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<document id="A106252E9F281752550F2A8678F9973B" ID-CLB-Dataset="26748" ID-DOI="10.11646/zootaxa.4413.1.2" ID-GBIF-Dataset="e9339680-76a0-4c9d-90fd-ecab97b50d9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3066636" ID-ZooBank="D65DAEB6-0499-44F9-A70D-0720BA71D520" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558414641282" checkinUser="plazi" docAuthor="Sassi, Davide" docDate="2018" docId="03DB87963221983BFF481676FE2DF84C" docLanguage="en" docName="zootaxa.4413.1.2.pdf" docOrigin="Zootaxa 4413 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metallactus regalini Sassi 2018, sp. nov." docType="treatment" docUuid="BCD7D90F-F8FC-4714-B986-95D6285D9032" docUuidSource="ZooBank" docVersion="12" lastPageNumber="100" masterDocId="FFE2FFEE320B9810FFDF1107FFBCFFD9" masterDocTitle="Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae)" masterLastPageNumber="110" masterPageNumber="57" pageNumber="99" updateTime="1698731295107" updateUser="ExternalLinkService">
<mods:mods id="BB0455BB72241AE2089199320816ED23" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7161D9E68F02ECBE25B6E32C8CA8AD10">
<mods:title id="5939DC4300066EEFC6166CBBBEDEBFDF">Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae)</mods:title>
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<mods:namePart id="B8FD56FF69E929B94286A65AB33D146B">Sassi, Davide</mods:namePart>
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<mods:title id="020EA79C0F80D357013CFF8310A86779">Zootaxa</mods:title>
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<mods:date id="74C4FBB5D0DBF505CADFF3E25EA8E886">2018</mods:date>
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<mods:number id="E6CE4AD76A28244EB67A4D9DA8A7F9EF">2018-04-20</mods:number>
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<mods:number id="F52058DE37E1DF3DF0AEACCC8FA98EDF">4413</mods:number>
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<mods:classification id="835BA84972175906E14D7448143AA2F6">journal article</mods:classification>
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<mods:identifier id="EC7F175F52C2D65AE3FE0C57A7B6F108" type="DOI">10.11646/zootaxa.4413.1.2</mods:identifier>
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<treatment id="03DB87963221983BFF481676FE2DF84C" ID-DOI="http://doi.org/10.5281/zenodo.5970590" ID-GBIF-Taxon="157232649" ID-Zenodo-Dep="5970590" LSID="urn:lsid:zoobank.org:act:BCD7D90F-F8FC-4714-B986-95D6285D9032" httpUri="http://treatment.plazi.org/id/03DB87963221983BFF481676FE2DF84C" lastPageId="43" lastPageNumber="100" pageId="42" pageNumber="99">
<subSubSection id="C368650B3221983AFF481676FE53F852" box="[151,495,1905,1931]" pageId="42" pageNumber="99" type="nomenclature">
<paragraph id="8BCD36803221983AFF481676FE53F852" blockId="42.[151,495,1905,1965]" box="[151,495,1905,1931]" pageId="42" pageNumber="99">
<heading id="D08581EC3221983AFF481676FE53F852" bold="true" box="[151,495,1905,1931]" fontSize="11" level="1" pageId="42" pageNumber="99" reason="1">
<emphasis id="B906EA923221983AFF481676FE53F852" bold="true" box="[151,495,1905,1931]" pageId="42" pageNumber="99">
<taxonomicName id="4C724D033221983AFF481676FE35F852" authority="Sassi, 2018" authorityName="Sassi" authorityYear="2018" box="[151,393,1905,1931]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="99" phylum="Arthropoda" rank="species" species="regalini" status="sp. nov.">
<emphasis id="B906EA923221983AFF481676FE35F852" bold="true" box="[151,393,1905,1931]" italics="true" pageId="42" pageNumber="99">Metallactus regalini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557E93221983AFE4E1676FE53F852" box="[401,495,1905,1931]" pageId="42" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368650B3221983AFF481693FE9BF875" box="[151,295,1940,1965]" pageId="42" pageNumber="99" type="description">
<paragraph id="8BCD36803221983AFF481693FE9BF875" blockId="42.[151,495,1905,1965]" box="[151,295,1940,1965]" pageId="42" pageNumber="99">
(
<figureCitation id="13492A053221983AFF401693FF4AF875" box="[159,246,1940,1965]" captionStart="FIGURES 1718" captionStartId="45.[151,264,1884,1907]" captionTargetBox="[246,1340,220,1843]" captionTargetId="figure@45.[205,1385,193,1868]" captionTargetPageId="45" captionText="FIGURES 1718. M. regalini (17), M. bezoar (18). 17ab: habitus; 17ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 17f: sclerites IIIIIIV of endophallus [0.5 mm]; 17g: sclerite I and dorsal spicule of endophallus [0.4 mm]. 18ab: habitus; 18ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 18f: sclerites IIIIIIV of endophallus [0.6 mm]; 18g: sclerite I and dorsal spicule of endophallus [0.5 mm]. 17ab: holotype; 18ab: holotype. hd: hairy dent: scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066654/files/figure.png" pageId="42" pageNumber="99">Figs 17</figureCitation>
;
<figureCitation id="13492A053221983AFEDD1693FEA3F875" box="[258,287,1940,1964]" captionStart="FIGURES 2936" captionStartId="51.[151,264,1934,1957]" captionTargetBox="[207,1393,222,1877]" captionTargetId="figure@51.[169,1418,193,1913]" captionTargetPageId="51" captionText="FIGURES 2936. Distribution maps for Metallactus kollari species-group. M. languidus (29); M. londonpridei (30); M. longicornis (31); M. patagonicus (32); M. pollens (33); M. regalini (34); M. rileyi (35); M. spilotus (36). Grey squares: localities reported in a generic way." httpUri="https://zenodo.org/record/3066660/files/figure.png" pageId="42" pageNumber="99">34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C368650B3220983BFF481190FAE9FF0C" pageId="43" pageNumber="100" type="etymology">
<paragraph id="8BCD36803220983BFF481190FAE9FF0C" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">
<emphasis id="B906EA923220983BFF481190FEA2FF69" bold="true" box="[151,286,151,176]" pageId="43" pageNumber="100">Etymology.</emphasis>
The species is dedicated to my friend and colleague Renato Regalin, valuable specialist in
<taxonomicName id="4C724D033220983BFF4811BBFEFDFF0C" authorityName="Latreille" authorityYear="1802" box="[151,321,188,213]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="family">Chrysomelidae</taxonomicName>
<taxonomicName id="4C724D033220983BFE9811BBFE1DFF0C" box="[327,417,188,213]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="tribe" tribe="Clytrini">Clytrini</taxonomicName>
, who passed away after a stouthearted struggle with cancer in the early part of 2016.
</paragraph>
</subSubSection>
<subSubSection id="C368650B3220983BFF1811D8FD59FEBC" pageId="43" pageNumber="100" type="materials_examined">
<paragraph id="8BCD36803220983BFF1811D8FBB0FE99" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">
<emphasis id="B906EA923220983BFF1811D8FECAFF21" bold="true" box="[199,374,223,248]" pageId="43" pageNumber="100">Type material.</emphasis>
<materialsCitation id="3B1A3CDD3220983BFEA211E7FB8AFEC5" ID-GBIF-Occurrence="2243028704" collectorName="Mallo do Hamer R. Longo &amp; F. Monros Collection" country="Brazil" location="Mallo do Hamer" pageId="43" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Parana" typeStatus="holotype">
<typeStatus id="54C988223220983BFEA211E7FE4BFF2E" box="[381,503,224,247]" pageId="43" pageNumber="100" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D74FD093220983BFDD811E7FDA2FF2E" box="[519,542,224,247]" pageId="43" pageNumber="100" type="male"></specimenCount>
, // “
<collectingCountry id="F36576103220983BFD9211D8FD2EFF21" box="[589,658,223,248]" name="Brazil" pageId="43" pageNumber="100">Brasil</collectingCountry>
<collectingRegion id="49B6F8623220983BFD4611E7FD59FF2E" box="[665,741,224,247]" country="Brazil" name="Parana" pageId="43" pageNumber="100">Parana</collectingRegion>
<collectorName id="268753563220983BFD3211D8FBA7FF21" box="[749,1051,223,248]" pageId="43" pageNumber="100">
<location id="8EAD605B3220983BFD3211D8FC14FF2E" LSID="urn:lsid:plazi:treatment:03DB87963221983BFF481676FE2DF84C:8EAD605B3220983BFD3211D8FC14FF2E" box="[749,936,223,248]" country="Brazil" name="Mallo do Hamer" pageId="43" pageNumber="100" stateProvince="Parana">Mallo do Hamer</location>
R. Longo
</collectorName>
leg.” [white label, handwritten] // “
<collectorName id="268753563220983BFF7D1003FE2EFEC4" box="[162,402,260,285]" pageId="43" pageNumber="100">F. Monrós Collection</collectorName>
1959” [white label, printed] // “
<taxonomicName id="4C724D033220983BFD261003FC61FEC5" authority="Sassi, 2018" authorityName="Sassi" authorityYear="2018" box="[761,989,260,284]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="species" species="regalini" status="sp. nov.">
<emphasis id="B906EA923220983BFD261003FC61FEC5" box="[761,989,260,284]" italics="true" pageId="43" pageNumber="100">Metallactus regalini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557E93220983BFC3C1002FB8AFEC5" box="[995,1078,261,284]" pageId="43" pageNumber="100" rank="species">sp. nov</taxonomicNameLabel>
</materialsCitation>
.
<materialsCitation id="3B1A3CDD3220983BFB9F100FFBB4FE99" ID-GBIF-Occurrence="2243028706" collectionCode="USNMNH" collectorName="D. Sassi" country="Brazil" pageId="43" pageNumber="100" specimenCount="2" stateProvince="Parana" typeStatus="holotype">
<typeStatus id="54C988223220983BFB9F100FFB79FEC2" box="[1088,1221,264,283]" pageId="43" pageNumber="100" type="holotype">HOLOTYPUS</typeStatus>
<collectorName id="268753563220983BFB131002FA94FEC4" box="[1228,1320,260,285]" pageId="43" pageNumber="100">D. Sassi</collectorName>
des.” [red label, printed] // (
<collectionCode id="ED63AE453220983BFE86102FFE65FE99" box="[345,473,296,320]" pageId="43" pageNumber="100">USNMNH</collectionCode>
). The
<typeStatus id="54C988223220983BFDC11020FD3CFE99" box="[542,640,295,320]" pageId="43" pageNumber="100" type="holotype">holotype</typeStatus>
is the only specimen known so far
</materialsCitation>
.
</paragraph>
<paragraph id="8BCD36803220983BFF18104BFD59FEBC" blockId="43.[151,1437,151,1941]" box="[199,741,332,357]" pageId="43" pageNumber="100">
<materialsCitation id="3B1A3CDD3220983BFF18104BFD5DFEBC" ID-GBIF-Occurrence="2243028689" box="[199,737,332,357]" country="Brazil" location="Mallo do Hamer" pageId="43" pageNumber="100" specimenCount="1" stateProvince="Parana" typeStatus="holotype">
<emphasis id="B906EA923220983BFF18104BFED9FEBC" bold="true" box="[199,357,332,357]" pageId="43" pageNumber="100">
<typeStatus id="54C988223220983BFF18104BFEBEFEBC" box="[199,258,332,357]" pageId="43" pageNumber="100">Type</typeStatus>
locality.
</emphasis>
<location id="8EAD605B3220983BFEB3104BFD99FEBD" LSID="urn:lsid:plazi:treatment:03DB87963221983BFF481676FE2DF84C:8EAD605B3220983BFEB3104BFD99FEBD" box="[364,549,332,357]" country="Brazil" name="Mallo do Hamer" pageId="43" pageNumber="100" stateProvince="Parana">Mallo do Hamer</location>
(
<collectingRegion id="49B6F8623220983BFDEB104BFD39FEBD" box="[564,645,332,356]" country="Brazil" name="Parana" pageId="43" pageNumber="100">Parana</collectingRegion>
,
<collectingCountry id="F36576103220983BFD50104BFD64FEBC" box="[655,728,332,357]" name="Brazil" pageId="43" pageNumber="100">Brazil</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C368650B3220983BFF181068FE0FFE51" box="[199,435,367,392]" pageId="43" pageNumber="100" type="distribution">
<paragraph id="8BCD36803220983BFF181068FE0FFE51" blockId="43.[151,1437,151,1941]" box="[199,435,367,392]" pageId="43" pageNumber="100">
<emphasis id="B906EA923220983BFF181068FEDCFE51" bold="true" box="[199,352,367,392]" pageId="43" pageNumber="100">Distribution.</emphasis>
<collectingCountry id="F36576103220983BFEB81068FE13FE51" box="[359,431,367,392]" name="Brazil" pageId="43" pageNumber="100">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C368650B3220983BFF181093FD27FD71" pageId="43" pageNumber="100" type="diagnosis">
<paragraph id="8BCD36803220983BFF181093FD27FD71" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">
<emphasis id="B906EA923220983BFF181093FEFDFE74" bold="true" box="[199,321,404,429]" pageId="43" pageNumber="100">Diagnosis.</emphasis>
A
<taxonomicName id="4C724D033220983BFEBB1093FE54FE75" authorityName="Suffrian" authorityYear="1866" box="[356,488,404,428]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA923220983BFEBB1093FE54FE75" box="[356,488,404,428]" italics="true" pageId="43" pageNumber="100">Metallactus</emphasis>
</taxonomicName>
of small size. It is quite similar to
<taxonomicName id="4C724D033220983BFCAE1092FC5FFE75" authorityName="Sassi" authorityYear="2018" box="[881,995,404,428]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="species" species="bezoar">
<emphasis id="B906EA923220983BFCAE1092FC5FFE75" box="[881,995,404,428]" italics="true" pageId="43" pageNumber="100">M. bezoar</emphasis>
</taxonomicName>
in size and dorsal colour pattern, but it differs in having legs completely black and head yellow colour pattern formed by a small triangular spot on lower part of vertex (in
<taxonomicName id="4C724D033220983BFE8010DAFE6DFE2D" authorityName="Sassi" authorityYear="2018" box="[351,465,476,500]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="species" species="bezoar">
<emphasis id="B906EA923220983BFE8010DAFE6DFE2D" box="[351,465,476,500]" italics="true" pageId="43" pageNumber="100">M. bezoar</emphasis>
</taxonomicName>
femora and basal half of tibiae are yellow and head yellow pattern is a x-shaped spot extended on lower part of frons and clypeus). Besides, pronotal outline in
<taxonomicName id="4C724D033220983BFBD71306FB35FDC1" authorityName="Sassi" authorityYear="2018" box="[1032,1161,512,536]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="species" species="regalini">
<emphasis id="B906EA923220983BFBD71306FB35FDC1" box="[1032,1161,512,536]" italics="true" pageId="43" pageNumber="100">M. regalini</emphasis>
</taxonomicName>
is more transverse and elliptical (slightly tronco-conical in
<taxonomicName id="4C724D033220983BFDEB1322FD14FDE5" authorityName="Sassi" authorityYear="2018" box="[564,680,548,572]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="species" species="bezoar">
<emphasis id="B906EA923220983BFDEB1322FD14FDE5" box="[564,680,548,572]" italics="true" pageId="43" pageNumber="100">M. bezoar</emphasis>
</taxonomicName>
). Aedeagal shape is also different comparing the two species.
<taxonomicName id="4C724D033220983BFAA21322FE94FDB9" authorityName="Sassi" authorityYear="2018" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="species" species="londonpridei">
<emphasis id="B906EA923220983BFAA21322FE94FDB9" italics="true" pageId="43" pageNumber="100">M. londonpridei</emphasis>
</taxonomicName>
is also similar in elytral and ventral patterns, but it differs in larger size, different pronotal coloration and different aedeagal shape. Pronotal outline is also different in
<taxonomicName id="4C724D033220983BFCB5136AFBA3FD5D" authorityName="Sassi" authorityYear="2018" box="[874,1055,620,644]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="100" phylum="Arthropoda" rank="species" species="londonpridei">
<emphasis id="B906EA923220983BFCB5136AFBA3FD5D" box="[874,1055,620,644]" italics="true" pageId="43" pageNumber="100">M. londonpridei</emphasis>
</taxonomicName>
, being slightly subconical and not elliptical, with almost straight lateral margins.
</paragraph>
</subSubSection>
<subSubSection id="C368650B3220983BFF1813B3FE2DF84C" pageId="43" pageNumber="100" type="description">
<paragraph id="8BCD36803220983BFF1813B3FDA8FD36" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">
<emphasis id="B906EA923220983BFF1813B3FE04FD14" bold="true" box="[199,440,692,717]" pageId="43" pageNumber="100">Description of male.</emphasis>
Habitus in
<figureCitation id="13492A053220983BFDE513B3FD3BFD14" box="[570,647,692,717]" captionStart="FIGURES 1718" captionStartId="45.[151,264,1884,1907]" captionTargetBox="[246,1340,220,1843]" captionTargetId="figure@45.[205,1385,193,1868]" captionTargetPageId="45" captionText="FIGURES 1718. M. regalini (17), M. bezoar (18). 17ab: habitus; 17ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 17f: sclerites IIIIIIV of endophallus [0.5 mm]; 17g: sclerite I and dorsal spicule of endophallus [0.4 mm]. 18ab: habitus; 18ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 18f: sclerites IIIIIIV of endophallus [0.6 mm]; 18g: sclerite I and dorsal spicule of endophallus [0.5 mm]. 17ab: holotype; 18ab: holotype. hd: hairy dent: scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066654/files/figure.png" pageId="43" pageNumber="100">Fig. 17</figureCitation>
ab (HT). BL =
<quantity id="4C8A9B653220983BFC9E13B3FC25FD15" box="[833,921,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="43" pageNumber="100" unit="mm" value="4.5">4.5 mm</quantity>
, BW =
<quantity id="4C8A9B653220983BFC2F13B3FBF4FD15" box="[1008,1096,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="43" pageNumber="100" unit="mm" value="2.7">2.7 mm</quantity>
, PL =
<quantity id="4C8A9B653220983BFB4C13B3FB57FD15" box="[1171,1259,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="43" pageNumber="100" unit="mm" value="1.6">1.6 mm</quantity>
, PW =
<quantity id="4C8A9B653220983BFAE013B3FA24FD15" box="[1343,1432,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="43" pageNumber="100" unit="mm" value="2.4">2.4 mm</quantity>
. Interocular distance 14.4% of BL.
</paragraph>
<paragraph id="8BCD36803220983BFF1813FBFE4EFC59" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">Head black, with a triangular yellow spot on lower part of vertex. Labrum blackish. Head surface with sparse pale setae. Mid-cranial suture short, blurred. Vertex almost impunctate. Frons and clypeus rather coarsely punctate in particular close to ocular rims and across midline. Antennae almost completely black, only first and second antennomeres partly brownish.</paragraph>
<paragraph id="8BCD36803220983BFF18128AFADCFC34" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">Pronotum yellow with a large, rectangular black plague not reaching margins. Pronotal shape sub-elliptical with lateral margins thin, regularly curved, barely visible from above. Posterolateral impressions faint, almost indistinct. Surface lustrous with scattered, fairly impressed punctation, slightly sparser and shallower on disc.</paragraph>
<paragraph id="8BCD36803220983BFF1812F0FDE9FBEC" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">Scutellum completely black, distinctly raised, sub-squared with truncated apex, almost bald and impunctate with tiny and sparse micropuncturation.</paragraph>
<paragraph id="8BCD36803220983BFF181538FCC0FB31" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">Elytra yellow with black pattern consisting of two large transverse bands and a pair of small rounded spots on apex. First transversal band including basal margin and not reaching lateral ones. Second transversal band, just behind half of elytra, extending from suture up to last interval but not reaching the lateral margins. Surface rather lustrous with punctation moderately impressed, denser on anterior half of disc, partly arranged in almost regular striae, fairly reduced and shallower on apical clivus. Intervals flat.</paragraph>
<paragraph id="8BCD36803220983BFF1815F3FD21FA8C" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">Pygidium yellow with a transverse black stripe on proximal margin and an elliptical black one on apex, with surface smooth, covered by sparse shallow punctures and whitish setae. Two little rounded pits close to each lateral margin of pygidium, roughly at half of length.</paragraph>
<paragraph id="8BCD36803220983BFF181458FCB5F9D1" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">Inferior parts of thorax completely black, only hypomera partly yellow on outer half. Abdominal ventrites black on midline, extensively yellow on sides, regularly and shallowly punctate and setose. Prothorax (prosternal process and hypomera) coarsely punctate; meso and metathorax coarsely punctate on sides, covered by whitish setae above all on midline. Prosternal process normally shaped, longitudinally grooved, apex blunt, surface covered with very long dense whitish setae. Legs black.</paragraph>
<paragraph id="8BCD36803220983BFF181713FAFFF941" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">
Fifth abdominal ventrite with a median, brownish, fairly impressed depression, smooth, glabrous, impunctate and posterior margin rather deeply notched. Median lobe of aedeagus (
<figureCitation id="13492A053220983BFC1F1730FBACF989" box="[960,1040,1591,1616]" captionStart="FIGURES 1718" captionStartId="45.[151,264,1884,1907]" captionTargetBox="[246,1340,220,1843]" captionTargetId="figure@45.[205,1385,193,1868]" captionTargetPageId="45" captionText="FIGURES 1718. M. regalini (17), M. bezoar (18). 17ab: habitus; 17ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 17f: sclerites IIIIIIV of endophallus [0.5 mm]; 17g: sclerite I and dorsal spicule of endophallus [0.4 mm]. 18ab: habitus; 18ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 18f: sclerites IIIIIIV of endophallus [0.6 mm]; 18g: sclerite I and dorsal spicule of endophallus [0.5 mm]. 17ab: holotype; 18ab: holotype. hd: hairy dent: scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066654/files/figure.png" pageId="43" pageNumber="100">Fig. 17</figureCitation>
ce) with sides slightly concave in ventral view, with apex wide, elongated, blunt, almost straight in lateral view. Hairy dents shallow, very poorly delimited, bearing quite long, straight and fairly close setae. Ventral profile weakly swollen in lateral view.
</paragraph>
<paragraph id="8BCD36803220983BFF1817A3FD6FF8A8" blockId="43.[151,1437,151,1941]" pageId="43" pageNumber="100">
Endophallus (
<figureCitation id="13492A053220983BFEBA17A3FE09F964" box="[357,437,1700,1725]" captionStart="FIGURES 1718" captionStartId="45.[151,264,1884,1907]" captionTargetBox="[246,1340,220,1843]" captionTargetId="figure@45.[205,1385,193,1868]" captionTargetPageId="45" captionText="FIGURES 1718. M. regalini (17), M. bezoar (18). 17ab: habitus; 17ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 17f: sclerites IIIIIIV of endophallus [0.5 mm]; 17g: sclerite I and dorsal spicule of endophallus [0.4 mm]. 18ab: habitus; 18ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [0.8 mm]; 18f: sclerites IIIIIIV of endophallus [0.6 mm]; 18g: sclerite I and dorsal spicule of endophallus [0.5 mm]. 17ab: holotype; 18ab: holotype. hd: hairy dent: scIIV: sclerites IIV of endophallus; ds: dorsal spicule. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066654/files/figure.png" pageId="43" pageNumber="100">Fig. 17</figureCitation>
fg) with sclerite I strongly developed and pigmented, denticle apparent, bent upwards and perceptibly sticking out laterally. Dorsal spicule strongly reduced, thorn-like. Sclerite II well developed, distinctly bent at base and gradually tapered towards apex. Arch of sclerite III slender, raised. Apex of sclerite III straight and pointed, fairly expanded on its proximal half, so that the sclerite looks like the head of a grebe. Branches of sclerite IV definitely shorter than sclerite III in the folded up structure, distinctly arched, with blunt, almost symmetrical apex and surface fairly rugose.
</paragraph>
<paragraph id="8BCD36803220983BFF18167BFE2DF84C" blockId="43.[151,1437,151,1941]" box="[199,401,1916,1941]" pageId="43" pageNumber="100">
<emphasis id="B906EA923220983BFF18167BFEA0F84D" box="[199,284,1916,1940]" italics="true" pageId="43" pageNumber="100">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
</treatment>
</document>