treatments-xml/data/84/70/87/84708794EF45FFA6FF709B726748F8FF.xml

<document id="554E79C0B3AD57FB3843573A8269B0CC" ID-DOI="10.5281/zenodo.211031" ID-GBIF-Dataset="db0a2f30-28ed-43b2-8483-cd280ee1541a" ID-ISSN="1175-5326" ID-Zenodo-Dep="211031" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460540065668" checkinUser="plazi" docAuthor="Luque, Carlos G. &amp; Labrada, Lucia" docDate="2012" docId="84708794EF45FFA6FF709B726748F8FF" docLanguage="en" docName="zt03506p042.pdf" docOrigin="Zootaxa 3506" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ischyropsalis cantabrica Luque &amp; Labrada, 2012, sp. n." docType="treatment" docVersion="7" lastPageNumber="34" masterDocId="7849FFECEF47FFAEFFE7992C6379FF94" masterDocTitle="A new cave-dwelling endemic Ischyropsalis C. L. Koch, 1839 (Opiliones: Dyspnoi: Ischyropsalididae) from the karstic region of Cantabria (Spain)" masterLastPageNumber="42" masterPageNumber="26" pageNumber="28" updateTime="1698316911919" updateUser="plazi">
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<mods:title id="2C70011583CADC6ADDAC1258E17125E4">A new cave-dwelling endemic Ischyropsalis C. L. Koch, 1839 (Opiliones: Dyspnoi: Ischyropsalididae) from the karstic region of Cantabria (Spain)</mods:title>
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<mods:namePart id="484D2E8DD212412EBAC257CB22072ABC">Luque, Carlos G.</mods:namePart>
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<mods:namePart id="824BE3B16DBEA4DFE34CE21123244353">Labrada, Lucia</mods:namePart>
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<mods:date id="54723F8C3F517B6F3A05E6E5A408DC42">2012</mods:date>
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<paragraph id="0C663682EF45FFACFF709B72617EFDEC" blockId="2.[151,519,606,665]" box="[151,519,606,632]" pageId="2" pageNumber="28">
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<taxonomicName id="CBD94D01EF45FFACFF709B7262C7FDEC" ID-CoL="7WRHM" box="[151,446,606,632]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF45FFACFF709B7262C7FDEC" bold="true" box="[151,446,606,632]" italics="true" pageId="2" pageNumber="28">Ischyropsalis cantabrica</emphasis>
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<taxonomicNameLabel id="259E57EBEF45FFACFE229B72617EFDEC" box="[453,519,606,632]" pageId="2" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="0C663682EF45FFACFF709BAD6261FD0D" blockId="2.[151,519,606,665]" box="[151,280,641,665]" pageId="2" pageNumber="28">
(
<figureCitation id="94E22A07EF45FFACFF789BAD6269FD0D" box="[159,272,641,665]" captionStart-0="FIGURES 1 – 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURES 6 – 10" captionStart-4="FIGURES 11 – 15" captionStart-5="FIGURES 16 – 17" captionStart-6="FIGURES 18 – 21" captionStart-7="FIGURES 22 – 24" captionStartId-0="2.[151,264,1921,1944]" captionStartId-1="3.[151,250,541,564]" captionStartId-2="3.[151,250,1446,1469]" captionStartId-3="4.[151,264,1921,1944]" captionStartId-4="5.[151,264,1781,1804]" captionStartId-5="7.[151,264,1258,1281]" captionStartId-6="8.[151,264,1248,1271]" captionStartId-7="9.[151,264,1335,1358]" captionTargetBox-0="[151,1435,721,1900]" captionTargetBox-1="[151,1436,155,520]" captionTargetBox-2="[151,1436,886,1422]" captionTargetBox-3="[152,1404,552,1902]" captionTargetBox-4="[151,1435,155,1767]" captionTargetBox-5="[151,1435,155,1247]" captionTargetBox-6="[151,1435,181,1227]" captionTargetBox-7="[151,1435,157,1319]" captionTargetId-0="figure@2.[151,1436,721,1900]" captionTargetId-1="figure@3.[151,1436,155,520]" captionTargetId-2="figure@3.[151,1436,886,1422]" captionTargetId-3="figure@4.[151,1436,535,1903]" captionTargetId-4="figure@5.[151,1436,155,1767]" captionTargetId-5="figure@7.[151,1436,155,1249]" captionTargetId-6="figure@8.[151,1436,181,1227]" captionTargetId-7="figure@9.[151,1436,157,1319]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionText-0="FIGURES 1 – 3. Type locality and habitus of Ischyropsalis cantabrica sp. n. 1: Entrance to ‘ Rogeria Cave’ in Spain, Prov. Cantabria. 2: ‘ Rogeria Cave’ Digital Orthophoto with UTM grid, Zone 30, ED 50; 3: Life female specimen from ‘ Rogeria Cave’ (CGL 369)." captionText-1="FIGURE 4. Geographical map of central to northern Spain and surroundings of the sampling area (left). Ischyropsalis collection sites, with new sampling in original localities (right). Numbers refer to the following regions and species (Provinces are indicated in parentheses): 1, Western Coastal Cantabria inhabited by I. cantabrica sp. n.; 2, Upper Asón and Miera Mountains (Cantabria) inhabited by I. gigantea Dresco, 1968; 3, Triano-Galdames Mountains (Vizcaya) inhabited by I. magdalenae Simon, 1881; 4, Sierra de Aralar Massif (Guipúzcoa-Navarra) inhabited by I. navarrensis Roewer, 1950; 5, Sierra de Arno – Izarraitz Massif (Guipúzcoa) inhabited by I. galani nomen nudum; 6, Eastern part of the Basque-Cantabrian Mountains (inhabited by the complex of I. dispar Simon, 1872), including the localities of 6 a, Laredo – Limpias – Rasines – Guriezo Mountains (Cantabria), 6 b, Castro Urdiales (Cantabria), 6 c, Soncillo (Burgos), 6 d, Sierra Salvada Massif (Burgos – Álava), 6 e, Sierra de Arkamo Massif (Álava), 6 f – g, Gorbea Massif (Vizcaya), and 6 h, Sierra de Aizkorri Massif (Guipúzcoa). For cave names, coordinates and altitudes see Appendix 1." captionText-2="FIGURE 5. Map of the central region of the Cantabrian Mountain (northern Spain) to show the outcrops of the rock systems (in gray) which contain limestone and geographical situation of the study area with the distribution of the troglobitic species according to Luque (1991) and sampling of new localities (all species are allopatric): Ischyropsalis cantabrica sp. n.: Locus typicus ‘ Rogeria Cave’ (1), Portillo Cave (2), Cerviz Cave (3), Cáscaras Cave (4); Ischyropsalis gigantea (see Appendix 2 for explanation of numbers 5 – 12): Cayuela or Cañuela Cave (5), Currutuerta or Posadía Cave (6), Becerral Cave (7), Mortero Pit (8), Calleja Lavalle Pit (9), Los Moros del Río Munio Cave (10), Recuistro Cave (11), and Canal de la Torca nº 3 Pit, The Peña del Trillo Cave karst system (12); and Ischyropsalis sp. aff. dispar: La Baja Cave (13), Hoyo Molino Cave (14), Perelada Cave (15), Rejuyo or ‘ S. E. I. I. - 8 ’ Cave (16), Tornillos or RN- 30 Cave (17), La Lastrilla Cave (18), La Pasada Caves (19 – 20). Source: Geologic maps of Spain 1: 50,000 - scale corresponding to the Province of Cantabria (IGME)." captionText-3="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." captionText-4="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." captionText-5="FIGURES 16 – 17. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Male cheliceral apophysis on distal basal segment, medial view." captionText-6="FIGURES 18 – 21. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Detail of the femoral plate with cuticular structures on legs II, showing the typical microtrichia and isolated slit sense organ (18), a group of deep-circular pits on the cuticle (19), the ‘ holes’ of unknown function and identity (20) and an unidentified deep funnel-shaped depression surrounded by four spines (21)." captionText-7="FIGURES 22 – 24. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Detail of cuticular structures on the coxal plate of the legs II (22 a), showing the slit sense organ (black arrows), an unidentified deep funnelshaped depression surrounded by four spines (white arrow), several long hairs (macrotrichia), which arise from a basal socket (22), and other types of sculptural elements (Figs 23 – 24): e. g. granules, tubercles and cones with long macrotrichia on the apex, which arise from a basal socket." httpUri-0="https://zenodo.org/record/211032/files/figure.png" httpUri-1="https://zenodo.org/record/211033/files/figure.png" httpUri-2="https://zenodo.org/record/211034/files/figure.png" httpUri-3="https://zenodo.org/record/211035/files/figure.png" httpUri-4="https://zenodo.org/record/211036/files/figure.png" httpUri-5="https://zenodo.org/record/211037/files/figure.png" httpUri-6="https://zenodo.org/record/211038/files/figure.png" httpUri-7="https://zenodo.org/record/211039/files/figure.png" pageId="2" pageNumber="28">Figs 1–24</figureCitation>
)
</paragraph>
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<paragraph id="0C663682EF45FFACFF709EAD6229F848" blockId="2.[151,1437,1921,2012]" pageId="2" pageNumber="28">
<emphasis id="3EADEA90EF45FFACFF709EAD6244F803" bold="true" box="[151,317,1921,1944]" pageId="2" pageNumber="28">FIGURES 1–3.</emphasis>
Type locality and habitus of 
<taxonomicName id="CBD94D01EF45FFACFD939EAD6008F803" box="[628,881,1921,1943]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="28" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF45FFACFD939EAD6008F803" box="[628,881,1921,1943]" italics="true" pageId="2" pageNumber="28">Ischyropsalis cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF45FFACFC9E9EAE60CCF803" box="[889,949,1922,1943]" pageId="2" pageNumber="28" rank="species">
<emphasis id="3EADEA90EF45FFACFC9E9EAE60CCF803" bold="true" box="[889,949,1922,1943]" pageId="2" pageNumber="28">sp. n.</emphasis>
</taxonomicNameLabel>
1: Entrance to ‘Rogeria Cave’ in Spain, Prov. Cantabria. 2: ‘Rogeria Cave’ Digital Orthophoto with UTM grid, Zone 30, ED50; 3: Life female specimen from ‘Rogeria Cave’ (CGL 369).
</paragraph>
</caption>
<caption id="58A6660AEF44FFADFF709B316763FCD3" httpUri="https://zenodo.org/record/211033/files/figure.png" pageId="3" pageNumber="29" targetBox="[151,1436,155,520]" targetPageId="3">
<paragraph id="0C663682EF44FFADFF709B316763FCD3" blockId="3.[151,1437,541,840]" pageId="3" pageNumber="29">
<emphasis id="3EADEA90EF44FFADFF709B316261FDA7" bold="true" box="[151,280,541,564]" pageId="3" pageNumber="29">FIGURE 4.</emphasis>
Geographical map of central to northern Spain and surroundings of the sampling area (left). 
<taxonomicName id="CBD94D01EF44FFADFAF19B3266E5FDA0" box="[1302,1436,542,564]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="3EADEA90EF44FFADFAF19B3266E5FDA0" box="[1302,1436,542,564]" italics="true" pageId="3" pageNumber="29">Ischyropsalis</emphasis>
</taxonomicName>
collection sites, with new sampling in original localities (right). Numbers refer to the following regions and species (Provinces are indicated in parentheses): 
<emphasis id="3EADEA90EF44FFADFE3F9B77629CFDE4" bold="true" box="[472,485,603,624]" pageId="3" pageNumber="29">1</emphasis>
, Western Coastal Cantabria inhabited by 
<taxonomicName id="CBD94D01EF44FFADFC429B706755FDE5" box="[933,1068,603,625]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF44FFADFC429B706755FDE5" box="[933,1068,603,625]" italics="true" pageId="3" pageNumber="29">I. cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF44FFADFBD09B70670CFDE5" box="[1079,1141,604,625]" pageId="3" pageNumber="29" rank="species">
<emphasis id="3EADEA90EF44FFADFBD09B70670CFDE5" bold="true" box="[1079,1141,604,625]" pageId="3" pageNumber="29">sp. n.</emphasis>
</taxonomicNameLabel>
; 
<emphasis id="3EADEA90EF44FFADFB609B7767EDFDE4" bold="true" box="[1159,1172,603,624]" pageId="3" pageNumber="29">2</emphasis>
, Upper Asón and Miera Mountains (Cantabria) inhabited by 
<taxonomicName id="CBD94D01EF44FFADFDFA9B576055FD04" authority="Dresco, 1968" authorityName="Dresco" authorityYear="1968" box="[541,812,634,656]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="3EADEA90EF44FFADFDFA9B5761E8FD04" box="[541,657,635,656]" italics="true" pageId="3" pageNumber="29">I. gigantea</emphasis>
Dresco, 1968
</taxonomicName>
; 
<emphasis id="3EADEA90EF44FFADFCDB9B566030FD1B" bold="true" box="[828,841,634,655]" pageId="3" pageNumber="29">3</emphasis>
, Triano-Galdames Mountains (Vizcaya) inhabited by 
<taxonomicName id="CBD94D01EF44FFADFA6A9B5762E4FD3A" authority="Simon, 1881" authorityName="Simon" authorityYear="1881" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="magdalenae">
<emphasis id="3EADEA90EF44FFADFA6A9B57626AFD3A" italics="true" pageId="3" pageNumber="29">I. magdalenae</emphasis>
Simon, 1881
</taxonomicName>
; 
<emphasis id="3EADEA90EF44FFADFE4F9BB562CCFD3A" bold="true" box="[424,437,665,686]" pageId="3" pageNumber="29">4</emphasis>
, Sierra de Aralar Massif (Guipúzcoa-Navarra) inhabited by 
<taxonomicName id="CBD94D01EF44FFADFBFE9BB66645FD3A" authority="Roewer, 1950" authorityName="Roewer" authorityYear="1950" box="[1049,1340,664,687]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="navarrensis">
<emphasis id="3EADEA90EF44FFADFBFE9BB667DCFD3B" box="[1049,1189,666,687]" italics="true" pageId="3" pageNumber="29">I. navarrensis</emphasis>
Roewer, 1950
</taxonomicName>
; 
<emphasis id="3EADEA90EF44FFADFAA19BB5662AFD3A" bold="true" box="[1350,1363,665,686]" pageId="3" pageNumber="29">5</emphasis>
, Sierra de Arno–Izarraitz Massif (Guipúzcoa) inhabited by 
<taxonomicName id="CBD94D01EF44FFADFD2C9B94605EFD59" box="[715,807,695,717]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="galani">
<emphasis id="3EADEA90EF44FFADFD2C9B94605EFD59" box="[715,807,695,717]" italics="true" pageId="3" pageNumber="29">I. galani</emphasis>
</taxonomicName>
nomen nudum; 
<emphasis id="3EADEA90EF44FFADFC079B9B6094FD58" bold="true" box="[992,1005,695,716]" pageId="3" pageNumber="29">6</emphasis>
, Eastern part of the Basque-Cantabrian Mountains (inhabited by the complex of 
<taxonomicName id="CBD94D01EF44FFADFD249BFB6092FD78" authority="Simon, 1872" authorityName="Simon" authorityYear="1872" box="[707,1003,726,748]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="3EADEA90EF44FFADFD249BFB6048FD78" box="[707,817,726,748]" italics="true" pageId="3" pageNumber="29">I. dispar</emphasis>
Simon, 1872
</taxonomicName>
), including the localities of 
<emphasis id="3EADEA90EF44FFADFA9B9BFA66EFFD7F" bold="true" box="[1404,1430,726,747]" pageId="3" pageNumber="29">6a</emphasis>
, Laredo–Limpias–Rasines–Guriezo Mountains (Cantabria), 
<emphasis id="3EADEA90EF44FFADFD119BD96068FC9E" bold="true" box="[758,785,757,778]" pageId="3" pageNumber="29">6b</emphasis>
, Castro Urdiales (Cantabria), 
<emphasis id="3EADEA90EF44FFADFBAC9BD9671BFC9E" bold="true" box="[1099,1122,757,778]" pageId="3" pageNumber="29">6c</emphasis>
, Soncillo (Burgos), 
<emphasis id="3EADEA90EF44FFADFAD19BD96629FC9E" bold="true" box="[1334,1360,757,778]" pageId="3" pageNumber="29">6d</emphasis>
, Sierra Salvada Massif (Burgos–Álava), 
<emphasis id="3EADEA90EF44FFADFE149A3F6172FCBC" bold="true" box="[499,523,787,808]" pageId="3" pageNumber="29">6e</emphasis>
, Sierra de Arkamo Massif (Álava), 
<emphasis id="3EADEA90EF44FFADFC699A3F60C5FCBD" bold="true" box="[910,956,787,810]" pageId="3" pageNumber="29">6f–g</emphasis>
, Gorbea Massif (Vizcaya), and 
<emphasis id="3EADEA90EF44FFADFAF59A3F6655FCBC" bold="true" box="[1298,1324,787,808]" pageId="3" pageNumber="29">6h</emphasis>
, Sierra de Aizkorri Massif (Guipúzcoa). For cave names, coordinates and altitudes see Appendix 1.
</paragraph>
</caption>
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<paragraph id="0C663682EF44FFADFF709C8A6746F927" blockId="3.[151,1437,1446,1715]" pageId="3" pageNumber="29">
<emphasis id="3EADEA90EF44FFADFF709C8A626AFA28" bold="true" box="[151,275,1446,1469]" pageId="3" pageNumber="29">FIGURE 5.</emphasis>
Map of the central region of the Cantabrian Mountain (northern Spain) to show the outcrops of the rock systems (in gray) which contain limestone and geographical situation of the study area with the distribution of the troglobitic species according to Luque (1991) and sampling of new localities (all species are allopatric): 
<taxonomicName id="CBD94D01EF44FFADFBF49CC86676FA6E" box="[1043,1295,1508,1530]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF44FFADFBF49CC86676FA6E" box="[1043,1295,1508,1530]" italics="true" pageId="3" pageNumber="29">Ischyropsalis cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF44FFADFAF09CC96628FA6E" box="[1303,1361,1509,1530]" pageId="3" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
: Locus typicus ‘Rogeria Cave’ (1), Portillo Cave (2), Cerviz Cave (3), Cáscaras Cave (4); 
<taxonomicName id="CBD94D01EF44FFADFC039F2F67B3F98D" box="[996,1226,1539,1561]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="3EADEA90EF44FFADFC039F2F67B3F98D" box="[996,1226,1539,1561]" italics="true" pageId="3" pageNumber="29">Ischyropsalis gigantea</emphasis>
</taxonomicName>
(see Appendix 2 for explanation of numbers 5–12): Cayuela or Cañuela Cave (5), Currutuerta or Posadía Cave (6), Becerral Cave (7), Mortero Pit (8), Calleja Lavalle Pit (9), Los Moros del Río Munio Cave (10), Recuistro Cave (11), and Canal de la Torca nº 3 Pit, The Peña del Trillo Cave karst system (12); and 
<taxonomicName id="CBD94D01EF44FFADFDFB9F7361DBF9E1" box="[540,674,1631,1653]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="3EADEA90EF44FFADFDFB9F7361DBF9E1" box="[540,674,1631,1653]" italics="true" pageId="3" pageNumber="29">Ischyropsalis</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="CBD94D01EF44FFADFD2B9F7360B5F9E0" authority="La Baja Cave" authorityName="La Baja Cave" box="[716,972,1631,1653]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="29" phylum="Arthropoda" rank="species" species="dispar">
aff. 
<emphasis id="3EADEA90EF44FFADFD139F73604DF9E1" box="[756,820,1631,1653]" italics="true" pageId="3" pageNumber="29">dispar</emphasis>
: La Baja Cave
</taxonomicName>
(13), Hoyo Molino Cave (14), Perelada Cave (15), Rejuyo or ‘S.E.I.I.-8’ Cave (16), Tornillos or RN-30 Cave (17), La Lastrilla Cave (18), La Pasada Caves (19–20). Source: Geologic maps of Spain 1:50,000-scale corresponding to the Province of Cantabria (IGME).
</paragraph>
</caption>
<subSubSection id="44C36509EF44FFAAFF709E2F673CFF40" lastPageId="4" lastPageNumber="30" pageId="3" pageNumber="29" type="materials_examined">
<paragraph id="0C663682EF44FFADFF709E2F67FDF867" blockId="3.[151,1436,1794,2035]" pageId="3" pageNumber="29">
<emphasis id="3EADEA90EF44FFADFF709E2F62C5F88F" bold="true" box="[151,444,1794,1819]" pageId="3" pageNumber="29">
<typeStatus id="D3628820EF44FFADFF709E2F63A8F88F" box="[151,209,1795,1819]" pageId="3" pageNumber="29">Type</typeStatus>
material: 
<typeStatus id="D3628820EF44FFADFEA89E2F62C5F88F" box="[335,444,1795,1819]" pageId="3" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
3 (
<collectionCode id="6AC8AE47EF44FFADFE0C9E2F6138F88F" LSID="urn:lsid:biocol.org:col:35275" box="[491,577,1795,1819]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="29">MNCN</collectionCode>
20.02/12828) and 
<emphasis id="3EADEA90EF44FFADFCFC9E2E60F9F88F" bold="true" box="[795,896,1794,1819]" pageId="3" pageNumber="29">Allotype</emphasis>
Ƥ (
<collectionCode id="6AC8AE47EF44FFADFC489E2F677CF88F" LSID="urn:lsid:biocol.org:col:35275" box="[943,1029,1795,1819]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="29">MNCN</collectionCode>
20.02/12829): 
<emphasis id="3EADEA90EF44FFADFB529E2E66E5F88F" bold="true" box="[1205,1436,1794,1819]" pageId="3" pageNumber="29">
<collectingCountry id="74CE7612EF44FFADFB529E2E6670F88F" box="[1205,1289,1794,1819]" name="Spain" pageId="3" pageNumber="29">SPAIN</collectingCountry>
, Cantabria:
</emphasis>
Alfoz de Lloredo, Oreña: Rogeria Cave (30TVP0867005271, 
<quantity id="CB219B67EF44FFADFC899E0B60D6F8AB" box="[878,943,1831,1855]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="29" unit="m" value="90.0">90 m</quantity>
), 
<date id="78671042EF44FFADFC2F9E0B6747F8AB" box="[968,1086,1831,1855]" pageId="3" pageNumber="29" value="1994-04-02">2-IV-1994</date>
, leg. C.G. Luque. 
<emphasis id="3EADEA90EF44FFADFAFC9E0B6206F8F7" bold="true" pageId="3" pageNumber="29">
<typeStatus id="D3628820EF44FFADFAFC9E0B66EEF8AB" box="[1307,1431,1831,1855]" pageId="3" pageNumber="29" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="74CE7612EF44FFADFF709E666392F8F7" box="[151,235,1866,1891]" name="Spain" pageId="3" pageNumber="29">SPAIN</collectingCountry>
, Cantabria:
</emphasis>
Alfoz de Lloredo: 5 3, 4 ƤƤ, same data as the 
<typeStatus id="D3628820EF44FFADFC3D9E676745F8F7" box="[986,1084,1867,1891]" pageId="3" pageNumber="29" type="holotype">holotype</typeStatus>
(1 3, 1 Ƥ 
<collectionCode id="6AC8AE47EF44FFADFB2F9E676659F8F7" LSID="urn:lsid:biocol.org:col:35275" box="[1224,1312,1867,1891]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="29">MNCN</collectionCode>
; 1 3, 1 Ƥ 
<collectionCode id="6AC8AE47EF44FFADFF709E5C638AF813" box="[151,243,1904,1927]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="29">NHMW</collectionCode>
, 1 3, 1 Ƥ 
<collectionCode id="6AC8AE47EF44FFADFE8B9E4362C1F813" box="[364,440,1903,1927]" pageId="3" pageNumber="29">AXLS</collectionCode>
, the rest CGL); Ruiloba, Casasola: 1 3, 1 Ƥ, Portillo Cave (30TVP9639204775, 20 m), 
<date id="78671042EF44FFADFF709EBF6265F83F" box="[151,284,1939,1963]" pageId="3" pageNumber="29" value="1995-03-25">25-III-1995</date>
, leg. C.G. Luque &amp; E. Muñoz (CGL); Cabezón de la Sal, Bustablado: 1 3 mounted on SEM stubs (
<collectionCode id="6AC8AE47EF44FFADFF799E9B638DF85B" LSID="urn:lsid:biocol.org:col:35275" box="[158,244,1975,1999]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="3" pageNumber="29">MNCN</collectionCode>
20.02/12857), Cerviz Cave (30TVP0286800238, 
<quantity id="CB219B67EF44FFADFCC09E9B6017F85B" box="[807,878,1975,1999]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="3" pageNumber="29" unit="m" value="365.0">365 m</quantity>
), 
<date id="78671042EF44FFADFC619E9B6085F85B" box="[902,1020,1975,1999]" pageId="3" pageNumber="29" value="1995-04-01">1-IV-1995</date>
, leg. C.G. Luque &amp; J. Ruiz; Alfoz de Lloredo, Oreña: 2 3, Calero Cave, 
<date id="78671042EF44FFADFDD09EF761A8F867" box="[567,721,2011,2035]" pageId="3" pageNumber="29" value="2003-08-12">12-VIII-2003</date>
, leg. L. Labrada &amp; C.G. Luque (CGL).
</paragraph>
<paragraph id="0C663682EF43FFAAFF2099BB673CFF40" blockId="4.[151,1436,151,500]" pageId="4" pageNumber="30">
<emphasis id="3EADEA90EF43FFAAFF2099BB607DFF24" bold="true" box="[199,772,151,176]" pageId="4" pageNumber="30">
Additional material studied: 
<collectingCountry id="74CE7612EF43FFAAFDF899BB6108FF24" box="[543,625,151,176]" name="Spain" pageId="4" pageNumber="30">SPAIN</collectingCountry>
, Cantabria:
</emphasis>
Udías, Canales: 1 Ƥ (
<collectionCode id="6AC8AE47EF43FFAAFC1F99BB6737FF3B" LSID="urn:lsid:biocol.org:col:35275" box="[1016,1102,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="4" pageNumber="30">MNCN</collectionCode>
20.02/12863), Cáscaras Cave (30TVP 
<quantity id="CB219B67EF43FFAAFF08999062ACFF40" box="[239,469,188,212]" metricMagnitude="12" metricUnit="m" metricValue="9.894201878130001" pageId="4" pageNumber="30" unit="m" value="9.89420187813E12">9894201878, 130 m</quantity>
), 
<date id="78671042EF43FFAAFE0E9990611BFF40" box="[489,610,188,212]" pageId="4" pageNumber="30" value="1915-09-02">2-IX-1915</date>
, leg. C. Bolívar (see 
<bibRefCitation id="68484B73EF43FFAAFCAA9990609AFF40" author="Bolivar" box="[845,995,188,212]" pageId="4" pageNumber="30" refString="Bolivar, C. (1916) Exploracion entomologica de algunas cuevas de la region cantabrica. Boletin de la Real Sociedad Espanola de Historia Natural, 16, 315 - 322." type="journal article" year="1916">Bolívar 1916</bibRefCitation>
, p. 319).
</paragraph>
</subSubSection>
<subSubSection id="44C36509EF43FFAAFF2099F36160FE60" pageId="4" pageNumber="30" type="diagnosis">
<paragraph id="0C663682EF43FFAAFF2099F36160FE60" blockId="4.[151,1436,151,500]" pageId="4" pageNumber="30">
<emphasis id="3EADEA90EF43FFAAFF2099F3623AFF6C" bold="true" box="[199,323,223,248]" pageId="4" pageNumber="30">Diagnosis:</emphasis>
A species of the 
<taxonomicName id="CBD94D01EF43FFAAFDFC99CC611FFF6C" box="[539,614,224,248]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="3EADEA90EF43FFAAFDFC99CC611FFF6C" box="[539,614,224,248]" italics="true" pageId="4" pageNumber="30">kollari</emphasis>
</taxonomicName>
group (
<emphasis id="3EADEA90EF43FFAAFD2099CD607CFF6C" box="[711,773,225,248]" italics="true" pageId="4" pageNumber="30">sensu</emphasis>
<bibRefCitation id="68484B73EF43FFAAFCF699CC60C8FF63" author="Martens" box="[785,945,223,247]" pageId="4" pageNumber="30" refString="Martens, J. (1969) Die Abgrenzung von Biospezies auf biologisch-etologischer und morphologischer Grundlage am Beispiel der Gattung Ischyropsalis C. L. Koch 1839 (Opiliones, Ischyropsalididae). Zoologische Jahrbuecher Abteilung fur Systematik, 96, 133 - 264." type="journal article" year="1969">Martens 1969</bibRefCitation>
) with long and slender chelicerae (spines completely absent in males); cheliceral apophysis high, not projected forwards, with rounded conical shape and the piliferous apex, with a series of deep parallel grooves, medial and characterized by an additional small piliferous and pocket-forming apophysis; dorsal region of the glans penis with a dense mass of very long bristles at its end; glans short and conical shaped at its apex; upper part of glans rounded-quadrangular shaped in cross-section; sclerite of glans long and narrow, almost parallel-sided, basal end tapering, median keel developed, bristle cover dense, widely separated into two lateral areas; body depigmented but scutum parvum uniformly dark colored (dorsal abdominal sclerites fused).
</paragraph>
</subSubSection>
<caption id="58A6660AEF43FFAAFF709EAD60EFF842" httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="4" pageNumber="30" targetBox="[152,1404,552,1902]" targetPageId="4">
<paragraph id="0C663682EF43FFAAFF709EAD60EFF842" blockId="4.[151,1436,1921,2006]" pageId="4" pageNumber="30">
<emphasis id="3EADEA90EF43FFAAFF709EAD6230F803" bold="true" box="[151,329,1921,1944]" pageId="4" pageNumber="30">FIGURES 6–10.</emphasis>
<taxonomicName id="CBD94D01EF43FFAAFEB59EAE6137F80C" box="[338,590,1922,1944]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="30" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF43FFAAFEB59EAE6137F80C" box="[338,590,1922,1944]" italics="true" pageId="4" pageNumber="30">Ischyropsalis cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF43FFAAFDB09EAE61EAF803" box="[599,659,1922,1943]" pageId="4" pageNumber="30" rank="species">
<emphasis id="3EADEA90EF43FFAAFDB09EAE61EAF803" bold="true" box="[599,659,1922,1943]" pageId="4" pageNumber="30">sp. n.</emphasis>
</taxonomicNameLabel>
, male holotype from Rogeria Cave (MNCN 20.02/12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8a, 9), female chelicerae (8b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7).
</paragraph>
</caption>
<caption id="58A6660AEF42FFABFF709FD9626EF8DC" httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="5" pageNumber="31" targetBox="[151,1435,155,1767]" targetPageId="5">
<paragraph id="0C663682EF42FFABFF709FD9626EF8DC" blockId="5.[151,1436,1781,1864]" pageId="5" pageNumber="31">
<emphasis id="3EADEA90EF42FFABFF709FD9622BF89E" bold="true" box="[151,338,1781,1804]" pageId="5" pageNumber="31">FIGURES 11–15.</emphasis>
<taxonomicName id="CBD94D01EF42FFABFEBF9FD9612BF89F" box="[344,594,1781,1803]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="31" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF42FFABFEBF9FD9612BF89F" box="[344,594,1781,1803]" italics="true" pageId="5" pageNumber="31">Ischyropsalis cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF42FFABFDBF9FDA61E9F89F" box="[600,656,1782,1803]" pageId="5" pageNumber="31" rank="species">
<emphasis id="3EADEA90EF42FFABFDBF9FDA61E9F89F" bold="true" box="[600,656,1782,1803]" pageId="5" pageNumber="31">sp. n.</emphasis>
</taxonomicNameLabel>
, male paratype from Cerviz Cave (MNCN 20.02/12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15).
</paragraph>
</caption>
<subSubSection id="44C36509EF42FFA8FF209E4362B4F969" lastPageId="6" lastPageNumber="32" pageId="5" pageNumber="31" type="description">
<paragraph id="0C663682EF42FFABFF209E436222F81C" blockId="5.[151,1436,1903,2037]" box="[199,347,1903,1928]" pageId="5" pageNumber="31">
<emphasis id="3EADEA90EF42FFABFF209E436222F81C" bold="true" box="[199,347,1903,1928]" pageId="5" pageNumber="31">Description:</emphasis>
</paragraph>
<paragraph id="0C663682EF42FFA8FF209EB860A0FE38" blockId="5.[151,1436,1903,2037]" lastBlockId="6.[151,1437,151,1963]" lastPageId="6" lastPageNumber="32" pageId="5" pageNumber="31">
<emphasis id="3EADEA90EF42FFABFF209EB86209F83F" bold="true" box="[199,368,1940,1963]" italics="true" pageId="5" pageNumber="31">Measurements</emphasis>
were taken for the 
<typeStatus id="D3628820EF42FFABFDBB9EB861B8F838" box="[604,705,1940,1964]" pageId="5" pageNumber="31" type="holotype">holotype</typeStatus>
, allotype and 
<typeStatus id="D3628820EF42FFABFC8E9EB960ACF839" box="[873,981,1941,1965]" pageId="5" pageNumber="31" type="paratype">paratypes</typeStatus>
(9 3 and 6 ƤƤ). Body length (with distended abdomen): 
<typeStatus id="D3628820EF42FFABFE6A9E946296F844" box="[397,495,1976,2000]" pageId="5" pageNumber="31" type="holotype">holotype</typeStatus>
5.01, 3 
<typeStatus id="D3628820EF42FFABFD889E9561A3F845" box="[623,730,1977,2001]" pageId="5" pageNumber="31" type="paratype">paratypes</typeStatus>
5.01–5.45, allotype 5.58, ƤƤ 
<typeStatus id="D3628820EF42FFABFBD99E9567D0F845" box="[1086,1193,1977,2001]" pageId="5" pageNumber="31" type="paratype">paratypes</typeStatus>
5.56–5.81; length of basal cheliceral segment (in parentheses, with length/width of distal cheliceral segment): 
<typeStatus id="D3628820EF42FFABFB9E9EF067A2F860" box="[1145,1243,2012,2036]" pageId="5" pageNumber="31" type="holotype">holotype</typeStatus>
5.20 (6.50/0.80), 3 
<typeStatus id="D3628820EF41FFA8FF2C99B4624FFF24" box="[203,310,152,176]" pageId="6" pageNumber="32" type="paratype">paratypes</typeStatus>
5.20–5.54 (6.50–6.88/0.80), allotype 5.53 (7.25/0.84), ƤƤ 
<typeStatus id="D3628820EF41FFA8FC0D99B4672CFF24" box="[1002,1109,152,176]" pageId="6" pageNumber="32" type="paratype">paratypes</typeStatus>
5.53–5.85 (7.25–7.68/0.84). Female show a wider range in body length due to different stages of gravidity but their chelicerae are less variable in size than in male. Total length of leg II of male 
<typeStatus id="D3628820EF41FFA8FCED99F36015FF63" box="[778,876,223,247]" pageId="6" pageNumber="32" type="holotype">holotype</typeStatus>
(
<typeStatus id="D3628820EF41FFA8FC6799F36094FF63" box="[896,1005,223,247]" pageId="6" pageNumber="32" type="paratype">paratypes</typeStatus>
in parentheses) and female allotype (
<typeStatus id="D3628820EF41FFA8FF7998286272FE88" box="[158,267,260,284]" pageId="6" pageNumber="32" type="paratype">paratypes</typeStatus>
in parentheses): 42.49 (42.49–44.82), 42.57 (42.57–43.78). Length of the II segments (in parentheses as before): femur: 8.82 (8.82–8.98), 9.32 (9.32–9.66); patella: 1.36 (1.36–1.41), 1.49 (1.49–1.56); tibia: 8.38 (8.38–8.94), 8.41 (8.41–8.78); metatarsus: 13.52 (13.52–13.89), 13.40 (13.40–13.81); tarsus: 10.41 (10.41–10.82), 9.95 (9.95–10.21). The measurements of the male (
<typeStatus id="D3628820EF41FFA8FD3B98436038FE13" box="[732,833,367,391]" pageId="6" pageNumber="32" type="holotype">holotype</typeStatus>
) and female (in parentheses) femurs from the first to the fourth pair of legs are 6.45 (6.84), 8.82 (9.32), 5.34 (5.56), 7.08 (7.35).
</paragraph>
<paragraph id="0C663682EF41FFA8FF20989B6195FC15" blockId="6.[151,1437,151,1963]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF20989B62C9FE44" bold="true" box="[199,432,439,464]" italics="true" pageId="6" pageNumber="32">Genital morphology</emphasis>
(
<figureCitation id="94E22A07EF41FFA8FE27989B615EFE5B" box="[448,551,439,463]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Figs 6–7</figureCitation>
, 
<figureCitation id="94E22A07EF41FFA8FDD2989B6104FE5B" box="[565,637,439,463]" captionStart="FIGURES 11 – 15" captionStartId="5.[151,264,1781,1804]" captionTargetBox="[151,1435,155,1767]" captionTargetId="figure@5.[151,1436,155,1767]" captionTargetPageId="5" captionText="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="6" pageNumber="32">11–15</figureCitation>
). 
<emphasis id="3EADEA90EF41FFA8FD74989B61AAFE44" bold="true" box="[659,723,439,464]" pageId="6" pageNumber="32">Penis</emphasis>
: Symmetrical, with robust truncus and the base long and split (
<figureCitation id="94E22A07EF41FFA8FF7898F5639DFE65" box="[159,228,473,497]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Fig. 6</figureCitation>
); the truncus is of medium length (L: 4.0). The coloration is dark, from the truncus to the base, the ventral side is depigmented; the glans in contrast is blackish in colour. The glans at its apex is short conical shaped, with a long stylus (
<figureCitation id="94E22A07EF41FFA8FEF89B306211FDA0" box="[287,360,540,564]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Figs 7</figureCitation>
, 
<figureCitation id="94E22A07EF41FFA8FE939B3062EAFDA0" box="[372,403,540,564]" captionStart="FIGURES 11 – 15" captionStartId="5.[151,264,1781,1804]" captionTargetBox="[151,1435,155,1767]" captionTargetId="figure@5.[151,1436,155,1767]" captionTargetPageId="5" captionText="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="6" pageNumber="32">13</figureCitation>
). On the dorsal side (
<figureCitation id="94E22A07EF41FFA8FD669B3061B5FDA0" box="[641,716,540,564]" captionStart="FIGURES 11 – 15" captionStartId="5.[151,264,1781,1804]" captionTargetBox="[151,1435,155,1767]" captionTargetId="figure@5.[151,1436,155,1767]" captionTargetPageId="5" captionText="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="6" pageNumber="32">Fig 11</figureCitation>
), the glans has a dense mass of very long bristles at its ends (the bristles become shorter nearer the apex of the glans). In the upper part of glans, the outline of the cross section is rounded-quadrangular shaped (
<figureCitation id="94E22A07EF41FFA8FE139B726147FDE2" box="[500,574,606,630]" captionStart="FIGURES 11 – 15" captionStartId="5.[151,264,1781,1804]" captionTargetBox="[151,1435,155,1767]" captionTargetId="figure@5.[151,1436,155,1767]" captionTargetPageId="5" captionText="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="6" pageNumber="32">Fig 11</figureCitation>
) and does not narrow gradually to acquire a triangular shape in lateral view. In contrast, the ventral side displays a sclerite of glans long and narrow, almost parallel-sided, basal end tapering, ventral median keel developed, bristle cover dense, widely separated into two lateral areas (
<figureCitation id="94E22A07EF41FFA8FB509B8D667FFD2D" box="[1207,1286,673,697]" captionStart="FIGURES 11 – 15" captionStartId="5.[151,264,1781,1804]" captionTargetBox="[151,1435,155,1767]" captionTargetId="figure@5.[151,1436,155,1767]" captionTargetPageId="5" captionText="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="6" pageNumber="32">Fig 12</figureCitation>
), and a large depigmented mouth indicating the attached muscular tendon (
<figureCitation id="94E22A07EF41FFA8FCA89BEE60E4FD4E" box="[847,925,706,730]" captionStart="FIGURES 11 – 15" captionStartId="5.[151,264,1781,1804]" captionTargetBox="[151,1435,155,1767]" captionTargetId="figure@5.[151,1436,155,1767]" captionTargetPageId="5" captionText="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="6" pageNumber="32">Fig 15</figureCitation>
). The glans is clearly inflated in lateral view, appearing swollen in mid-section. This spaces the ventro-lateral bristle areas wide apart from each other. The midarea has a thin longitudinal row of short bristles (
<figureCitation id="94E22A07EF41FFA8FD0C9A29600BFC89" box="[747,882,773,797]" captionStart="FIGURES 11 – 15" captionStartId="5.[151,264,1781,1804]" captionTargetBox="[151,1435,155,1767]" captionTargetId="figure@5.[151,1436,155,1767]" captionTargetPageId="5" captionText="FIGURES 11 – 15. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Distal part of penis with glans and stylus: dorsal view (11), ventral view (12), lateral view (13), dorsolateral view (14), detail of the funnel-like sclerite (15)." httpUri="https://zenodo.org/record/211036/files/figure.png" pageId="6" pageNumber="32">Figs 13–14</figureCitation>
). 
<emphasis id="3EADEA90EF41FFA8FC6C9A296773FC8A" bold="true" box="[907,1034,773,798]" pageId="6" pageNumber="32">Ovipositor</emphasis>
: bilobulate, with the ventral apex covered with few hairs; the seminal receptacle has four short tubes. In this respect not differentiated from other 
<taxonomicName id="CBD94D01EF41FFA8FF709A646255FCF4" box="[151,300,840,864]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="3EADEA90EF41FFA8FF709A646255FCF4" box="[151,300,840,864]" italics="true" pageId="6" pageNumber="32">Ischyropsalis</emphasis>
</taxonomicName>
species (
<bibRefCitation id="68484B73EF41FFA8FE779A6B6156FCCB" author="Martens" box="[400,559,839,863]" pageId="6" pageNumber="32" refString="Martens, J. (1969) Die Abgrenzung von Biospezies auf biologisch-etologischer und morphologischer Grundlage am Beispiel der Gattung Ischyropsalis C. L. Koch 1839 (Opiliones, Ischyropsalididae). Zoologische Jahrbuecher Abteilung fur Systematik, 96, 133 - 264." type="journal article" year="1969">Martens 1969</bibRefCitation>
; 
<bibRefCitation id="68484B73EF41FFA8FDDD9A6461AEFCCB" author="Dresco" box="[570,727,839,863]" pageId="6" pageNumber="32" refString="Dresco, E. (1968 a) Recherches sur les Opilions du genre Ischyropsalis (Fam. Ischyropsalidae). VII. Ischyropsalis gigantea sp. nov. Bulletin du Museum Natural d'Historie Naturalle, 2 ser., 40 (2), 308 - 314." type="journal article" year="1968" yearSuffix="a">Dresco 1968a</bibRefCitation>
, p. 313, fig. 19; 
<bibRefCitation id="68484B73EF41FFA8FC6C9A646753FCCB" author="Dresco" box="[907,1066,839,863]" pageId="6" pageNumber="32" refString="Dresco, E. (1968 b) Recherches sur les Opilions du genre Ischyropsalis (Fam. Ischyropsalidae). IX. Ischyropsalis espagnoli sp. nov. Bulletin du Museum Natural d'Historie Naturalle, 2 ser., 40 (5), 962 - 967." type="journal article" year="1968" yearSuffix="b">Dresco 1968b</bibRefCitation>
, p. 966, fig. 20; 
<bibRefCitation id="68484B73EF41FFA8FB3A9A646605FCCB" author="Dresco" box="[1245,1404,839,863]" pageId="6" pageNumber="32" refString="Dresco, E. (1972 b) Recherches sur les Opilions du genre Ischyropsalis (Fam. Ischyropsalidae). XII. Ischyropsalis dispar Simon. Annales de Speleologie, 27 (2), 351 - 361." type="journal article" year="1972" yearSuffix="b">Dresco 1972b</bibRefCitation>
, p. 356, fig. 25; see also 
<bibRefCitation id="68484B73EF41FFA8FE629A45616EFC15" author="Gruber" box="[389,535,873,897]" pageId="6" pageNumber="32" refString="Gruber, J. (2007) Ischyropsalididae Simon, 1879. In: R. Pinto-da-Rocha, G. Machado &amp; G. Giribet (Eds), Harvestmen: The Biology of Opiliones. Harvard University Press, Cambridge, 145 - 148." type="book chapter" year="2007">Gruber 2007</bibRefCitation>
, p. 147, fig. 4.15i).
</paragraph>
<paragraph id="0C663682EF41FFA8FF209AA66153FA9E" blockId="6.[151,1437,151,1963]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209AA6623DFC37" bold="true" box="[199,324,906,931]" italics="true" pageId="6" pageNumber="32">Chelicerae</emphasis>
(
<figureCitation id="94E22A07EF41FFA8FEB49AA662C0FC36" box="[339,441,906,930]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Figs 8–9</figureCitation>
, 
<figureCitation id="94E22A07EF41FFA8FE219AA66169FC36" box="[454,528,906,930]" captionStart="FIGURES 16 – 17" captionStartId="7.[151,264,1258,1281]" captionTargetBox="[151,1435,155,1247]" captionTargetId="figure@7.[151,1436,155,1249]" captionTargetPageId="7" captionText="FIGURES 16 – 17. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Male cheliceral apophysis on distal basal segment, medial view." httpUri="https://zenodo.org/record/211037/files/figure.png" pageId="6" pageNumber="32">16–17</figureCitation>
): Sexually dimorph. In general, long and slender, black or dark brown. In males basal segment with a longitudinal row of small and flat spines with bristles ventrally and with 4 or 5 piliferous tubercles dorsally (
<figureCitation id="94E22A07EF41FFA8FE919AFE62B8FC7E" box="[374,449,978,1002]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Figs 8</figureCitation>
a, 9). Basal segment narrow at the base, widens distally to a pronounced triangular apophysis (lateral view; 
<figureCitation id="94E22A07EF41FFA8FE4D9ADA6296FB9A" box="[426,495,1014,1038]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Figs 8</figureCitation>
a, 9, 16–17). The apophysis does not project proximal but dorsal, and has a rounded conical shape with the piliferous apex, with a piliferous pocket-forming apophysis and a series of deep parallel grooves on the inner side (
<figureCitation id="94E22A07EF41FFA8FE279D126171FBC2" box="[448,520,1086,1110]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Figs 9</figureCitation>
, 
<figureCitation id="94E22A07EF41FFA8FDF39D12614AFBC2" box="[532,563,1086,1110]" captionStart="FIGURES 16 – 17" captionStartId="7.[151,264,1258,1281]" captionTargetBox="[151,1435,155,1247]" captionTargetId="figure@7.[151,1436,155,1249]" captionTargetPageId="7" captionText="FIGURES 16 – 17. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Male cheliceral apophysis on distal basal segment, medial view." httpUri="https://zenodo.org/record/211037/files/figure.png" pageId="6" pageNumber="32">17</figureCitation>
). The distal segment is completely smooth, only few short hairs present at the base (
<figureCitation id="94E22A07EF41FFA8FF3C9D4E625FFBEE" box="[219,294,1122,1146]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Figs 8</figureCitation>
, 
<figureCitation id="94E22A07EF41FFA8FED19D4E6206FBEE" box="[310,383,1122,1146]" captionStart="FIGURES 16 – 17" captionStartId="7.[151,264,1258,1281]" captionTargetBox="[151,1435,155,1247]" captionTargetId="figure@7.[151,1436,155,1249]" captionTargetPageId="7" captionText="FIGURES 16 – 17. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Male cheliceral apophysis on distal basal segment, medial view." httpUri="https://zenodo.org/record/211037/files/figure.png" pageId="6" pageNumber="32">16–17</figureCitation>
). They are generally bristle-like and are set into a basal socket (
<figureCitation id="94E22A07EF41FFA8FB8B9D4E67C4FBEE" box="[1132,1213,1122,1146]" captionStart="FIGURES 32 – 37" captionStartId="12.[151,264,1847,1870]" captionTargetBox="[154,1436,187,1826]" captionTargetId="figure@12.[154,1439,187,1826]" captionTargetPageId="12" captionText="FIGURES 32 – 37. Ischyropsalis gigantea, male from Recuistro Cave (MNCN 20.02 / 9957, Upper Miera Mountains). Male cheliceral apophysis on distal basal segment, medial view (32 – 33). Detail of bristle with basal socket structure revealing the whorled striae on the external surface and the lack of a discernable pore at the tip (34). Distal part of penis with glans and stylus: dorsal view (35), ventral view (36) and lateral view (37)." httpUri="https://zenodo.org/record/211043/files/figure.png" pageId="6" pageNumber="32">Fig 34</figureCitation>
). The chelicerae of females are larger, with basal segment more robust and armed with long spines of different size over its whole length (six shorter spines on dorsal surface, ventro-medially with six, and ventro-laterally with five spines; 
<figureCitation id="94E22A07EF41FFA8FAA39D866607FB56" box="[1348,1406,1194,1218]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Fig 8</figureCitation>
b). The distal segment is covered with small rounded piliferous tubercles at the dorsal knee, and also with some short hairs towards the base of the pincer.
</paragraph>
<paragraph id="0C663682EF41FFA8FF209C3B6798FAE2" blockId="6.[151,1437,151,1963]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209C3B624FFABB" bold="true" box="[199,310,1303,1327]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209C3B6257FABA" bold="true" box="[199,302,1303,1326]" italics="true" pageId="6" pageNumber="32">Prosoma</emphasis>
:
</emphasis>
Cephalothorax flat (
<figureCitation id="94E22A07EF41FFA8FDCD9C3A6103FABA" box="[554,634,1302,1326]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1921,1944]" captionTargetBox="[152,1404,552,1902]" captionTargetId="figure@4.[151,1436,535,1903]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Ischyropsalis cantabrica sp. n., male holotype from Rogeria Cave (MNCN 20.02 / 12828). Cephalothorax, lateral view (10). Male cheliceral apophysis on distal basal segment, medial view (8 a, 9), female chelicerae (8 b). Total view of ventral penis (6); distal part of penis with glans and stylus, ventral view (7)." httpUri="https://zenodo.org/record/211035/files/figure.png" pageId="6" pageNumber="32">Fig 10</figureCitation>
), black or dark brown, moderately ascending from border of second thoracic tergite, and ornamented with a transversal row of bristles with six denticles of different sizes; eye mound weakly developed, eyes widely separated, with small lenses in comparison to 
<taxonomicName id="CBD94D01EF41FFA8FC199C4C6701FAE2" box="[1022,1144,1375,1399]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="3EADEA90EF41FFA8FC199C4C6701FAE2" box="[1022,1144,1375,1399]" italics="true" pageId="6" pageNumber="32">I. gigantea</emphasis>
</taxonomicName>
(
<figureCitation id="94E22A07EF41FFA8FB609C7267ACFAE2" box="[1159,1237,1374,1398]" captionStart="FIGURES 27 – 31" captionStartId="11.[151,264,984,1007]" captionTargetBox="[195,1384,163,962]" captionTargetId="figure@11.[190,1395,143,970]" captionTargetPageId="11" captionText="FIGURES 27 – 31. Ischyropsalis gigantea, male from Los Moros del Río Munio Cave (Upper Asón Mountain). Cephalothorax, lateral view (27). Male cheliceral apophysis on distal basal segment, medial view (Figs 28 a – 29, female 28 b). Total penis, ventral view (30); distal part of penis with glans and stylus, ventral view (31)." httpUri="https://zenodo.org/record/211042/files/figure.png" pageId="6" pageNumber="32">Fig 27</figureCitation>
).
</paragraph>
<paragraph id="0C663682EF41FFA8FF209CAE67EEFA2A" blockId="6.[151,1437,151,1963]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209CAE621EFA0F" bold="true" box="[199,359,1410,1435]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209CAE6227FA0F" bold="true" box="[199,350,1410,1435]" italics="true" pageId="6" pageNumber="32">Opisthosoma</emphasis>
:
</emphasis>
Males and females with scutum parvum, the last three tergites always free. Sclerotised area finely grained, surrounded by slight grooves, each with a transversal line of bristles (
<figureCitation id="94E22A07EF41FFA8FBAC9C8A67F1FA2A" box="[1099,1160,1446,1470]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1921,1944]" captionTargetBox="[151,1435,721,1900]" captionTargetId="figure@2.[151,1436,721,1900]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Type locality and habitus of Ischyropsalis cantabrica sp. n. 1: Entrance to ‘ Rogeria Cave’ in Spain, Prov. Cantabria. 2: ‘ Rogeria Cave’ Digital Orthophoto with UTM grid, Zone 30, ED 50; 3: Life female specimen from ‘ Rogeria Cave’ (CGL 369)." httpUri="https://zenodo.org/record/211032/files/figure.png" pageId="6" pageNumber="32">Fig 3</figureCitation>
).
</paragraph>
<paragraph id="0C663682EF41FFA8FF209CE766E5F9DA" blockId="6.[151,1437,151,1963]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209CE7627FFA77" bold="true" box="[199,262,1483,1507]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209CE76384FA76" bold="true" box="[199,253,1483,1506]" italics="true" pageId="6" pageNumber="32">Legs</emphasis>
:
</emphasis>
Slender, of medium length, all segments rounded, light brown, metatarsus and tarsus of lighter colour, base of femora whitish to yellowish; femora without sculpture elements (spines or tubercles), only with microtrichia cover and setae; surface of basal segments (trochanter and coxa) with several 
<typeStatus id="D3628820EF41FFA8FB2C9F3F667FF9BF" box="[1227,1286,1555,1579]" pageId="6" pageNumber="32">types</typeStatus>
of sculpture elements (e.g. granules, tubercles and cones), with long macrotrichia, which arise from a basal socket (
<figureCitation id="94E22A07EF41FFA8FAEA9F1A66E9F9DA" box="[1293,1424,1590,1614]" captionStart-0="FIGURES 18 – 21" captionStart-1="FIGURES 22 – 24" captionStartId-0="8.[151,264,1248,1271]" captionStartId-1="9.[151,264,1335,1358]" captionTargetBox-0="[151,1435,181,1227]" captionTargetBox-1="[151,1435,157,1319]" captionTargetId-0="figure@8.[151,1436,181,1227]" captionTargetId-1="figure@9.[151,1436,157,1319]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 18 – 21. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Detail of the femoral plate with cuticular structures on legs II, showing the typical microtrichia and isolated slit sense organ (18), a group of deep-circular pits on the cuticle (19), the ‘ holes’ of unknown function and identity (20) and an unidentified deep funnel-shaped depression surrounded by four spines (21)." captionText-1="FIGURES 22 – 24. Ischyropsalis cantabrica sp. n., male paratype from Cerviz Cave (MNCN 20.02 / 12857). Detail of cuticular structures on the coxal plate of the legs II (22 a), showing the slit sense organ (black arrows), an unidentified deep funnelshaped depression surrounded by four spines (white arrow), several long hairs (macrotrichia), which arise from a basal socket (22), and other types of sculptural elements (Figs 23 – 24): e. g. granules, tubercles and cones with long macrotrichia on the apex, which arise from a basal socket." httpUri-0="https://zenodo.org/record/211038/files/figure.png" httpUri-1="https://zenodo.org/record/211039/files/figure.png" pageId="6" pageNumber="32">Figs 18–24</figureCitation>
).
</paragraph>
<paragraph id="0C663682EF41FFA8FF209F7662B4F969" blockId="6.[151,1437,151,1963]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209F766229F9E7" bold="true" box="[199,336,1626,1651]" pageId="6" pageNumber="32">Variability:</emphasis>
There seems to be no extraordinary variation despite the general differences in 
<taxonomicName id="CBD94D01EF41FFA8FAEF9F7766E5F9E7" box="[1288,1436,1627,1651]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="3EADEA90EF41FFA8FAEF9F7766E5F9E7" box="[1288,1436,1627,1651]" italics="true" pageId="6" pageNumber="32">Ischyropsalis</emphasis>
</taxonomicName>
species, e.g. in case of females spination is slightly more pronounced in larger specimens (see 
<bibRefCitation id="68484B73EF41FFA8FB2A9F526615F901" author="Martens" box="[1229,1388,1661,1685]" pageId="6" pageNumber="32" refString="Martens, J. (1969) Die Abgrenzung von Biospezies auf biologisch-etologischer und morphologischer Grundlage am Beispiel der Gattung Ischyropsalis C. L. Koch 1839 (Opiliones, Ischyropsalididae). Zoologische Jahrbuecher Abteilung fur Systematik, 96, 133 - 264." type="journal article" year="1969">Martens 1969</bibRefCitation>
, pp. 160–165). The structure of the glans penis is relatively constant. This is also the case for the other species of the Iberian 
<taxonomicName id="CBD94D01EF41FFA8FF099FEF6241F94F" box="[238,312,1731,1755]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="3EADEA90EF41FFA8FF099FEF6241F94F" box="[238,312,1731,1755]" italics="true" pageId="6" pageNumber="32">kollari</emphasis>
</taxonomicName>
group (
<emphasis id="3EADEA90EF41FFA8FE739FE862ABF94F" box="[404,466,1732,1755]" italics="true" pageId="6" pageNumber="32">sensu</emphasis>
<bibRefCitation id="68484B73EF41FFA8FE3D9FEF6103F94E" author="Martens" box="[474,634,1730,1754]" pageId="6" pageNumber="32" refString="Martens, J. (1969) Die Abgrenzung von Biospezies auf biologisch-etologischer und morphologischer Grundlage am Beispiel der Gattung Ischyropsalis C. L. Koch 1839 (Opiliones, Ischyropsalididae). Zoologische Jahrbuecher Abteilung fur Systematik, 96, 133 - 264." type="journal article" year="1969">Martens 1969</bibRefCitation>
), except for the heterogeneous 
<taxonomicName id="CBD94D01EF41FFA8FC3A9FE86744F94F" box="[989,1085,1731,1755]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="3EADEA90EF41FFA8FC3A9FE86744F94F" box="[989,1085,1731,1755]" italics="true" pageId="6" pageNumber="32">I. dispar</emphasis>
</taxonomicName>
group as shown in 
<figureCitation id="94E22A07EF41FFA8FAF99FEE66E5F94E" box="[1310,1436,1730,1754]" captionStart="FIGURES 38 – 41" captionStartId="13.[151,264,731,754]" captionTargetBox="[151,1435,193,699]" captionTargetId="figure@13.[151,1436,193,710]" captionTargetPageId="13" captionText="FIGURES 38 – 41. Ischyropsalis sp. aff. dispar from La Baja Cave (MNCN 20.02 / 12858, Laredo, Hacha Mountain). Male cheliceral apophysis on distal basal segment, medial view (38). Distal part of penis with glans and stylus: dorsal view (39), ventral view (40) and lateral view (41)." httpUri="https://zenodo.org/record/211044/files/figure.png" pageId="6" pageNumber="32">Figs 38–41</figureCitation>
and 
<figureCitation id="94E22A07EF41FFA8FF219FC96275F969" box="[198,268,1765,1789]" captionStart="FIGURES 50 – 57" captionStartId="14.[151,264,1221,1244]" captionTargetBox="[153,1435,195,1200]" captionTargetId="figure@14.[151,1436,195,1200]" captionTargetPageId="14" captionText="FIGURES 50 – 57. Ischyropsalis cf. noltei (or Ischyropsalis sp. aff. dispar) from Abaro Pit (MNCN 20.02 / 12860, Indusi Mountain; 50 – 53); Ischyropsalis sp. aff. dispar from La Pasada Cave (MNCN 20.02 / 12859, Guriezo; 54 – 57). Male cheliceral apophysis on distal basal segment, medial view (50, 54). Distal part of penis with glans and stylus: dorsal view (51, 55), ventral view (52, 56) and lateral view (53, 57)." httpUri="https://zenodo.org/record/211046/files/figure.png" pageId="6" pageNumber="32">50–57</figureCitation>
(see discussion).
</paragraph>
</subSubSection>
<subSubSection id="44C36509EF41FFA8FF209E246702F88B" box="[199,1147,1799,1824]" pageId="6" pageNumber="32" type="etymology">
<paragraph id="0C663682EF41FFA8FF209E246702F88B" blockId="6.[151,1437,151,1963]" box="[199,1147,1799,1824]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209E246228F8B4" bold="true" box="[199,337,1800,1824]" pageId="6" pageNumber="32">Etymology:</emphasis>
The species is named for the Cantabria Region, for which it is endemic.
</paragraph>
</subSubSection>
<subSubSection id="44C36509EF41FFA6FF209E066748F8FF" lastPageId="8" lastPageNumber="34" pageId="6" pageNumber="32" type="distribution">
<paragraph id="0C663682EF41FFA8FF209E0666E5F83F" blockId="6.[151,1437,151,1963]" pageId="6" pageNumber="32">
<emphasis id="3EADEA90EF41FFA8FF209E06622DF8D7" bold="true" box="[199,340,1834,1859]" pageId="6" pageNumber="32">Distribution</emphasis>
(
<figureCitation id="94E22A07EF41FFA8FE809E0662B7F8D6" box="[359,462,1834,1858]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="3.[151,250,541,564]" captionStartId-1="3.[151,250,1446,1469]" captionTargetBox-0="[151,1436,155,520]" captionTargetBox-1="[151,1436,886,1422]" captionTargetId-0="figure@3.[151,1436,155,520]" captionTargetId-1="figure@3.[151,1436,886,1422]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionText-0="FIGURE 4. Geographical map of central to northern Spain and surroundings of the sampling area (left). Ischyropsalis collection sites, with new sampling in original localities (right). Numbers refer to the following regions and species (Provinces are indicated in parentheses): 1, Western Coastal Cantabria inhabited by I. cantabrica sp. n.; 2, Upper Asón and Miera Mountains (Cantabria) inhabited by I. gigantea Dresco, 1968; 3, Triano-Galdames Mountains (Vizcaya) inhabited by I. magdalenae Simon, 1881; 4, Sierra de Aralar Massif (Guipúzcoa-Navarra) inhabited by I. navarrensis Roewer, 1950; 5, Sierra de Arno – Izarraitz Massif (Guipúzcoa) inhabited by I. galani nomen nudum; 6, Eastern part of the Basque-Cantabrian Mountains (inhabited by the complex of I. dispar Simon, 1872), including the localities of 6 a, Laredo – Limpias – Rasines – Guriezo Mountains (Cantabria), 6 b, Castro Urdiales (Cantabria), 6 c, Soncillo (Burgos), 6 d, Sierra Salvada Massif (Burgos – Álava), 6 e, Sierra de Arkamo Massif (Álava), 6 f – g, Gorbea Massif (Vizcaya), and 6 h, Sierra de Aizkorri Massif (Guipúzcoa). For cave names, coordinates and altitudes see Appendix 1." captionText-1="FIGURE 5. Map of the central region of the Cantabrian Mountain (northern Spain) to show the outcrops of the rock systems (in gray) which contain limestone and geographical situation of the study area with the distribution of the troglobitic species according to Luque (1991) and sampling of new localities (all species are allopatric): Ischyropsalis cantabrica sp. n.: Locus typicus ‘ Rogeria Cave’ (1), Portillo Cave (2), Cerviz Cave (3), Cáscaras Cave (4); Ischyropsalis gigantea (see Appendix 2 for explanation of numbers 5 – 12): Cayuela or Cañuela Cave (5), Currutuerta or Posadía Cave (6), Becerral Cave (7), Mortero Pit (8), Calleja Lavalle Pit (9), Los Moros del Río Munio Cave (10), Recuistro Cave (11), and Canal de la Torca nº 3 Pit, The Peña del Trillo Cave karst system (12); and Ischyropsalis sp. aff. dispar: La Baja Cave (13), Hoyo Molino Cave (14), Perelada Cave (15), Rejuyo or ‘ S. E. I. I. - 8 ’ Cave (16), Tornillos or RN- 30 Cave (17), La Lastrilla Cave (18), La Pasada Caves (19 – 20). Source: Geologic maps of Spain 1: 50,000 - scale corresponding to the Province of Cantabria (IGME)." httpUri-0="https://zenodo.org/record/211033/files/figure.png" httpUri-1="https://zenodo.org/record/211034/files/figure.png" pageId="6" pageNumber="32">Figs 4–5</figureCitation>
): Endemic to the northwestern Cantabrian coastal region in 
<collectingCountry id="74CE7612EF41FFA8FB989E0667C6F8D6" box="[1151,1215,1834,1858]" name="Spain" pageId="6" pageNumber="32">Spain</collectingCountry>
, and restricted to a narrow band of karst systems along the coast within the municipal districts of Alfoz de Lloredo, Ruiloba, Udías and Cabezón de la Sal. Epigean species, like 
<taxonomicName id="CBD94D01EF41FFA8FD869E5D61ACF81C" box="[609,725,1904,1928]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="nodifera">
<emphasis id="3EADEA90EF41FFA8FD869E5D61ACF81C" box="[609,725,1904,1928]" italics="true" pageId="6" pageNumber="32">I. nodifera</emphasis>
</taxonomicName>
and 
<taxonomicName id="CBD94D01EF41FFA8FCE99E5D60EEF81C" box="[782,919,1905,1928]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="petiginosa">
<emphasis id="3EADEA90EF41FFA8FCE99E5D60EEF81C" box="[782,919,1905,1928]" italics="true" pageId="6" pageNumber="32">I. petiginosa</emphasis>
</taxonomicName>
, occur in all caves visited along the Cantabria coast (see, e.g., 
<bibRefCitation id="68484B73EF41FFA8FED89EBF62BDF83E" author="Luque" box="[319,452,1938,1962]" pageId="6" pageNumber="32" refString="Luque, C. G. (1991) Los Ischyropsalidoidea de la Cornisa Cantabrica: Cantabria (Arachnida: Opiliones: Palpatores: Ischyropsalididae y Sabaconidae). Arquenas, 1, 1 - 96." type="journal article" year="1991">Luque 1991</bibRefCitation>
), while restricted to the shallower zone (&lt;
<quantity id="CB219B67EF41FFA8FC639EBE60C7F83E" box="[900,958,1938,1962]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="32" unit="m" value="30.0">30 m</quantity>
) and not interacting with 
<taxonomicName id="CBD94D01EF41FFA8FB379EB86622F83F" box="[1232,1371,1939,1963]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="32" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF41FFA8FB379EB86622F83F" box="[1232,1371,1939,1963]" italics="true" pageId="6" pageNumber="32">I. cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF41FFA8FA869EBF66E5F83F" box="[1377,1436,1939,1963]" pageId="6" pageNumber="32" rank="species">
<emphasis id="3EADEA90EF41FFA8FA869EBF66E5F83F" bold="true" box="[1377,1436,1939,1963]" pageId="6" pageNumber="32">sp. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
<caption id="58A6660AEF40FFA9FF709DC66104FA8B" httpUri="https://zenodo.org/record/211037/files/figure.png" pageId="7" pageNumber="33" targetBox="[151,1435,155,1247]" targetPageId="7">
<paragraph id="0C663682EF40FFA9FF709DC66104FA8B" blockId="7.[151,1436,1258,1311]" pageId="7" pageNumber="33">
<emphasis id="3EADEA90EF40FFA9FF709DC6622FFA94" bold="true" box="[151,342,1258,1281]" pageId="7" pageNumber="33">FIGURES 16–17.</emphasis>
<taxonomicName id="CBD94D01EF40FFA9FEB99DC76122FA95" box="[350,603,1259,1281]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF40FFA9FEB99DC76122FA95" box="[350,603,1259,1281]" italics="true" pageId="7" pageNumber="33">Ischyropsalis cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF40FFA9FD849DC761E7FA94" box="[611,670,1259,1280]" pageId="7" pageNumber="33" rank="species">
<emphasis id="3EADEA90EF40FFA9FD849DC761E7FA94" bold="true" box="[611,670,1259,1280]" pageId="7" pageNumber="33">sp. n.</emphasis>
</taxonomicNameLabel>
, male paratype from Cerviz Cave (MNCN 20.02/12857). Male cheliceral apophysis on distal basal segment, medial view.
</paragraph>
</caption>
<paragraph id="0C663682EF40FFA9FF209C4F60A3F817" blockId="7.[151,1436,1379,1923]" pageId="7" pageNumber="33">
<emphasis id="3EADEA90EF40FFA9FF209C4F6256FAEF" bold="true" box="[199,303,1379,1403]" pageId="7" pageNumber="33">Ecology:</emphasis>
The caves inhabited by 
<taxonomicName id="CBD94D01EF40FFA9FDBF9C486197FAEF" box="[600,750,1379,1403]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF40FFA9FDBF9C486197FAEF" box="[600,750,1379,1403]" italics="true" pageId="7" pageNumber="33">I. cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF40FFA9FD1D9C4F6047FAEF" box="[762,830,1379,1403]" pageId="7" pageNumber="33" rank="species">
<emphasis id="3EADEA90EF40FFA9FD1D9C4F6047FAEF" bold="true" box="[762,830,1379,1403]" pageId="7" pageNumber="33">sp. n.</emphasis>
</taxonomicNameLabel>
developed in the Lower Cretaceous (urgonian-type facies) limestones (Aptian age). All specimens were collected in the lightless zones of caves (especially in “deep cave” sites), about 
<quantity id="CB219B67EF40FFA9FE899C8462CCFA54" box="[366,437,1448,1472]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="33" unit="m" value="300.0">300 m</quantity>
from the entrance of Rogeria Cave (
<figureCitation id="94E22A07EF40FFA9FCB49C8460ECFA54" box="[851,917,1448,1472]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1921,1944]" captionTargetBox="[151,1435,721,1900]" captionTargetId="figure@2.[151,1436,721,1900]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Type locality and habitus of Ischyropsalis cantabrica sp. n. 1: Entrance to ‘ Rogeria Cave’ in Spain, Prov. Cantabria. 2: ‘ Rogeria Cave’ Digital Orthophoto with UTM grid, Zone 30, ED 50; 3: Life female specimen from ‘ Rogeria Cave’ (CGL 369)." httpUri="https://zenodo.org/record/211032/files/figure.png" pageId="7" pageNumber="33">Fig. 1</figureCitation>
), and about 
<quantity id="CB219B67EF40FFA9FBC49C846712FA54" box="[1059,1131,1448,1472]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="33" unit="m" value="150.0">150 m</quantity>
from the entrances of both Portillo and Cerviz Caves. The specimens were usually found close to water, on cave walls with abundant speleothems or mud-cover, and between 
<quantity id="CB219B67EF40FFA9FD839CC161AEF991" box="[612,727,1517,1541]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="7" pageNumber="33" unit="m" value="1.25" valueMax="2.0" valueMin="0.5">0.5 to 2 m</quantity>
above water level. In the Calero Cave 
<taxonomicName id="CBD94D01EF40FFA9FB779CC3665BF992" box="[1168,1314,1518,1542]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF40FFA9FB779CC3665BF992" box="[1168,1314,1518,1542]" italics="true" pageId="7" pageNumber="33">I. cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF40FFA9FACE9CC26610F992" box="[1321,1385,1518,1542]" pageId="7" pageNumber="33" rank="species">
<emphasis id="3EADEA90EF40FFA9FACE9CC26610F992" bold="true" box="[1321,1385,1518,1542]" pageId="7" pageNumber="33">sp. n.</emphasis>
</taxonomicNameLabel>
was observed to prey on the small aquatic gastropod 
<taxonomicName id="CBD94D01EF40FFA9FD029F3C6047F9BC" box="[741,830,1552,1576]" class="Gastropoda" family="Planorbidae" genus="Ancylus" kingdom="Animalia" order="Hygrophila" pageId="7" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis id="3EADEA90EF40FFA9FD029F3C6047F9BC" box="[741,830,1552,1576]" italics="true" pageId="7" pageNumber="33">Ancylus</emphasis>
</taxonomicName>
sp. aff. 
<emphasis id="3EADEA90EF40FFA9FC4D9F3C676AF9BC" box="[938,1043,1552,1576]" italics="true" pageId="7" pageNumber="33">fluviatilis</emphasis>
(in spring of late to mid-1970s; 
<bibRefCitation id="68484B73EF40FFA9FF709F1F6228F9DF" author="Menendez" box="[151,337,1587,1611]" pageId="7" pageNumber="33" refString="Menendez, R. (1973) Cinecologia de la cueva del Calero (Orena, Santander). Cuadernos de Espeleologia, 7, 61 - 82." type="journal article" year="1973">Menéndez 1973</bibRefCitation>
). Although they are mainly active predators, in some cases they do not disdain carrion. Sixteen other invertebrates were recorded from the deep cave zone of all the caves inhabited by 
<taxonomicName id="CBD94D01EF40FFA9FB509F7B6635F9FA" box="[1207,1356,1622,1646]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF40FFA9FB509F7B6635F9FA" box="[1207,1356,1622,1646]" italics="true" pageId="7" pageNumber="33">I. cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF40FFA9FABF9F7A66E5F9FA" box="[1368,1436,1622,1646]" pageId="7" pageNumber="33" rank="species">
<emphasis id="3EADEA90EF40FFA9FABF9F7A66E5F9FA" bold="true" box="[1368,1436,1622,1646]" pageId="7" pageNumber="33">sp. n.</emphasis>
</taxonomicNameLabel>
(
<bibRefCitation id="68484B73EF40FFA9FF799F54622FF904" author="Menendez" box="[158,342,1656,1680]" pageId="7" pageNumber="33" refString="Menendez, R. (1973) Cinecologia de la cueva del Calero (Orena, Santander). Cuadernos de Espeleologia, 7, 61 - 82." type="journal article" year="1973">Menéndez 1973</bibRefCitation>
; 
<bibRefCitation id="68484B73EF40FFA9FE869F546112F904" author="Labrada" box="[353,619,1656,1680]" pageId="7" pageNumber="33" refString="Labrada, L. &amp; Luque, C. G. (2004) Inventory of Bat Community Composition and of the Invertebrate Cave Fauna at Sites of Community Importance of the Cueva del Rejo and Cueva de la Rogeria (&quot; Natura 2000 sites &quot;), Cantabria, Spain. Unpublished research and technical report on behalf of IMPRESS Group Consulting (Pty) Ltd to the General Direction for Biodiversity of the Regional Government of Cantabria (Spain)." type="journal volume" year="2004">Labrada &amp; Luque 2004</bibRefCitation>
): 
<taxonomicName id="CBD94D01EF40FFA9FD9B9F5460C0F904" box="[636,953,1656,1680]" class="Malacostraca" family="Trichoniscidae" genus="Trichoniscoides" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="3EADEA90EF40FFA9FD9B9F5460C0F904" box="[636,953,1656,1680]" italics="true" pageId="7" pageNumber="33">Trichoniscoides cavernicola</emphasis>
</taxonomicName>
(class Crustacea, order 
<taxonomicName id="CBD94D01EF40FFA9FB259F546659F904" box="[1218,1312,1656,1680]" class="Malacostraca" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Isopoda</taxonomicName>
); 
<taxonomicName id="CBD94D01EF40FFA9FAD59F546391F927" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="3EADEA90EF40FFA9FAD59F546391F927" italics="true" pageId="7" pageNumber="33">Lithobius validus</emphasis>
</taxonomicName>
and 
<taxonomicName id="CBD94D01EF40FFA9FECF9FB062D2F927" box="[296,427,1692,1715]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="tricuspis">
<emphasis id="3EADEA90EF40FFA9FECF9FB062D2F927" box="[296,427,1692,1715]" italics="true" pageId="7" pageNumber="33">L. tricuspis</emphasis>
</taxonomicName>
(both class Chilopoda, order 
<taxonomicName id="CBD94D01EF40FFA9FCE89FB760B3F927" box="[783,970,1691,1715]" class="Chilopoda" kingdom="Animalia" order="Lithobiomorpha" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Lithobiomorpha</taxonomicName>
); 
<taxonomicName id="CBD94D01EF40FFA9FC059FB76798F927" box="[994,1249,1691,1715]" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="3EADEA90EF40FFA9FC059FB76798F927" box="[994,1249,1691,1715]" italics="true" pageId="7" pageNumber="33">Polydesmus coriaceus</emphasis>
</taxonomicName>
, 
<taxonomicName id="CBD94D01EF40FFA9FB139FB76397F942" class="Diplopoda" family="Glomeridae" genus="Trachysphaera" kingdom="Animalia" order="Glomerida" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="rousseti">
<emphasis id="3EADEA90EF40FFA9FB139FB76397F942" italics="true" pageId="7" pageNumber="33">Trachysphaera rousseti</emphasis>
</taxonomicName>
and 
<taxonomicName id="CBD94D01EF40FFA9FECF9F926296F942" box="[296,495,1726,1750]" class="Diplopoda" family="Blaniulidae" genus="Blaniulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="dollfusi">
<emphasis id="3EADEA90EF40FFA9FECF9F926296F942" box="[296,495,1726,1750]" italics="true" pageId="7" pageNumber="33">Blaniulus dollfusi</emphasis>
</taxonomicName>
(all three class Diplopoda, orders 
<taxonomicName id="CBD94D01EF40FFA9FC9F9F916775F941" box="[888,1036,1725,1749]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
, 
<taxonomicName id="CBD94D01EF40FFA9FBFF9F9167E9F941" box="[1048,1168,1725,1749]" class="Diplopoda" kingdom="Animalia" order="Glomerida" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Glomerida</taxonomicName>
and Iulida); 
<taxonomicName id="CBD94D01EF40FFA9FAC39F93638FF96C" class="Entognatha" family="Campodeidae" genus="Litocampa" kingdom="Animalia" order="Diplura" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="espanoli">
<emphasis id="3EADEA90EF40FFA9FAC39F93638FF96C" italics="true" pageId="7" pageNumber="33">Litocampa espanoli</emphasis>
</taxonomicName>
(class Insecta, order 
<taxonomicName id="CBD94D01EF40FFA9FE099FCC6130F96C" box="[494,585,1760,1784]" class="Entognatha" kingdom="Animalia" order="Diplura" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Diplura</taxonomicName>
); 
<taxonomicName id="CBD94D01EF40FFA9FDB99FCC60ECF96C" box="[606,917,1760,1784]" class="Entognatha" family="Entomobryidae" genus="Pseudosinella" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="superoculata">
<emphasis id="3EADEA90EF40FFA9FDB99FCC60ECF96C" box="[606,917,1760,1784]" italics="true" pageId="7" pageNumber="33">Pseudosinella superoculata</emphasis>
</taxonomicName>
and 
<taxonomicName id="CBD94D01EF40FFA9FC359FCC6726F96C" box="[978,1119,1760,1784]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="3EADEA90EF40FFA9FC359FCC6726F96C" box="[978,1119,1760,1784]" italics="true" pageId="7" pageNumber="33">Arrhopalites</emphasis>
</taxonomicName>
sp. aff. 
<emphasis id="3EADEA90EF40FFA9FB269FCC666EF96C" box="[1217,1303,1760,1784]" italics="true" pageId="7" pageNumber="33">elegans</emphasis>
(both class Insecta, order 
<taxonomicName id="CBD94D01EF40FFA9FE869E2F6290F88F" box="[353,489,1795,1819]" class="Entognatha" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Collembola</taxonomicName>
); 
<taxonomicName id="CBD94D01EF40FFA9FDF69E2F6056F88F" box="[529,815,1795,1819]" class="Arachnida" family="Neobisiidae" genus="Neobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="3EADEA90EF40FFA9FDF69E2F6056F88F" box="[529,815,1795,1819]" italics="true" pageId="7" pageNumber="33">Neobisium (Neobisium)</emphasis>
</taxonomicName>
sp. aff. 
<emphasis id="3EADEA90EF40FFA9FC2E9E286729F88F" box="[969,1104,1796,1819]" italics="true" pageId="7" pageNumber="33">cavernarum</emphasis>
(class Arachnida, order Pseudoscorpionida); 
<emphasis id="3EADEA90EF40FFA9FE6F9E0B6055F8AA" box="[392,812,1830,1854]" italics="true" pageId="7" pageNumber="33">
<taxonomicName id="CBD94D01EF40FFA9FE6F9E0B6175F8AA" box="[392,524,1831,1854]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="33" phylum="Arthropoda" rank="genus">Nemastoma</taxonomicName>
(s. lat.) sexmucronatum
</emphasis>
(class Arachnida, order 
<taxonomicName id="CBD94D01EF40FFA9FBB19E0967B1F8A9" box="[1110,1224,1829,1853]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
); 
<taxonomicName id="CBD94D01EF40FFA9FB069E0A6262F8F4" class="Arachnida" family="Linyphiidae" genus="Troglohyphantes" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="cantabricus">
<emphasis id="3EADEA90EF40FFA9FB069E0A6262F8F4" italics="true" pageId="7" pageNumber="33">Troglohyphantes cantabricus</emphasis>
</taxonomicName>
and 
<taxonomicName id="CBD94D01EF40FFA9FE859E65616FF8F4" box="[354,534,1864,1888]" class="Arachnida" family="Nesticidae" genus="Nesticus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="luquei">
<emphasis id="3EADEA90EF40FFA9FE859E65616FF8F4" box="[354,534,1864,1888]" italics="true" pageId="7" pageNumber="33">Nesticus luquei</emphasis>
</taxonomicName>
(both class Arachnida, order 
<taxonomicName id="CBD94D01EF40FFA9FC6B9E646097F8F4" box="[908,1006,1864,1888]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
); 
<taxonomicName id="CBD94D01EF40FFA9FBEE9E65661DF8F4" box="[1033,1380,1864,1888]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="arcanus" subGenus="Quaestus">
<emphasis id="3EADEA90EF40FFA9FBEE9E65661DF8F4" box="[1033,1380,1864,1888]" italics="true" pageId="7" pageNumber="33">Quaestus (Quaestus) arcanus</emphasis>
</taxonomicName>
and 
<taxonomicName id="CBD94D01EF40FFA9FF709E40614DF817" box="[151,564,1899,1923]" class="Insecta" family="Carabidae" genus="Laemostenus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="33" phylum="Arthropoda" rank="species" species="pelaeus" subGenus="Antisphodrus">
<emphasis id="3EADEA90EF40FFA9FF709E40614DF817" box="[151,564,1899,1923]" italics="true" pageId="7" pageNumber="33">Laemostenus (Antisphodrus) pelaeus</emphasis>
</taxonomicName>
(both class Insect, order 
<taxonomicName id="CBD94D01EF40FFA9FCA99E4760B7F817" box="[846,974,1899,1923]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="33" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
).
</paragraph>
<caption id="58A6660AEF4FFFA6FF709DCC6131FAC7" httpUri="https://zenodo.org/record/211038/files/figure.png" pageId="8" pageNumber="34" targetBox="[151,1435,181,1227]" targetPageId="8">
<paragraph id="0C663682EF4FFFA6FF709DCC6131FAC7" blockId="8.[151,1436,1248,1363]" pageId="8" pageNumber="34">
<emphasis id="3EADEA90EF4FFFA6FF709DCC6221FB62" bold="true" box="[151,344,1248,1271]" pageId="8" pageNumber="34">FIGURES 18–21.</emphasis>
<taxonomicName id="CBD94D01EF4FFFA6FE859DCD6119FB63" box="[354,608,1249,1271]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF4FFFA6FE859DCD6119FB63" box="[354,608,1249,1271]" italics="true" pageId="8" pageNumber="34">Ischyropsalis cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF4FFFA6FD8C9DCD61D1FB62" box="[619,680,1249,1270]" pageId="8" pageNumber="34" rank="species">
<emphasis id="3EADEA90EF4FFFA6FD8C9DCD61D1FB62" bold="true" box="[619,680,1249,1270]" pageId="8" pageNumber="34">sp. n.</emphasis>
</taxonomicNameLabel>
, male paratype from Cerviz Cave (MNCN 20.02/12857). Detail of the femoral plate with cuticular structures on legs II, showing the typical microtrichia and isolated slit sense organ (18), a group of deep-circular pits on the cuticle (19), the ‘holes’ of unknown function and identity (20) and an unidentified deep funnel-shaped depression surrounded by four spines (21).
</paragraph>
</caption>
<paragraph id="0C663682EF4FFFA6FF209C8F6748F8FF" blockId="8.[151,1436,1443,1899]" pageId="8" pageNumber="34">
<emphasis id="3EADEA90EF4FFFA6FF209C8F6208FA28" bold="true" box="[199,369,1443,1468]" pageId="8" pageNumber="34">Conservation:</emphasis>
Although 
<taxonomicName id="CBD94D01EF4FFFA6FE109C8961F6FA28" box="[503,655,1444,1468]" class="Arachnida" family="Ischyropsalididae" genus="Ischyropsalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="cantabrica" status="sp. nov.">
<emphasis id="3EADEA90EF4FFFA6FE109C8961F6FA28" box="[503,655,1444,1468]" italics="true" pageId="8" pageNumber="34">I. cantabrica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="259E57EBEF4FFFA6FD799C88619FFA28" box="[670,742,1444,1468]" pageId="8" pageNumber="34" rank="species">
<emphasis id="3EADEA90EF4FFFA6FD799C88619FFA28" bold="true" box="[670,742,1444,1468]" pageId="8" pageNumber="34">sp. n.</emphasis>
</taxonomicNameLabel>
seems frequent in suitable habitats, which are numerous throughout its area, it is an extremely narrow endemic species and therefore requires particular conservational attention. So far, the species is not listed as ‘vulnerable’ under the Threatened Species Act of 2008 (enacted under Decree No. 120/2008, and regionally known as ‘Catalogue of Threatened Species from Cantabria’) or the Nature Conservation Act of 2006 (under the Cantabria Regional Government legislation; 
<bibRefCitation id="68484B73EF4FFFA6FBBE9F1F663AF9DF" author="Labrada" box="[1113,1347,1587,1611]" pageId="8" pageNumber="34" refString="Labrada, L., Salgado, J. M. &amp; Luque, C. G. (2010) Fauna invertebrada de las cavidades de Cantabria. Locustella, Anuario de la Naturaleza de Cantabria, 7, 29 - 43." type="journal article" year="2010">Labrada et al. 2010</bibRefCitation>
). These legislations cover the protection of a wide range of species (e.g. cave-dwelling bats and invertebrates such as cavedwelling beetles 
<taxonomicName id="CBD94D01EF4FFFA6FE839F50614DF900" box="[356,564,1660,1684]" class="Insecta" family="Leiodidae" genus="Quaestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="34" phylum="Arthropoda" rank="species" species="arcanus">
<emphasis id="3EADEA90EF4FFFA6FE839F50614DF900" box="[356,564,1660,1684]" italics="true" pageId="8" pageNumber="34">Quaestus arcanus</emphasis>
</taxonomicName>
) and habitats (e.g., in the case of Rogeria Cave). This cave is further acknowledged as ‘important bat underground habitat’ (
<bibRefCitation id="68484B73EF4FFFA6FCFB9FB360A3F923" author="Menendez" box="[796,986,1695,1719]" pageId="8" pageNumber="34" refString="Menendez, R. (1973) Cinecologia de la cueva del Calero (Orena, Santander). Cuadernos de Espeleologia, 7, 61 - 82." type="journal article" year="1973">Menéndez 1973</bibRefCitation>
; 
<bibRefCitation id="68484B73EF4FFFA6FC0D9FB3667AF923" author="Labrada" box="[1002,1283,1695,1720]" pageId="8" pageNumber="34" refString="Labrada, L. &amp; Luque, C. G. (2004) Inventory of Bat Community Composition and of the Invertebrate Cave Fauna at Sites of Community Importance of the Cueva del Rejo and Cueva de la Rogeria (&quot; Natura 2000 sites &quot;), Cantabria, Spain. Unpublished research and technical report on behalf of IMPRESS Group Consulting (Pty) Ltd to the General Direction for Biodiversity of the Regional Government of Cantabria (Spain)." type="journal volume" year="2004">Labrada &amp; Luque 2004</bibRefCitation>
, unpublished results). Access to the cave is currently restricted, and occasionally prohibited, to protect threatened cave-dwelling bats —e.g. 
<taxonomicName id="CBD94D01EF4FFFA6FEFE9FC4617AF894" box="[281,515,1768,1792]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="34" phylum="Chordata" rank="species" species="euryale">
<emphasis id="3EADEA90EF4FFFA6FEFE9FC4617AF894" box="[281,515,1768,1792]" italics="true" pageId="8" pageNumber="34">Rhinolophus euryale</emphasis>
</taxonomicName>
, 
<taxonomicName id="CBD94D01EF4FFFA6FDF49FC5605EF894" box="[531,807,1768,1792]" class="Mammalia" family="Miniopteridae" genus="Miniopterus" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="34" phylum="Chordata" rank="species" species="schreibersii">
<emphasis id="3EADEA90EF4FFFA6FDF49FC5605EF894" box="[531,807,1768,1792]" italics="true" pageId="8" pageNumber="34">Miniopterus schreibersii</emphasis>
</taxonomicName>
and 
<emphasis id="3EADEA90EF4FFFA6FC869FC56087F894" box="[865,1022,1769,1792]" italics="true" pageId="8" pageNumber="34">Myotis myotis</emphasis>
— particularly during their breeding season (from June to August-September). For this reason, Rogeria Cave is included in the list of Areas or Sites of Community Importance for the Atlantic biogeographical region as SCI ES-1300017 (Commission Decision 2010/ 43/EU of 
<date id="78671042EF4FFFA6FEE29E7F62A3F8FF" box="[261,474,1875,1899]" pageId="8" pageNumber="34" value="2009-12-22">22 December 2009</date>
adopting, pursuant to Council Directive 92/43/EEC).
</paragraph>
</subSubSection>
</treatment>
</document>