176 lines
20 KiB
XML
176 lines
20 KiB
XML
<document id="0D92F19EB9A9CB9D58D43214E9F2551B" ID-DOI="10.11646/zootaxa.3955.3.9" ID-GBIF-Dataset="a09f0c7f-f65f-47af-bebc-9371363b9653" ID-ISSN="1175-5326" ID-Zenodo-Dep="233117" ID-ZooBank="D018AFA7-2EA9-44BF-AB81-ECBAF132E7AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461051944299" checkinUser="plazi" docAuthor="Häckel, Martin & Anichtchenko, Alexandr" docDate="2015" docId="B85E89401001990CFF5C10A6FD62A9BB" docLanguage="en" docName="zt03955p434.pdf" docOrigin="Zootaxa 3955 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspedophorus luzonensis Häckel & Anichtchenko, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="428" masterDocId="4467F1381000990DFFCB117FFFEEAE78" masterDocTitle="Two new Craspedophorus species (Coleoptera: Carabidae: Panagaeini) from the Philippines" masterLastPageNumber="434" masterPageNumber="427" pageNumber="428" updateTime="1698579237837" updateUser="plazi">
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<mods:mods id="7F0BC8CA485AC9BA2685673046E55544" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EA1083168976CC4F389F45E12DAC6540">
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<mods:title id="5340850BFDD3DD373858722E7DA60C86">Two new Craspedophorus species (Coleoptera: Carabidae: Panagaeini) from the Philippines</mods:title>
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</mods:titleInfo>
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<mods:name id="E1777DEFA860742F19406AC9FD43ED53" type="personal">
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<mods:role id="8483ADD757C3F093E36F4A05DC417E98">
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<mods:namePart id="2DDD2B4931A587511F1164E9B1F85FAD">Häckel, Martin</mods:namePart>
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<mods:name id="83104A181949E5BF5D1BB21A85B5FB2E" type="personal">
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<mods:roleTerm id="5F85651C6834C41F857092DE6B9FC6E7">Author</mods:roleTerm>
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<mods:namePart id="8B6BA098974DB95FD643239897162817">Anichtchenko, Alexandr</mods:namePart>
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<mods:typeOfResource id="3367D81129C67C932E1B311EA88EDD47">text</mods:typeOfResource>
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<mods:title id="CF52FF45B442BCF7A634680D19C78C44">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2933447E04804E904C4DF65225756F17">
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<mods:date id="FCA4AD14E941050B097A51301AC7FB8C">2015</mods:date>
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<mods:detail id="6F89C222CB0EE0EE6C151CFBC06A4704" type="volume">
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<mods:number id="6B9FFAAC03CBBA4B0E6CE478FE72B6E6">3955</mods:number>
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</mods:detail>
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<mods:detail id="9DC30256516074AFDF9D89578C09378A" type="issue">
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<mods:number id="E67519B7E6751ACB6761DAFE75102626">3</mods:number>
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<mods:end id="96E80F7E2A8DF56CB65E87B8467F4890">434</mods:end>
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<mods:classification id="CBB7A1A5C0029F4F3B3C0CAC1FA2BC52">journal article</mods:classification>
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<mods:identifier id="BA53E7B5084181D2DF878F8E80DC13AB" type="DOI">10.11646/zootaxa.3955.3.9</mods:identifier>
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<mods:identifier id="EDDF2270165ED03990D8CAA7F9394FB0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="87AD01FEE764159EED13ED7CEFC04381" type="Zenodo-Dep">233117</mods:identifier>
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</mods:mods>
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<treatment id="B85E89401001990CFF5C10A6FD62A9BB" ID-DOI="http://doi.org/10.5281/zenodo.6111076" ID-GBIF-Taxon="119612163" ID-Zenodo-Dep="6111076" LSID="urn:lsid:plazi:treatment:B85E89401001990CFF5C10A6FD62A9BB" httpUri="http://treatment.plazi.org/id/B85E89401001990CFF5C10A6FD62A9BB" lastPageNumber="428" pageId="1" pageNumber="428">
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<subSubSection id="78ED6BDD1001990CFF5C10A6FF05AC6C" pageId="1" pageNumber="428" type="nomenclature">
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<paragraph id="304838561001990CFF5C10A6FDA6AF8B" blockId="1.[151,584,473,533]" box="[151,584,473,499]" pageId="1" pageNumber="428">
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<heading id="6B008F3A1001990CFF5C10A6FDA6AF8B" bold="true" box="[151,584,473,499]" fontSize="11" level="1" pageId="1" pageNumber="428" reason="1">
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<emphasis id="0283E4441001990CFF5C10A6FDA6AF8B" bold="true" box="[151,584,473,499]" pageId="1" pageNumber="428">
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<taxonomicName id="F7F743D51001990CFF5C10A6FE0AAF8B" ID-CoL="Z5X4" box="[151,484,473,499]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
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<emphasis id="0283E4441001990CFF5C10A6FE0AAF8B" bold="true" box="[151,484,473,499]" italics="true" pageId="1" pageNumber="428">Craspedophorus luzonensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="19B0593F1001990CFE2010A6FDA6AF8B" box="[491,584,473,499]" pageId="1" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="304838561001990CFF5C1083FF05AC6C" blockId="1.[151,584,473,533]" box="[151,235,508,533]" pageId="1" pageNumber="428">
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(
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<figureCitation id="A8CC24D31001990CFF541083FF0EAC6C" box="[159,224,508,533]" captionStart="FIGURE 1" captionStartId="2.[151,250,1957,1979]" captionTargetBox="[390,1212,213,1932]" captionTargetId="figure@2.[340,1248,195,1933]" captionTargetPageId="2" captionText="FIGURE 1. Craspedophorus luzonensis sp. nov.: A. Holotype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233118/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="78ED6BDD1001990CFF5C133BFF31ACDC" pageId="1" pageNumber="428" type="materials_examined">
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<paragraph id="304838561001990CFF5C133BFF31ACDC" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
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<emphasis id="0283E4441001990CFF5C133BFEAAAC25" bold="true" box="[151,324,580,605]" pageId="1" pageNumber="428">
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<typeStatus id="EF4C86F41001990CFF5C133BFF3CAC25" box="[151,210,580,605]" pageId="1" pageNumber="428">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="EF4C86F41001990CFE80133BFE5DAC25" box="[331,435,580,605]" pageId="1" pageNumber="428" type="holotype">Holotype</typeStatus>
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♂: “
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<collectingCountry id="48E078C61001990CFE23133BFD85AC25" box="[488,619,580,605]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
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, N Luzon, Mt. Province, Mt. Polis, 7. 2014, lgt. loc. collector (
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<collectionCode id="56E6A0931001990CFAEC133BFA95AC24" box="[1319,1403,580,604]" pageId="1" pageNumber="428">DUBC</collectionCode>
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); 1
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<typeStatus id="EF4C86F41001990CFF5C1317FF12ACF8" box="[151,252,616,640]" pageId="1" pageNumber="428" type="paratype">Paratype</typeStatus>
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, ♀:
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<collectingCountry id="48E078C61001990CFEE71318FE44ACF8" box="[300,426,615,640]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
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[C Luzon Island], Nueva Vizcaya, Dupax del Sur,
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<date id="44491E961001990CFC261318FB48ACF8" box="[1005,1190,615,640]" pageId="1" pageNumber="428" value="2012-11">November 2012</date>
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, lgt. Ismael Lumawig (
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<collectionCode id="56E6A0931001990CFF5413F3FF3DACDC" box="[159,211,652,676]" httpUri="http://grbio.org/cool/vxkr-andy" name="Centro de Investigaciones Biologicas del Noroeste, S.C. (Mexico)" pageId="1" pageNumber="428">CIB</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="78ED6BDD1001990CFF0C13D0FBDAA903" pageId="1" pageNumber="428" type="description">
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<paragraph id="304838561001990CFF0C13D0FD39AC94" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
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<emphasis id="0283E4441001990CFF0C13D0FE08ACB0" bold="true" box="[199,486,687,712]" pageId="1" pageNumber="428">
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Description of
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<typeStatus id="EF4C86F41001990CFEB313D0FE0FACB0" box="[376,481,687,712]" pageId="1" pageNumber="428" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Length
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<quantity id="F70F95B31001990CFD8F13CFFD78ACBF" box="[580,662,688,712]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="428" unit="mm" value="17.0">17 mm</quantity>
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, width
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<quantity id="F70F95B31001990CFD2113D0FCADACBF" box="[746,835,687,712]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="428" unit="mm" value="6.5">6.5 mm</quantity>
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. Proportions: Pronotum 1.27x wider than long, 1.67x wider than head, elytra 1.41x wider than pronotum.
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</paragraph>
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<paragraph id="304838561001990CFF0C1388FC0FADF9" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">Colour: Head, pronotum and elytra black, glossy; mandibles, palps, tarsi, base of labrum, base of antennomere IV lighter, reddish. Legs and antennae ferrugineous, proximal parts of femur darkish. Elytron with two yellow spots; humeral spot reaches from 5th interval (4th stria) to margin and to epipleura, preapical spot reaches from 4th to 8th interval. Dorsal and ventral part of body black, covered by yellow setae.</paragraph>
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<paragraph id="304838561001990CFF0C12F2FF11AA22" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
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<emphasis id="0283E4441001990CFF0C12F2FEEAADDD" box="[199,260,909,933]" italics="true" pageId="1" pageNumber="428">Head</emphasis>
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large, wide, only 1.67x narrower than pronotum; eyes moderately convex; temples short; labrum strongly excavated at anterior side; clypeus smooth, with four long, yellow setae; vertex convex, sparsely punctured and covered by long, reddish setae; anterior part of frons smooth, fairly convex at middle, laterally depressed, rugate, lateral margin with sulcus rimming it from mandibular base to anterior margin of eyes, anterior side arcuate, sinuate before eyes, moderately impressed, smooth, sharply bordered externally, indistinctly bordered medially.
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</paragraph>
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<paragraph id="304838561001990CFF0C1519FC13AB4A" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
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<emphasis id="0283E4441001990CFF0C1519FED8AA05" box="[199,310,1126,1149]" italics="true" pageId="1" pageNumber="428">Pronotum</emphasis>
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weakly transverse (length to width ratio 1.27), almost hexagonal, widest at middle; lateral angles widely rounded, weakly arcuate anteriorly; anterior angles weakly rounded and distinctly protruded; anterior margin as wide as neck, flat between angular lobes; lateral margins weakly sinuate toward base; base parallel, oblique laterally near posterior angle; lateral rims punctured, narrowing anteriorly, posteriorly wider and more elevated; disc densely rugate and punctured; sagittal line distinctly impressed, reaching neither anterior margin nor base; basal pits elongate, narrowly and deeply impressed, coarsely punctured.
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</paragraph>
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<paragraph id="304838561001990CFF0C1442FDB2ABBA" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
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<emphasis id="0283E4441001990CFF0C1442FEE3AB2D" box="[199,269,1341,1365]" italics="true" pageId="1" pageNumber="428">Elytra</emphasis>
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almost parallel-sided, widest at middle, elongate, slightly convex, weakly tuberose in front of apex; disc flattened; humerus weakly developed; margins rounded, widening posterio-laterally; basal rim indistinct striae deeply impressed and very slightly punctured; intervals moderately convex, sparsely but coarsely punctured, interspaces with distinct microsculpture.
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</paragraph>
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<paragraph id="304838561001990CFF0C14B1FBF2A871" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
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<emphasis id="0283E4441001990CFF0C14B1FEE2AB9D" box="[199,268,1486,1509]" italics="true" pageId="1" pageNumber="428">Venter</emphasis>
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black, metepisternum transverse, punctured, weakly elongated and narrowed towards posterior. Ventrites laterally rugate, finely rugate at middle, densely covered by long setae.
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</paragraph>
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<paragraph id="304838561001990CFF0C176AFAE0A856" blockId="1.[151,1437,580,1987]" box="[199,1294,1557,1582]" pageId="1" pageNumber="428">
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Aedeagal median lobe stout, strongly curved laterally, apical lamella short, apex rounded (
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<figureCitation id="A8CC24D31001990CFB70176AFB12A856" box="[1211,1276,1557,1582]" captionStart="FIGURE 1" captionStartId="2.[151,250,1957,1979]" captionTargetBox="[390,1212,213,1932]" captionTargetId="figure@2.[340,1248,195,1933]" captionTargetPageId="2" captionText="FIGURE 1. Craspedophorus luzonensis sp. nov.: A. Holotype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233118/files/figure.png" pageId="1" pageNumber="428">Fig. 1</figureCitation>
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).
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</paragraph>
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<paragraph id="304838561001990CFF0C1746FBDAA903" blockId="1.[151,1437,580,1987]" pageId="1" pageNumber="428">
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<emphasis id="0283E4441001990CFF0C1746FE3FA82A" bold="true" box="[199,465,1593,1618]" pageId="1" pageNumber="428">Differential diagnosis.</emphasis>
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Based on similar habitus and elytral pattern,
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<taxonomicName id="F7F743D51001990CFC201745FB6BA829" box="[1003,1157,1593,1617]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
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<emphasis id="0283E4441001990CFC201745FB6BA829" box="[1003,1157,1593,1617]" italics="true" pageId="1" pageNumber="428">C. luzonensis</emphasis>
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</taxonomicName>
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<emphasis id="0283E4441001990CFB441746FB03A82A" bold="true" box="[1167,1261,1593,1618]" pageId="1" pageNumber="428">
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<taxonomicNameLabel id="19B0593F1001990CFB441746FB03A82A" box="[1167,1261,1593,1618]" pageId="1" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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belongs to the
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<emphasis id="0283E4441001990CFF5C1722FEE2A80D" box="[151,268,1629,1653]" italics="true" pageId="1" pageNumber="428">“obscurus</emphasis>
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species group sensu
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<bibRefCitation id="546645A71001990CFDEA1722FC47A80E" author="Hackel" box="[545,937,1629,1654]" pageId="1" pageNumber="428" refString="Hackel, M. & Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel & Kirschenhofer (2014)</bibRefCitation>
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. It resembles
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<taxonomicName id="F7F743D51001990CFBA91721FE90A8E2" authority="Hackel & Kirschenhofer, 2014" authorityName="Hackel & Kirschenhofer" authorityYear="2014" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="cenwanglao">
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<emphasis id="0283E4441001990CFBA91721FAFAA80D" box="[1122,1300,1629,1653]" italics="true" pageId="1" pageNumber="428">C. cenwanglao</emphasis>
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<bibRefCitation id="546645A71001990CFAEE1722FE90A8E2" author="Hackel" pageId="1" pageNumber="428" refString="Hackel, M. & Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel & Kirschenhofer, 2014</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="A8CC24D31001990CFE4417FEFE3AA8E2" box="[399,468,1665,1690]" captionStart="FIGURE 2. C" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[412,1147,238,1901]" captionTargetId="figure@3.[369,1218,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. C. cenwanglao Häckel & Kirschenhofer, 2014: A. Paratype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233119/files/figure.png" pageId="1" pageNumber="428">Fig. 2</figureCitation>
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) from western Guangxi,
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<collectingCountry id="48E078C61001990CFD3E17FEFCD5A8E2" box="[757,827,1665,1690]" name="China" pageId="1" pageNumber="428">China</collectingCountry>
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, and
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<taxonomicName id="F7F743D51001990CFCB217FDFA72A8E2" authority="Hackel & Kirschenhofer, 2014" authorityName="Hackel & Kirschenhofer" authorityYear="2014" box="[889,1436,1665,1690]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="phoupanensis">
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<emphasis id="0283E4441001990CFCB217FDFBDBA8E1" box="[889,1077,1665,1689]" italics="true" pageId="1" pageNumber="428">C. phoupanensis</emphasis>
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<bibRefCitation id="546645A71001990CFBF517FEFA72A8E2" author="Hackel" box="[1086,1436,1665,1690]" pageId="1" pageNumber="428" refString="Hackel, M. & Kirchenhofer, E. (2014) A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Asia. Part 2. Genus Craspedophorus Hope, 1838, east Palearctic and Oriental species, and genus Tinoderus Chaudoir, 1879 (Coleoptera: Carabidae). Studies and Reports, Taxonomical Series, 10, 275 - 392." type="journal article" year="2014">Häckel & Kirschenhofer, 2014</bibRefCitation>
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</taxonomicName>
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from northern
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<collectingCountry id="48E078C61001990CFEF717D9FE9BA8C5" box="[316,373,1702,1725]" name="Laos" pageId="1" pageNumber="428">Laos</collectingCountry>
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, particularly in the pronotum. The new species differs from them by somewhat more elongate habitus, less convex elytra, elytra with more grossly and deeply punctured striae and more extended spots; the humeral spot extends from 4th stria (external part of 4th interval) to margin and the preapical spot extends from 3rd stria (4th interval) to 9th interval, contrary to both continental species in which the humeral spot reaches from external part of 5th interval and the preapical spot from 4th stria (5th interval) to 9th interval (
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<figureCitation id="A8CC24D31001990CFB721642FAEFA92E" box="[1209,1281,1853,1878]" captionStart="FIGURE 2. C" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[412,1147,238,1901]" captionTargetId="figure@3.[369,1218,193,1933]" captionTargetPageId="3" captionText="FIGURE 2. C. cenwanglao Häckel & Kirschenhofer, 2014: A. Paratype, B. Lateral view of aedeagus." httpUri="https://zenodo.org/record/233119/files/figure.png" pageId="1" pageNumber="428">Fig. 2</figureCitation>
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). In addition, intervals are less dense and more coarsely punctured than in
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<taxonomicName id="F7F743D51001990CFCF5161DFC3AA902" box="[830,980,1890,1914]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="428" phylum="Arthropoda" rank="species" species="luzonensis" status="sp. nov.">
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<emphasis id="0283E4441001990CFCF5161DFC3AA902" box="[830,980,1890,1914]" italics="true" pageId="1" pageNumber="428">C. luzonensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="0283E4441001990CFC10161DFBDAA903" bold="true" box="[987,1076,1890,1915]" pageId="1" pageNumber="428">
|
|
<taxonomicNameLabel id="19B0593F1001990CFC10161DFBDAA903" box="[987,1076,1890,1915]" pageId="1" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="78ED6BDD1001990CFF0C16FAFB61A9E6" box="[199,1167,1925,1950]" pageId="1" pageNumber="428" type="etymology">
|
|
<paragraph id="304838561001990CFF0C16FAFB61A9E6" blockId="1.[151,1437,580,1987]" box="[199,1167,1925,1950]" pageId="1" pageNumber="428">
|
|
<emphasis id="0283E4441001990CFF0C16FAFEA0A9E6" bold="true" box="[199,334,1925,1950]" pageId="1" pageNumber="428">Etymology.</emphasis>
|
|
Named after the place of its occurrence - Luzon, island in the
|
|
<collectingCountry id="48E078C61001990CFBC216FAFB65A9E6" box="[1033,1163,1925,1950]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="78ED6BDD1001990CFF0C16D5FD62A9BB" box="[199,652,1962,1987]" pageId="1" pageNumber="428" type="distribution">
|
|
<paragraph id="304838561001990CFF0C16D5FD62A9BB" blockId="1.[151,1437,580,1987]" box="[199,652,1962,1987]" pageId="1" pageNumber="428">
|
|
<emphasis id="0283E4441001990CFF0C16D5FE8EA9BB" bold="true" box="[199,352,1962,1987]" pageId="1" pageNumber="428">Distribution.</emphasis>
|
|
Luzon island,
|
|
<collectingCountry id="48E078C61001990CFDCD16D5FD66A9BB" box="[518,648,1962,1987]" name="Philippines" pageId="1" pageNumber="428">Philippines</collectingCountry>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |