2A01D605AE5D554D066608C0D6DE" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr 2018</bibRefCitation>
).
</paragraph>
<paragraph id="FB4DD2333F180AAE7BAF6440F18C39C3" pageId="0" pageNumber="177">
<taxonomicName id="891625B8CEAC6C60593DE77CFAA3A260" family="Carabidae" genus="Amblytelina" higherTaxonomySource="treatment-meta" infraspecific-rank="subgenus" kingdom="Animalia" lsidName="Amblytelina (Eucyclothorax)" order="Coleoptera" pageId="0" pageNumber="177" rank="subGenus" subGenus="Eucyclothorax">subgenus Eucyclothorax</taxonomicName>
Liebherr, 2018: 12 (type species
<taxonomicName id="56CEF3172BFA5DF808BB5E6C0EC087DF" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax blackburni" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="blackburni">Cyclothorax blackburni</taxonomicName>
Sloane by original designation).
</paragraph>
<paragraph id="108A5CEE67CA2F200A144873D729A0F8" pageId="0" pageNumber="177">
<taxonomicName id="B608B99C111ED5ECF5BACE391CE94026" family="Carabidae" genus="Amblytelina" higherTaxonomySource="treatment-meta" infraspecific-rank="subgenus" kingdom="Animalia" lsidName="Amblytelina (Qecyclothorax)" order="Coleoptera" pageId="0" pageNumber="177" rank="subGenus" subGenus="Qecyclothorax">subgenus Qecyclothorax</taxonomicName>
Liebherr, 2018: 14 (type species
<taxonomicName id="63F86E1869C1554ABA01023EFBA6DD3F" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax storeyi" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="storeyi">Mecyclothorax storeyi</taxonomicName>
Moore by original designation).
</paragraph>
</subSubSection>
<subSubSection id="C6345C1FE264886DB46BB785C087E1F7" pageId="0" pageNumber="177" type="diagnosis">
<paragraph id="28EDB9F8A8E18EC27C003F52CBCAE907" pageId="0" pageNumber="177">Diagnosis.</paragraph>
<paragraph id="85CADA8C872C2BFC28D205318DCDDA69" pageId="0" pageNumber="177">
<bibRefCitation id="DF4EB8A669518DE9746850E7B7AC1F38" author="Moore, BP" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="177" pagination="277 - 290" title="Studies on Australian Carabidae (Coleoptera) - 3. the Psydrinae." url="https://doi.org/10.1111/j.1365-2311.1963.tb00810.x" volume="115" year="1963">Moore (1963)</bibRefCitation>
diagnosed
<taxonomicName id="5BC0239D220417064AD4E9C81036FE29" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
based on: 1, apical palpomeres glabrous; 2, pronotum cordate; 3, male parameres narrowed apically. These characters were chosen to differentiate
<taxonomicName id="859506AF41DD5DC67F02CAB0912856BE" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
from
<taxonomicName id="0C519EB4E91E84DACEE6D8373F57700F" class="Insecta" family="Carabidae" genus="Neonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neonomius" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Neonomius</taxonomicName>
Moore, a genus now placed far from
<taxonomicName id="AE1F266436DC2E16ABD67CFF053B1470" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
in the subtribe
<taxonomicName id="9CE529603518F24726D7CA6B248CD5AA" lsidName="Moriomorphina" pageId="0" pageNumber="177" rank="subTribe" subTribe="Moriomorphina">Moriomorphina</taxonomicName>
. The first character works well for
<taxonomicName id="91DE92604BF2E7FF1D70620416BEE308" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
, with the rare occurrence of very short setae on the apical palpomeres, those setae about 0.10
<normalizedToken id="D48D0C731A3C4FDF93DC94E6563779FB" originalValue="×">x</normalizedToken>
the
<normalizedToken id="9FE85E275FBFFB8427F7674E16C6BE41" originalValue="palpʼs">palpʼs</normalizedToken>
maximal diameter. Since
<normalizedToken id="C8FFD0F3DC661F994D6451B05D00FC70" originalValue="Mooreʼs">Mooreʼs</normalizedToken>
pioneering exposition, pronotal shape has been shown worthless for diagnosing
<taxonomicName id="B638165DD9EC5760DB885BE83276AC5E" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
given the great diversity in shape among the Tahitian (
<bibRefCitation id="E0A87CBB711C1AE7B23EAA329609D212" author="Perrault, GG" journalOrPublisher=")" pageId="0" pageNumber="177" pagination="19 - 31" title="La faune des Carabidae de Tahiti VI. revision du genre Mecyclothorax (Sharp) (Psydrini). 1. le groupe de M. muriauxi Perrault (Coleoptera). Nouvelle Revue dʼEntomologie (nouv. ser." volume="1" year="1984">Perrault 1984</bibRefCitation>
,
<bibRefCitation id="5E9B2DD244C61E2C824F4C3956AD1CFA" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="177" pagination="1 - 170" title="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Tahiti, Society Islands." url="https://doi.org/10.3897/zookeys.322.5492" volume="322" year="2013">Liebherr 2013</bibRefCitation>
), Hawaiian (
<bibRefCitation id="A809446F396C2D3303C27961D6F2E22B" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="177" pagination="1 - 407" title="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation." url="https://doi.org/10.3897/zookeys.544.6074" volume="544" year="2015">Liebherr 2015</bibRefCitation>
), and New Caledonian species (
<bibRefCitation id="DBF2C8990DC4F21A06CCF2CB2763644C" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr 2018</bibRefCitation>
). The male parameres differ in configuration among the various subgenera of
<taxonomicName id="CAF3262FF7CEE3D044340DC6F6506256" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
, but as Moore proposed, both right and left parameres are elongate and setose apically (
<bibRefCitation id="6C57F3983A7C57036F779DEF1A5BA6F3" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="177" pagination="1 - 63" title="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel." url="https://doi.org/10.3897/dez.65.21000" volume="65" year="2018">Liebherr 2018</bibRefCitation>
, fig. 5). Other characters that diagnose
<taxonomicName id="D299BDF3B8B598A98CB53316CA1F90C8" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
from various other taxa in tribe
<taxonomicName id="DD54222F0FA87148A922D84CC6A3966C" lsidName="" pageId="0" pageNumber="177" rank="tribe" tribe="Moriomorphini">Moriomorphini</taxonomicName>
, both symplesiomorphically and synapomorphically (
<bibRefCitation id="EA15E248D2A06BB626C6B5285A0D5E39" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="177" pagination="277 - 335" title="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoyceaglacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)." url="https://doi.org/10.3897/zookeys.147.1898" volume="147" year="2011 a">Liebherr 2011a</bibRefCitation>
), include: 1, labrum truncate to moderately and broadly emarginate apically; 2, ligula truncate medially between the 2 marginal setae, paraglossae elongate; 3, pronotal base at least moderately punctate; 4, elytral humeral margin smooth, without tooth at humeral angle; 5, female apical ventrite with median patch of 4-5 smaller setae.
</paragraph>
</subSubSection>
<subSubSection id="CA9101FE225CE715B4A01E921073876D" pageId="0" pageNumber="177" type="nomenclatural note">
<paragraph id="1A6CCA2241C10BBEB501B3B88C49041F" pageId="0" pageNumber="177">Nomenclatural note.</paragraph>
<paragraph id="7B337B14E1CF7E39B26CE0521F508DD1" pageId="0" pageNumber="177">
<bibRefCitation id="F8DA71A7FF5A3FD3AD30C9FE7AE44B48" author="Sloane, TG" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="177" pagination="438 - 473" title="Studies in Australian Entomology. no. XVII. New genera and species of Carabidae (Pamborini, Migadopini, Broscini, Cuneipectini, Nomiini, Pterostichini, Platynini, Oodini, Harpalini, and Lebiini)." url="https://doi.org/10.5962/bhl.part.18880" volume="40" year="1915">Sloane (1915)</bibRefCitation>
described
<taxonomicName id="C1395E40E1A932A595F1EA7E584A8F4F" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax ovalis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="ovalis">Mecyclothorax ovalis</taxonomicName>
from Manjimup, WA, with label data (ANIC): Manjimup, W.A. / 31.12.13 T. G. S. //
<taxonomicName id="0C2E50702D62FBD83FDD6A145266B28D" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
/
<taxonomicName id="CC9E489DABCB207CDC2895FB48554115" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moriomorphina ovalis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="ovalis">ovalis</taxonomicName>
Sl. Type // Holotype [pink label]. The pink holotype label is consistent with those used by P. J. Darlington, Jr. during his curation of the Sloane collection in 1957 (
<bibRefCitation id="BB69B099DD185226C1B2429E77A66BCF" author="Darlington, PJ Jr" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="0" pageNumber="177" pagination="321 - 564" title="The carabid beetles of New Guinea, part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini." volume="126" year="1962">Darlington 1962</bibRefCitation>
: 328). The specimen is mostly destroyed, with only partial remnants of the legs remaining glued to the card. As such, because the holotype specimen can no longer be used to diagnose the species nor the
<normalizedToken id="AEA7B1E82BB13B7E0CA47BCC4664D1DB" originalValue="species’">species'</normalizedToken>
generic placement, a neotype is designated. As this neotype designation is required to stabilize both the concept of this species and the
<normalizedToken id="6BDE6A3637AE5297AEDFBB3F1E3410C8" originalValue="species’">species'</normalizedToken>
new generic combination, such an action is valid under Article 75 of The Code (
<bibRefCitation id="A0E7379E8A76D62F910CD236708FF65C" author="Huber, BA" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="0" pageNumber="177" title="International Code of Zoological Nomenclature." url="http://iczn.org/iczn/index.jsp" year="1999">ICZN 1999</bibRefCitation>
). Therefore a Neotype male (MCZ specimen deposited in ANIC) is hereby designated: Mullewa, W.A. / Sept. 19 1931 // Australia / Harvard Exp., / Darlington // MCZ // Neotype /
<taxonomicName id="15363C5E0650FDD42C088A8DF8AD3837" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
/
<taxonomicName id="BB33F6E8A803B9E593CC02CC2240AD46" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moriomorphina ovalis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="ovalis">ovalis</taxonomicName>
Sloane / det. J.K. Liebherr 2004 [black-margined red label]. These specimens, both neotype and others in the series (MCZ) exhibit densely setose apical palpomeres as well as the small ovoid body diagnostic for
<taxonomicName id="B132FEDD8400E35F1614A2BECCA80FF7" class="Insecta" family="Carabidae" genus="Neonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neonomius" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Neonomius</taxonomicName>
<bibRefCitation id="B0C4E0380811A84290C0E90B99A12CD7" author="Moore, BP" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="177" pagination="277 - 290" title="Studies on Australian Carabidae (Coleoptera) - 3. the Psydrinae." url="https://doi.org/10.1111/j.1365-2311.1963.tb00810.x" volume="115" year="1963">Moore (1963)</bibRefCitation>
. The male genitalia conform to the configuration observed in the subtribe
<taxonomicName id="BE1307DC72D3A5AA1B28688F3FDAFFED" lsidName="Moriomorphina" pageId="0" pageNumber="177" rank="subTribe" subTribe="Moriomorphina">Moriomorphina</taxonomicName>
(
<bibRefCitation id="5F35807D11FD503CEFCC26AA321C0C2C" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="177" pagination="277 - 335" title="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoyceaglacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae)." url="https://doi.org/10.3897/zookeys.147.1898" volume="147" year="2011 a">Liebherr 2011a</bibRefCitation>
), with both parameres broad, parallel-sided with rounded apices, and the right paramere ventrally setose, as observed in the generic type species
<taxonomicName id="452AE6B85ED209B347DE9042E4418159" lsidName="N. laevicollis" pageId="0" pageNumber="177" rank="species" species="laevicollis">N. laevicollis</taxonomicName>
(Sloane) (
<bibRefCitation id="DE81B50EB24E85CF414A8A864CE1A188" author="Moore, BP" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="177" pagination="277 - 290" title="Studies on Australian Carabidae (Coleoptera) - 3. the Psydrinae." url="https://doi.org/10.1111/j.1365-2311.1963.tb00810.x" volume="115" year="1963">Moore 1963</bibRefCitation>
, fig. 7). Based on these characters,
<normalizedToken id="DAF85369B0544DBFEBA93F5692A25EFF" originalValue="Sloaneʼs">Sloaneʼs</normalizedToken>
species is newly combined as
<taxonomicName id="787EE7283CBEFB377870FD11B6E7C3CE" class="Insecta" family="Carabidae" genus="Neonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neonomius ovalis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="ovalis">Neonomius ovalis</taxonomicName>
(Sloane), comb. n. Among species of
<taxonomicName id="C001F5F2933ECEB3524A8C89DE07A743" class="Insecta" family="Carabidae" genus="Neonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neonomius" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Neonomius</taxonomicName>
,
<taxonomicName id="AE0B0435F27D6E9D6DB46CE352550777" lsidName="N. ovalis" pageId="0" pageNumber="177" rank="species" species="ovalis">N. ovalis</taxonomicName>
is diagnosable from the sympatric
<taxonomicName id="233CD54025EA0C8CD66D5A2F8A5D8915" lsidName="N. australis" pageId="0" pageNumber="177" rank="species" species="australis">N. australis</taxonomicName>
(Sloane) by: 1, the smaller body size, standardized body length 3.2-3.6 mm for the
<taxonomicName id="44D11D4A9DB7231420F4EB5F75AFA0DA" lsidName="N. ovalis" pageId="0" pageNumber="177" rank="species" species="ovalis">N. ovalis</taxonomicName>
neotype series - agreeing with
<bibRefCitation id="8C27761B82A2F54018CB77B264834A4E" author="Sloane, TG" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="177" pagination="438 - 473" title="Studies in Australian Entomology. no. XVII. New genera and species of Carabidae (Pamborini, Migadopini, Broscini, Cuneipectini, Nomiini, Pterostichini, Platynini, Oodini, Harpalini, and Lebiini)." url="https://doi.org/10.5962/bhl.part.18880" volume="40" year="1915">Sloane (1915</bibRefCitation>
; 451) - versus 5.0 mm for
<taxonomicName id="7BFDB4D21DF4340EF537FE0B82281388" lsidName="N. australis" pageId="0" pageNumber="177" rank="species" species="australis">N. australis</taxonomicName>
(
<bibRefCitation id="788FFC9A20AA14E2BC453AD940756312" author="Sloane, TG" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="177" pagination="438 - 473" title="Studies in Australian Entomology. no. XVII. New genera and species of Carabidae (Pamborini, Migadopini, Broscini, Cuneipectini, Nomiini, Pterostichini, Platynini, Oodini, Harpalini, and Lebiini)." url="https://doi.org/10.5962/bhl.part.18880" volume="40" year="1915">Sloane 1915</bibRefCitation>
: 450); and 2, the reddish-brown body color versus &quot;black, nitid; legs piceous-red; antennae reddish (
<bibRefCitation id="8396A554BAB775D44CB0D5108F6ED6B5" author="Sloane, TG" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="177" pagination="438 - 473" title="Studies in Australian Entomology. no. XVII. New genera and species of Carabidae (Pamborini, Migadopini, Broscini, Cuneipectini, Nomiini, Pterostichini, Platynini, Oodini, Harpalini, and Lebiini)." url="https://doi.org/10.5962/bhl.part.18880" volume="40" year="1915">Sloane 1915</bibRefCitation>
, 450)&quot; for
<taxonomicName id="153B86A5A2430CEAF7A042036E24E377" lsidName="N. australis" pageId="0" pageNumber="177" rank="species" species="australis">N. australis</taxonomicName>
.
<taxonomicName id="FA6DF7C20B4E45EFD4D2D4A869F587B7" class="Insecta" family="Carabidae" genus="Neonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neonomius ovalis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="ovalis">Neonomius ovalis</taxonomicName>
can be diagnosed from the other two southeastern Australian species placed in the genus -
<taxonomicName id="3F7255C3CD1B7E79A56255F18C84162C" lsidName="N. laevicollis" pageId="0" pageNumber="177" rank="species" species="laevicollis">N. laevicollis</taxonomicName>
(Sloane) and
<taxonomicName id="E5E02A46782A8B805EF78FD8283BE428" lsidName="N. laticollis" pageId="0" pageNumber="177" rank="species" species="laticollis">N. laticollis</taxonomicName>
(Sloane) - by the apically less convex elytral interval 8, versus the laterally compressed, subcarinate interval 8 characterizing those two species. The type locality for
<taxonomicName id="4437DAF0C74A0CD5E093A2970737B6E4" lsidName="M. ovalis" pageId="0" pageNumber="177" rank="species" species="ovalis">M. ovalis</taxonomicName>
(Sloane) becomes Mullewa. That
<bibRefCitation id="5729F3BB4429A59077DF8E24C5B43FF7" pageId="0" pageNumber="177">Moore et al. (1987)</bibRefCitation>
did not recognize
<normalizedToken id="CDD8EF35428D2B7C9F9C51B93020FB8C" originalValue="Sloane’s">Sloane's</normalizedToken>
<taxonomicName id="E9C6427A89196164F26156C21D642FF2" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax ovalis" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="ovalis">Mecyclothorax ovalis</taxonomicName>
as a member taxon of
<taxonomicName id="8BFFA64F6E942466B82618615754ECE2" class="Insecta" family="Carabidae" genus="Neonomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neonomius" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Neonomius</taxonomicName>
Moore is based on the lack of any other specimens in the ANIC beyond the destroyed holotype of
<taxonomicName id="72D55D7B8CC6B7BE071656E9BE1744F0" lsidName="M. ovalis" pageId="0" pageNumber="177" rank="species" species="ovalis">M. ovalis</taxonomicName>
Sloane, whereas Darlington made this nomenclatural connection, but only for specimens he deposited in the MCZ.
</paragraph>
<paragraph id="C875E3852DB6B5DCEF03A3C6DA637F75" pageId="0" pageNumber="177">
All other names placed under
<taxonomicName id="19CC0504BE5B7909F44D249EFC6FDFF6" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
Sharp in
<bibRefCitation id="057C1AC3611568A0AF2583803A8DDF16" pageId="0" pageNumber="177">Moore et al. (1987</bibRefCitation>
: 147-149) are treated below.
</paragraph>
</subSubSection>
</treatment>
</document>
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