182 lines
20 KiB
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182 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115688" ID-Pensoft-Pub="2747-8106-97-29" ID-Pensoft-UUID="CA664780046A5F938793904BB13AEB75" ID-ZooBank="BA2DA07949064AC8AE1105E21BBF12B9" ModsDocID="2747-8106-97-29" checkinTime="1702462890041" checkinUser="pensoft" docAuthor="Nuetzel, Alexander, Ebbestad, Jan Ove, Seuss, Barbara, Munnecke, Axel, Mapes, Royal H. & Cook, Alex G." docDate="2023" docId="B218C5B73C58578AA3889ED788410843" docLanguage="en" docName="Zitteliana 97: 29-51" docOrigin="Zitteliana 97" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.97.115688" docTitle="Orthonychia Hall 1843" docType="treatment" docVersion="1" id="CA664780046A5F938793904BB13AEB75" lastPageNumber="29" masterDocId="CA664780046A5F938793904BB13AEB75" masterDocTitle="On Paleozoic platycerate gastropods" masterLastPageNumber="51" masterPageNumber="29" pageNumber="29" updateTime="1702462890041" updateUser="pensoft">
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<mods:titleInfo id="1E859B278FFFDDB819BEC1D67E230643">
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<mods:title id="B4FAAF69EF2338D81820D16ED65D1299">On Paleozoic platycerate gastropods</mods:title>
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<mods:namePart id="B75A96A70B50749AB698110DCBFF1451">Nuetzel, Alexander</mods:namePart>
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<mods:nameIdentifier id="97CC0B2F8B74BB46805B661233C0AA4A" type="ORCID">https://orcid.org/0000-0002-8852-7688</mods:nameIdentifier>
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<mods:affiliation id="9AF0256B1926C72B4A20AE8266971BC9">SNSB-Bayerische Staatssammlung fuer Palaeontologie und Geologie, Richard-Wagner-Str. 10, 80333 Muenchen, Germany & Department of Earth and Environmental Sciences, Paleontology and Geobiology, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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<mods:nameIdentifier id="A72593372E7A5A40E81411A7492B6157" type="email">nuetzel@snsb.de</mods:nameIdentifier>
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<mods:namePart id="EB301FA9E771327D8EBF91CAFBB07E29">Ebbestad, Jan Ove</mods:namePart>
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<mods:nameIdentifier id="9CC60A677514DA74688BCD27C168652B" type="ORCID">https://orcid.org/0000-0001-8769-3572</mods:nameIdentifier>
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<mods:affiliation id="161F2DEB238917970819D4D81879883C">GeoBio-Center, Ludwig-Maximilians-Universitaet Muenchen, Richard-Wagner-Str. 10, 80333 Muenchen, Germany</mods:affiliation>
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<mods:namePart id="9ED0A2DC5BE498A41E9EC623F5F040AF">Seuss, Barbara</mods:namePart>
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<mods:nameIdentifier id="6F4569BE3A0BD2C79E44B5A024079277" type="ORCID">https://orcid.org/0000-0001-6615-8108</mods:nameIdentifier>
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<mods:affiliation id="946261E5025E689CD4E3472C030E10E5">Museum of Evolution, Uppsala University, Norbyvaegen 16, SE 75236 Uppsala, Sweden</mods:affiliation>
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<mods:namePart id="E11763679D907188E52EAE31E9A30D51">Munnecke, Axel</mods:namePart>
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<mods:nameIdentifier id="3B9F43EC8869F1A04BF15B7A1938C8C1" type="ORCID">https://orcid.org/0000-0002-6898-1082</mods:nameIdentifier>
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<mods:affiliation id="8042171054C5BC0B14098156EAB3218D">Museum of Evolution, Uppsala University, Norbyvaegen 16, SE 75236 Uppsala, Sweden</mods:affiliation>
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<mods:namePart id="FAEE1E0A63C4B393F3274A40E10FCCD0">Mapes, Royal H.</mods:namePart>
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<mods:affiliation id="5EE94331DD41D87EA3A63279E863C375">Friedrich-Alexander-Universitaet Erlangen-Nuernberg, GeoZentrum Nordbayern, Loewenichstrasse 28, 91054 Erlangen, Germany</mods:affiliation>
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<mods:namePart id="C84EBE90776E1A0CD453110FA5382375">Cook, Alex G.</mods:namePart>
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<mods:affiliation id="7ADCF89D2B96552B17EB296C5759CE55">Division of Paleontology, American Museum of Natural History, Central Park West at 79 th Street, New York, NY, 10024, USA</mods:affiliation>
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<mods:typeOfResource id="9B028FC5EFC9C336E82F48658AE12C59">text</mods:typeOfResource>
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<mods:title id="ED93440F7C0B15CE49F118813DE2DE85">Zitteliana</mods:title>
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<mods:part id="0544A58FA2ADAE0FCC89D1F5D1D0E9DD">
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<mods:date id="DC069502B3A2022F1AD3288CA30A57FF">2023</mods:date>
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<mods:number id="E1C70EC15DDEFD32EF22BA0D13C57752">2023-12-12</mods:number>
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<mods:detail id="635DBDB6A057380C4CAD1FCEA8DB4373" type="volume">
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<mods:number id="85C4A3243DE32582717F1C149DAC9D2F">97</mods:number>
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<mods:start id="AC0F03416B4F18B9FD996D7788513DE2">29</mods:start>
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<mods:end id="F40875EA7C2F4F0563D81E99ACE033DF">51</mods:end>
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<mods:location id="9AF54F44160C9B093AC40DFCC84326DF">
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<mods:url id="D947A551954B225B9BBAB89964D8C8C4">http://dx.doi.org/10.3897/zitteliana.97.115688</mods:url>
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<mods:classification id="A790B17E2749BF3ED25FFA017D7775DD">journal article</mods:classification>
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<mods:identifier id="7B193AA4C39B0FFB0687286BABC05686" type="DOI">http://dx.doi.org/10.3897/zitteliana.97.115688</mods:identifier>
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<treatment id="B218C5B73C58578AA3889ED788410843" LSID="urn:lsid:plazi:treatment:B218C5B73C58578AA3889ED788410843" httpUri="http://treatment.plazi.org/id/B218C5B73C58578AA3889ED788410843" lastPageNumber="29" pageId="0" pageNumber="29">
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<subSubSection id="2252A28975A4FD4F553FE159C2EC7B16" pageId="0" pageNumber="29" type="nomenclature">
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<paragraph id="1BAC53C3C27800DCBC7275AB79B2B23D" pageId="0" pageNumber="29">
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Genus
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<taxonomicName id="B11580E54FBAF61DD07CD6F36C3DE58E" ID-CoL="8YJL2" LSID="B218C5B7-3C58-578A-A388-9ED788410843" authority="Hall, 1843" authorityName="Hall" authorityYear="1843" class="Gastropoda" family="Orthonychiidae" genus="Orthonychia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Orthonychia" order="Cyrtoneritida" pageId="0" pageNumber="29" phylum="Mollusca" rank="genus">Orthonychia Hall, 1843</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="1BCBD77D433EB5274C282D73679CC7EC" pageId="0" pageNumber="29" type="type species">
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<paragraph id="71AFCD81F34BA6CC58ECD5D7AA041569" pageId="0" pageNumber="29">Type species.</paragraph>
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<paragraph id="39804CD7EA3AD4A537AF83122010BB66" pageId="0" pageNumber="29">
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<taxonomicName id="76DBD843273D66B3D9E1CBC78B13EAF7" baseAuthorityName="Hall" baseAuthorityYear="1859" class="Gastropoda" family="Orthonychiidae" genus="Platyceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyceras subrectum" order="Vetigastropoda" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="subrectum">
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<emphasis id="D388A34E66A748BC14314BD778E0AF26" italics="true" pageId="0" pageNumber="29">Platyceras subrectum</emphasis>
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</taxonomicName>
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Hall, 1859; by subsequent monotypy by
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<bibRefCitation id="4B365429F929FAE734ED00E9FF8F7B13" author="Hall, J" journalOrPublisher="Carroll and Cook, Albany" pageId="0" pageNumber="29" refId="B37" refString="Hall, J, 1859. Contributions to the palaeontology of New-York; being some results of investigations made during the years 1855, '56, '57, & '58: Twelfth Annual Report of the Regents of the University of the State of New-York, on the Condition of the State Cabinet of Natural History, and the Historical and Antiquarian Collection Connected therewith, State of New York in Assembly Document 186, C. van Benthuysen, Albany, 8-110." title="Contributions to the palaeontology of New-York; being some results of investigations made during the years 1855, ' 56, ' 57, & ' 58: Twelfth Annual Report of the Regents of the University of the State of New-York, on the Condition of the State Cabinet of Natural History, and the Historical and Antiquarian Collection Connected therewith, State of New York in Assembly Document 186, C. van Benthuysen, Albany, 8 - 110." year="1859">Hall (1859)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="6DF95F938F593ED440CE76D652CDADE4" pageId="0" pageNumber="29" type="remarks">
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<paragraph id="8A2B5E56AABD323AC34A7C9794FEBAC6" pageId="0" pageNumber="29">Remarks.</paragraph>
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<paragraph id="E26995597D2B5EB2B274AFAA617527AB" pageId="0" pageNumber="29">
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<taxonomicName id="8AEFFEED9E99ED7F4B6E48B0FDF401AD" authorityName="Hall" authorityYear="1843" class="Gastropoda" family="Orthonychiidae" genus="Orthonychia" kingdom="Animalia" lsidName="Orthonychia" order="Cyrtoneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="genus">
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<emphasis id="631337461D1E08F44801A5D719486CFC" italics="true" pageId="0" pageNumber="29">Orthonychia</emphasis>
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</taxonomicName>
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is a cap-shaped or elongate tube-like mollusk which has traditionally been placed in the in the Paleozoic gastropod family
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<taxonomicName id="7CF97024E0E9F595F991EDEB3F720234" class="Gastropoda" family="Platyceratidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" pageId="0" pageNumber="29" phylum="Mollusca" rank="family">Platyceratidae</taxonomicName>
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and was also considered a subgenus of
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<taxonomicName id="064D5AD14F73F0B36E33EC4B88542D0D" class="Gastropoda" family="Orthonychiidae" genus="Platyceras" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyceras" order="Vetigastropoda" pageId="0" pageNumber="29" phylum="Mollusca" rank="genus">
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<emphasis id="618FFF2C4C1A45EAF08EE1BA62A6BA87" italics="true" pageId="0" pageNumber="29">Platyceras</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="850DB49C27F596FA402FA5F573399707" author="Knight, JB" journalOrPublisher="Geological Society of America Special Papers" pageId="0" pageNumber="29" refId="B53" refString="Knight, JB, Cox, LR, Keen, AM, Batten, RL, Yochelson, EL, Robertson, R, 1960. Systematic descriptions. In: Moore RC (Ed.) Treatise on invertebrate paleontology, Part 1, Mollusca 1. Geological Society of America and University of Kansas Press, Kansas, I169-I310." title="Systematic descriptions. In: Moore RC (Ed.) Treatise on invertebrate paleontology, Part 1, Mollusca 1. Geological Society of America and University of Kansas Press, Kansas, I 169 - I 310." year="1960">Knight et al. 1960</bibRefCitation>
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). The Devonian type species of
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<taxonomicName id="6050F0EB5BA33AD36EB481AA0253EA07" authorityName="Hall" authorityYear="1843" class="Gastropoda" family="Orthonychiidae" genus="Orthonychia" kingdom="Animalia" lsidName="Orthonychia" order="Cyrtoneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="genus">
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<emphasis id="0607A0D571F1D42F953E1C20C1889D9F" italics="true" pageId="0" pageNumber="29">Orthonychia</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="B840ED68991CEBECD3CE6044264A77DE" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. subrecta" order="Cycloneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="subrecta">
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<emphasis id="1B26458E8F220B4E8E465BBBAAF3C20A" italics="true" pageId="0" pageNumber="29">O. subrecta</emphasis>
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</taxonomicName>
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(Hall, 1859), is known from steinkerns only (
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<bibRefCitation id="029EE95466ACE602192137EC544EF2CF" DOI="https://doi.org/10.1130/SPE32-p1" author="Knight, JB" journalOrPublisher="Geological Society of America Special Papers" pageId="0" pageNumber="29" pagination="1 - 510" refId="B52" refString="Knight, JB, 1941. Palaeozoic gastropod genotypes. Geological Society of America Special Papers 32: 1 - 510, DOI: https://doi.org/10.1130/SPE32-p1" title="Palaeozoic gastropod genotypes." url="https://doi.org/10.1130/SPE32-p1" volume="32" year="1941">Knight 1941</bibRefCitation>
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) and therefore, the status of shell ornamentation and protoconch morphology cannot be assessed. The general habitus (elongated slightly curved shell with only the early teleoconch being coiled) is close to that of
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<taxonomicName id="80218A16DE2117858CB8639933BECEDF" baseAuthorityName="Lindstroem" baseAuthorityYear="1884" class="Gastropoda" family="Orthonychiidae" genus="Orthonychia" kingdom="Animalia" lsidName="Orthonychia enorme" order="Cyrtoneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="enorme">
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<emphasis id="443899B30266AFE947C61A35D533BDAA" italics="true" pageId="0" pageNumber="29">Orthonychia enorme</emphasis>
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</taxonomicName>
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(
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<normalizedToken id="9B358CA763B702F7C2C4422CAC49DE00" originalValue="Lindström">Lindstroem</normalizedToken>
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, 1884) and
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<taxonomicName id="9968259DE8A0E232BDEBC1CA6FF7EE8C" authorityName="Ebbestad" authorityYear="2023" class="Gastropoda" family="Orthonychiidae" genus="Orthonychia" kingdom="Animalia" lsidName="Orthonychia yutaroi" order="Cyrtoneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="yutaroi">
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<emphasis id="975879246C5592B6DEE1D01B45AB33AB" italics="true" pageId="0" pageNumber="29">Orthonychia yutaroi</emphasis>
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</taxonomicName>
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Ebbestad, sp. nov. as described below, especially regarding the tube-shaped, stretched late teleoconch. Based on an openly coiled fish-hook-like protoconch, present in the Pennsylvanian species
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<taxonomicName id="04EB31D4D724117570A352B5552C5BAC" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. parva" order="Cycloneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="parva">
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<emphasis id="0AD3021C1612D0502F0759B029C5208A" italics="true" pageId="0" pageNumber="29">O. parva</emphasis>
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</taxonomicName>
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,
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<bibRefCitation id="E2FB633F7F5560C05147B8AB02CA87B3" author="Bandel, K" journalOrPublisher="Geologica et Palaeontologica" pageId="0" pageNumber="29" pagination="219 - 235" refId="B7" refString="Bandel, K, Fryda, J, 1999. Notes on the evolution and higher classification of the subclass Neritimorpha (Gastropoda) with the description of some new taxa. Geologica et Palaeontologica 33: 219 - 235" title="Notes on the evolution and higher classification of the subclass Neritimorpha (Gastropoda) with the description of some new taxa." volume="33" year="1999">
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Bandel and
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<normalizedToken id="4DE50E0446530A82D539DD37E8D46A03" originalValue="Frýda">Fryda</normalizedToken>
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(1999)
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</bibRefCitation>
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erected the family
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<taxonomicName id="19536874DA9BCEA368F59AF3AE20E64B" authorityName="Bandel & Fryda" authorityYear="1999" class="Gastropoda" family="Orthonychiidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Cyrtoneritida" pageId="0" pageNumber="29" phylum="Mollusca" rank="family">Orthonychiidae</taxonomicName>
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and placed it in a new order
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<taxonomicName id="520975AD3017806C4ADC8F55F273C69B" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Cyrtoneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="order">Cyrtoneritimorpha</taxonomicName>
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, forming the fossil sister group of modern
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<taxonomicName id="6121F5076D2AAB9239E07D12ABBC0C51" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" pageId="0" pageNumber="29" phylum="Mollusca" rank="subClass" subClass="Neritimorpha">Neritimorpha</taxonomicName>
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(
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<taxonomicName id="4542E3EB6F5377052E02CDC58C022333" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Cycloneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="order">Cycloneritimorpha</taxonomicName>
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).
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</paragraph>
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<paragraph id="154B047D9F5F39788F7FF04A9ADDDC85" pageId="0" pageNumber="29">
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The tall shell morphology of some
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<taxonomicName id="8CCCC4CC5BEAF372DE293E07469E1B5B" authorityName="Hall" authorityYear="1843" class="Gastropoda" family="Orthonychiidae" genus="Orthonychia" kingdom="Animalia" lsidName="Orthonychia" order="Cyrtoneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="genus">
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<emphasis id="88C7A26E0128E324F02BDC5BEBB71CB2" italics="true" pageId="0" pageNumber="29">Orthonychia</emphasis>
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</taxonomicName>
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species including
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<taxonomicName id="C6E49F1FD809CAC30253646027502AC0" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. yutaroi" order="Cycloneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="yutaroi">
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<emphasis id="ECD685BC4252B8CF1E4EF82E47218295" italics="true" pageId="0" pageNumber="29">O. yutaroi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AB82BEF56AE17ABBB9851E43FA89D5E1" class="Gastropoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. enorme" order="Cycloneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="species" species="enorme">
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<emphasis id="C16E9B316ED6623AB75A5D5D8AA8FC0E" italics="true" pageId="0" pageNumber="29">O. enorme</emphasis>
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</taxonomicName>
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as reported herein superficially resembles that of the Ordovician archaeogastropod
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<taxonomicName id="FC0435D517DBD0A120CADD642D61E734" class="Gastropoda" family="Pollicinidae" genus="Pollicina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pollicina" pageId="0" pageNumber="29" phylum="Mollusca" rank="genus">
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<emphasis id="89013D66E99EC8610183A7358AE77668" italics="true" pageId="0" pageNumber="29">Pollicina</emphasis>
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</taxonomicName>
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, described by
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<bibRefCitation id="E590F5C1630B954FF74E1EA521D3B7ED" DOI="https://doi.org/10.3176/earth.2020.02" author="Peel, JS" journalOrPublisher="Estonian Journal of Earth Sciences" pageId="0" pageNumber="29" pagination="20 - 36" refId="B75" refString="Peel, JS, 2020a. Muscle scars, mode of life and systematics of Pollicina (Mollusca) from the Ordovician of Baltica. Estonian Journal of Earth Sciences 69: 20 - 36, DOI: https://doi.org/10.3176/earth.2020.02" title="Muscle scars, mode of life and systematics of Pollicina (Mollusca) from the Ordovician of Baltica." url="https://doi.org/10.3176/earth.2020.02" volume="69" year="2020 a">Peel (2020a</bibRefCitation>
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,
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<bibRefCitation id="24598E8FE46DD2D857AFD685226ED600" DOI="https://doi.org/10.1080/11035897.2020.1728374" author="Peel, JS" journalOrPublisher="GFF" pageId="0" pageNumber="29" pagination="139 - 146" refId="B76" refString="Peel, JS, 2020b. The Dala thumb: shell morphology and failed predation in Pollicina cyathina (Gastropoda) from the Ordovician of Dalarna, Sweden. GFF 142: 139 - 146, DOI: https://doi.org/10.1080/11035897.2020.1728374" title="The Dala thumb: shell morphology and failed predation in Pollicina cyathina (Gastropoda) from the Ordovician of Dalarna, Sweden." url="https://doi.org/10.1080/11035897.2020.1728374" volume="142" year="2020 b">b</bibRefCitation>
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). This genus can reach a height of at least 3 cm, has a thick shell, with even co-marginal ribs, and an apex truncated by septa. The shell is bilaterally symmetrical, in contrast to that of
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<taxonomicName id="915B5AA4168711D9FB4250FC077EA0AE" authorityName="Hall" authorityYear="1843" class="Gastropoda" family="Orthonychiidae" genus="Orthonychia" kingdom="Animalia" lsidName="Orthonychia" order="Cyrtoneritimorpha" pageId="0" pageNumber="29" phylum="Mollusca" rank="genus">
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<emphasis id="589AD0DB63FD3C95678A7A68B16B85E6" italics="true" pageId="0" pageNumber="29">Orthonychia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |