treatments-xml/data/03/DB/87/03DB878DFF9DD537CEB5FADDD773F9F5.xml

<document id="389CC7564A2C491486887AA60B5D5B84" ID="10.11646/zootaxa.4273.4.1SLASH11260" ID-CLB-Dataset="32893" ID-DOI="10.11646/zootaxa.4273.4.1" ID-GBIF-Dataset="3f4e48f2-f9a0-434e-8ec3-be455a1ec995" ID-ISSN="1175-5326" ID-Zenodo-Dep="803644" ID-ZooBank="310C1107-096C-4C9B-A131-B0F69DB938E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496819464485" checkinUser="plazi" docAuthor="Bortoluzzi, Sidnei, Caron, Edilson &amp; Ribeiro-Costa, Cibele S." docDate="2017" docId="03DB878DFF9DD537CEB5FADDD773F9F5" docLanguage="en" docName="zootaxa.4273.4.1SLASH11260.pdf" docOrigin="Zootaxa 4273 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypotelus brevitarsus Bortoluzzi &amp; Caron, sp. nov." docType="treatment" docVersion="7" lastPageNumber="463" masterDocId="FFE2FFF5FF96D53BCE22FF82D22AFFA5" masterDocTitle="Revision and phylogeny of Hypotelus Erichson: a Neotropical genus of minute rove beetles (Coleoptera, Staphylinidae, Piestinae)" masterLastPageNumber="487" masterPageNumber="451" pageNumber="462" updateTime="1698440233452" updateUser="ExternalLinkService">
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<mods:title id="6BCCEE065A95AE09425E691AD10E929C">Revision and phylogeny of Hypotelus Erichson: a Neotropical genus of minute rove beetles (Coleoptera, Staphylinidae, Piestinae)</mods:title>
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<mods:namePart id="1076E13860F27AADE40657C577C29A96">Bortoluzzi, Sidnei</mods:namePart>
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<mods:namePart id="D6BB25C58019EC0C3E53D1ADF371C882">Caron, Edilson</mods:namePart>
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<mods:namePart id="DCD797D1A577DF151C5384140C57D583">Ribeiro-Costa, Cibele S.</mods:namePart>
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<treatment id="03DB878DFF9DD537CEB5FADDD773F9F5" ID-DOI="http://doi.org/10.5281/zenodo.6017728" ID-GBIF-Taxon="131812440" ID-Zenodo-Dep="6017728" LSID="urn:lsid:plazi:treatment:03DB878DFF9DD537CEB5FADDD773F9F5" httpUri="http://treatment.plazi.org/id/03DB878DFF9DD537CEB5FADDD773F9F5" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
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<paragraph id="8BCD369BFF9DD530CEB5FADDD128FADF" blockId="11.[151,770,1375,1435]" box="[151,770,1375,1402]" pageId="11" pageNumber="462">
<heading id="D08581F7FF9DD530CEB5FADDD128FADF" bold="true" box="[151,770,1375,1402]" fontSize="11" level="1" pageId="11" pageNumber="462" reason="1">
<emphasis id="B906EA89FF9DD530CEB5FADDD128FADF" bold="true" box="[151,770,1375,1402]" pageId="11" pageNumber="462">
<taxonomicName id="4C724D18FF9DD530CEB5FADDD0B7FADF" ID-CoL="3NZD4" authority="Bortoluzzi &amp; Caron" authorityName="Bortoluzzi &amp; Caron" box="[151,669,1375,1402]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="brevitarsus" status="sp. nov.">
<emphasis id="B906EA89FF9DD530CEB5FADDD3B7FADC" bold="true" box="[151,413,1375,1401]" italics="true" pageId="11" pageNumber="462">Hypotelus brevitarsus</emphasis>
Bortoluzzi &amp; Caron
</taxonomicName>
<taxonomicNameLabel id="A23557F2FF9DD530CC87FAE2D128FADF" box="[677,770,1376,1402]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BCD369BFF9DD530CEB5FA00D04BFA3E" blockId="11.[151,770,1375,1435]" box="[151,609,1410,1435]" pageId="11" pageNumber="462">
(
<figureCitation id="13492A1EFF9DD530CEBDFA00D2D6FA3E" box="[159,252,1410,1435]" captionStart="FIGURES 9 – 19" captionStartId="9.[151,264,1913,1935]" captionTargetBox="[216,1355,230,1832]" captionTargetId="figure@9.[151,1436,157,1945]" captionTargetPageId="9" captionText="FIGURES 9 – 19. Habitus, dorsal view. 9, Hypotelus insulanus, syntype, male; 10, H. praecox, lectotype, female; 11, H. scheerpeltzi sp. nov., holotype, male; 12, H. testaceus, syntype, male; 13, H. melanodelta sp. nov., holotype, female; 14, H. castaneus sp. nov., holotype, male; 15, H. brevitarsus sp. nov., holotype, male; 16, H. corniculatus sp. nov., holotype, female; 17, H. marginatus, syntype, male; 18, H. laevis, male; 19, H. andinus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/803652/files/figure.png" pageId="11" pageNumber="462">Figs. 15</figureCitation>
, 
<figureCitation id="13492A1EFF9DD530CF2AFA01D30CFA3E" box="[264,294,1411,1435]" captionStart="FIGURES 36 – 47. 36 – 38" captionStartId="15.[151,264,1841,1863]" captionTargetBox="[213,1360,260,1787]" captionTargetId="figure@15.[176,1398,193,1850]" captionTargetPageId="15" captionText="FIGURES 36 – 47. 36 – 38, Hypotelus pusillus. 36, anterior leg, posterior view; 37, median leg, posterior view; 38, hind leg, posterior view. 39, H. brevitarsus sp. nov., metatarsus, lateral view. 40 – 45, H. pusillus. 40, tergites 1 – 3, dorsal view; 41, sternites 1 – 3, ventral view; 42, tergite 8, female, dorsal view, setae removed (drawn under microscope slides, plates were separated); 43, sternite 8, lateral view; 44, sternite 9, male, ventral view; 45, ovipositor, female, ventral view. 46 – 47, H. castaneus sp. nov. 46, abdominal segments 7 – 9, male, ventral view; 47, abdominal segments 7 – 8, female, ventral view. Scale bars = 0.125 mm, except Fig. 40 = 0.25 mm and Figs. 46 – 47 = 0.5 mm." httpUri="https://zenodo.org/record/803658/files/figure.png" pageId="11" pageNumber="462">39</figureCitation>
, 
<figureCitation id="13492A1EFF9DD530CF10FA00D37BFA3E" box="[306,337,1410,1435]" captionStart="FIGURES 48 – 58" captionStartId="16.[151,264,1844,1866]" captionTargetBox="[200,1385,250,1781]" captionTargetId="figure@16.[151,1436,193,1823]" captionTargetPageId="16" captionText="FIGURES 48 – 58. Tergites and sternites of males in dorsal and ventral view, respectively and right setae removed. 48, Hypotelus pusillus, tergite 8; 49, H. scheerpeltzi sp. nov., tergite 8; 50, H. castaneus sp. nov., tergite 8; 51, H. pusillus, sternite 8; 52, H. insulanus, sternite 8; 53, H. testaceus, sternite 8; 54, H. castaneus sp. nov., sternite 8; 55, H. pusillus, tergites 9 and 10; 56, H. insulanus, tergite 10; 57, H. castaneus sp. nov., tergite 10; 58, H. brevitarsus sp. nov., tergite 10. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803660/files/figure.png" pageId="11" pageNumber="462">58</figureCitation>
, 
<figureCitation id="13492A1EFF9DD530CF7FFA00D38EFA3E" box="[349,420,1410,1435]" captionStart="FIGURES 59 – 74" captionStartId="17.[151,264,1924,1946]" captionTargetBox="[290,1268,229,1830]" captionTargetId="figure@17.[237,1336,193,1893]" captionTargetPageId="17" captionText="FIGURES 59 – 74. Aedeagus, for each species with lateral view and ventral view, respectively. 59 – 60, Hypotelus pusillus; 61 – 62, H. insulanus; 63 – 64, H. marginatus; 65 – 66, H. testaceus; 67 – 68, H. praecox; 69 – 70, H. brevitarsus sp. nov., parameres removed in ventral view; 71 – 72, H. castaneus sp. nov.; 73 – 74, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803662/files/figure.png" pageId="11" pageNumber="462">69, 70</figureCitation>
, 
<figureCitation id="13492A1EFF9DD530CF92FA01D3E4FA3E" box="[432,462,1411,1435]" captionStart="FIGURES 75 – 82" captionStartId="18.[151,264,1403,1425]" captionTargetBox="[194,1368,254,1322]" captionTargetId="figure@18.[151,1436,193,1381]" captionTargetPageId="18" captionText="FIGURES 75 – 82. Sternite 8 of females in ventral view, right setae removed. 75, Hypotelus pusillus; 76, H. insulanus; 77, H. marginatus; 78 H. testaceus; 79, H. melanodelta sp. nov.; 80, H. brevitarsus sp. nov.; 81, H. castaneus sp. nov.; 82, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803664/files/figure.png" pageId="11" pageNumber="462">80</figureCitation>
, 
<figureCitation id="13492A1EFF9DD530CFF8FA01D009FA3E" box="[474,547,1411,1435]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="11" pageNumber="462">98, 99</figureCitation>
, 
<figureCitation id="13492A1EFF9DD530CC0CFA01D072FA3E" box="[558,600,1411,1435]" captionStart="FIGURE 114" captionStartId="31.[151,250,1701,1723]" captionTargetBox="[190,1375,215,1655]" captionTargetId="figure@31.[150,1437,153,1679]" captionTargetPageId="31" captionText="FIGURE 114. Geographical records of Hypotelus. Hypotelus andinus, Colombia, Ecuador and Bolivia; H. insulanus, Caribbean islands; H. marginatus, Colombia, Venezuela and Guatemala; H. testaceus, Panama; H. brevitarsus sp. nov., Panama. Countries without dots but colored represent the occurrence but no locality given." httpUri="https://zenodo.org/record/803674/files/figure.png" pageId="11" pageNumber="462">114</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BCD369BFF9DD530CEB5FA48D2CEF9D6" blockId="11.[151,1437,1482,2011]" pageId="11" pageNumber="462">
<materialsCitation id="3B1A3CC6FF9DD530CEB5FA48D2CEF9D6" ID-GBIF-Occurrence="1563407731" collectionCode="FMNH" collectorName="H. S. Dybas &amp; Bortoluzzi &amp; Caron" country="Panama" county="Lerida" elevation="1722" location="Chiriqui Prov." municipality="Finca" pageId="11" pageNumber="462" specimenCount="3" typeStatus="holotype">
<emphasis id="B906EA89FF9DD530CEB5FA48D36CFA46" bold="true" box="[151,326,1482,1507]" pageId="11" pageNumber="462">
<typeStatus id="54C98839FF9DD530CEB5FA48D2F8FA46" box="[151,210,1482,1507]" pageId="11" pageNumber="462">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C98839FF9DD530CF6CFA48D39CFA46" box="[334,438,1482,1507]" pageId="11" pageNumber="462" type="holotype">Holotype</typeStatus>
deposited in 
<collectionCode id="ED63AE5EFF9DD530CC70FA49D08CFA47" box="[594,678,1483,1506]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="462">FMNH</collectionCode>
, male (photo, 
<figureCitation id="13492A1EFF9DD530CD68FA48D1B7FA46" box="[842,925,1482,1507]" captionStart="FIGURES 9 – 19" captionStartId="9.[151,264,1913,1935]" captionTargetBox="[216,1355,230,1832]" captionTargetId="figure@9.[151,1436,157,1945]" captionTargetPageId="9" captionText="FIGURES 9 – 19. Habitus, dorsal view. 9, Hypotelus insulanus, syntype, male; 10, H. praecox, lectotype, female; 11, H. scheerpeltzi sp. nov., holotype, male; 12, H. testaceus, syntype, male; 13, H. melanodelta sp. nov., holotype, female; 14, H. castaneus sp. nov., holotype, male; 15, H. brevitarsus sp. nov., holotype, male; 16, H. corniculatus sp. nov., holotype, female; 17, H. marginatus, syntype, male; 18, H. laevis, male; 19, H. andinus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/803652/files/figure.png" pageId="11" pageNumber="462">Fig. 15</figureCitation>
), with labels: (1) “ 
<collectingCountry id="F365760BFF9DD530CA50FA49D6DAFA46" box="[1138,1264,1483,1507]" name="Panama" pageId="11" pageNumber="462">PANAMA</collectingCountry>
: 
<location id="8EAD6040FF9DD530CADEFA48D7B7FA47" LSID="urn:lsid:plazi:treatment:03DB878DFF9DD537CEB5FADDD773F9F5:8EAD6040FF9DD530CADEFA48D7B7FA47" box="[1276,1437,1482,1507]" country="Panama" county="Lerida" municipality="Finca" name="Chiriqui Prov." pageId="11" pageNumber="462">Chiriqui Prov.</location>
“ 
<location id="8EAD6040FF9DD530CE83FA6DD2CDF9A2" LSID="urn:lsid:plazi:treatment:03DB878DFF9DD537CEB5FADDD773F9F5:8EAD6040FF9DD530CE83FA6DD2CDF9A2" box="[161,231,1519,1543]" country="Panama" county="Lerida" municipality="Finca" name="Barca" pageId="11" pageNumber="462">Barca</location>
” area, 
<collectingMunicipality id="6BA9ACE1FF9DD530CF1FFA6CD351F9A2" box="[317,379,1518,1543]" pageId="11" pageNumber="462">Finca</collectingMunicipality>
/ 
<collectingCounty id="62AC4E17FF9DD530CFA1FA6CD3E6F9A2" box="[387,460,1518,1543]" pageId="11" pageNumber="462">Lerida</collectingCounty>
nr. Boquete./III:12:1959 
<elevation id="005FD1A8FF9DD530CCD7FA6CD165F9A2" box="[757,847,1518,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.72212" pageId="11" pageNumber="462" unit="ft" value="5650.0">
<quantity id="4C8A9B7EFF9DD530CCD7FA6CD165F9A2" box="[757,847,1518,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.72212" pageId="11" pageNumber="462" unit="ft" value="5650.0">5650 ft</quantity>
</elevation>
.” [white label, printed in black; day and elevation handwritten]; (2) “under slab on/pile of cut chips/and bark” [white label, printed in black]; (3) “leg./ 
<collectorName id="2687534DFF9DD530CB25F990D7B8F98E" box="[1287,1426,1554,1579]" pageId="11" pageNumber="462">H. S. Dybas</collectorName>
” [white label, printed in black]; (4) “ 
<typeStatus id="54C98839FF9DD530CC05F9B4D094F9EA" box="[551,702,1590,1615]" pageId="11" pageNumber="462" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="4C724D18FF9DD530CCE4F9B4D190F9EA" box="[710,954,1590,1615]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="brevitarsus">Hypotelus brevitarsus</taxonomicName>
/ 
<collectorName id="2687534DFF9DD530CDE1F9B4D612F9EA" box="[963,1080,1590,1615]" pageId="11" pageNumber="462">Bortoluzzi</collectorName>
&amp; 
<collectorName id="2687534DFF9DD530CA7FF9B5D68EF9EA" box="[1117,1188,1591,1615]" pageId="11" pageNumber="462">Caron</collectorName>
” [red label, printed in black].
</materialsCitation>
</paragraph>
<paragraph id="8BCD369BFF9DD537CEE5F9FDD3B6FF5D" blockId="11.[151,1437,1482,2011]" lastBlockId="12.[151,1437,151,1616]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
Paratypes: 11 specimens, deposited in FMNH. 
<materialsCitation id="3B1A3CC6FF9DD530CCF5F9FDD0DEF97A" ID-GBIF-Occurrence="1563407733" collectorName="Bortoluzzi &amp; Caron" country="Canada" location="Canada" pageId="11" pageNumber="462" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<specimenCount id="9D74FD12FF9DD530CCF5F9FDD10FF932" box="[727,805,1662,1687]" pageId="11" pageNumber="462" type="male">1 male</specimenCount>
, dissected, whole specimen fixed on acetate plastic card covered with 
<collectingCountry id="F365760BFF9DD530CF12F920D3AEF91E" box="[304,388,1698,1723]" name="Canada" pageId="11" pageNumber="462">Canada</collectingCountry>
balsam, with the same three first labels of 
<typeStatus id="54C98839FF9DD530CD45F920D1E4F91E" box="[871,974,1698,1723]" pageId="11" pageNumber="462" type="holotype">holotype</typeStatus>
: (4) “ 
<typeStatus id="54C98839FF9DD530CA2BF920D6B0F91E" box="[1033,1178,1698,1723]" pageId="11" pageNumber="462" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C724D18FF9DD530CA81F920D7B9F91E" box="[1187,1427,1698,1723]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="brevitarsus">Hypotelus brevitarsus</taxonomicName>
/ 
<collectorName id="2687534DFF9DD530CEB5F944D327F97A" box="[151,269,1734,1759]" pageId="11" pageNumber="462">Bortoluzzi</collectorName>
&amp; 
<collectorName id="2687534DFF9DD530CF17F945D356F97A" box="[309,380,1735,1759]" pageId="11" pageNumber="462">Caron</collectorName>
” [yellow label, printed in black]
</materialsCitation>
. 
<materialsCitation id="3B1A3CC6FF9DD530CD22F944D0DAF882" ID-GBIF-Occurrence="1563407730" collectorName="Bortoluzzi &amp; Caron" country="Canada" location="Canada" pageId="11" pageNumber="462" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="9D74FD12FF9DD530CD22F944D177F97A" box="[768,861,1734,1759]" pageId="11" pageNumber="462" type="female">1female</specimenCount>
, dissected, terminalia fixed on acetate plastic card covered with 
<collectingCountry id="F365760BFF9DD530CF14F968D3A0F8A6" box="[310,394,1770,1795]" name="Canada" pageId="11" pageNumber="462">Canada</collectingCountry>
balsam, with same three first labels of 
<typeStatus id="54C98839FF9DD530CD7FF968D1EEF8A6" box="[861,964,1770,1795]" pageId="11" pageNumber="462" type="holotype">holotype</typeStatus>
: (4) “ 
<typeStatus id="54C98839FF9DD530CA24F968D6BDF8A6" box="[1030,1175,1770,1795]" pageId="11" pageNumber="462" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C724D18FF9DD530CABDF968D7B9F8A6" box="[1183,1427,1770,1795]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="brevitarsus">Hypotelus brevitarsus</taxonomicName>
/ 
<collectorName id="2687534DFF9DD530CEB5F88CD327F882" box="[151,269,1806,1831]" pageId="11" pageNumber="462">Bortoluzzi</collectorName>
&amp; 
<collectorName id="2687534DFF9DD530CF16F88DD351F882" box="[308,379,1807,1831]" pageId="11" pageNumber="462">Caron</collectorName>
” [yellow label, printed in black]
</materialsCitation>
. 
<materialsCitation id="3B1A3CC6FF9DD530CCDEF88DD695F8CA" ID-GBIF-Occurrence="1563407726" collectorName="Bortoluzzi &amp; Caron" country="Canada" location="Canada" pageId="11" pageNumber="462" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="9D74FD12FF9DD530CCDEF88DD14BF882" box="[764,865,1806,1831]" pageId="11" pageNumber="462" type="female">1 female</specimenCount>
, dissected, terminalia fixed on acetate plastic card covered with 
<collectingCountry id="F365760BFF9DD530CF1AF8B0D3A6F8EE" box="[312,396,1842,1867]" name="Canada" pageId="11" pageNumber="462">Canada</collectingCountry>
balsam, with same three first labels of 
<typeStatus id="54C98839FF9DD530CD47F8B0D1E0F8EE" box="[869,970,1842,1867]" pageId="11" pageNumber="462" type="holotype">holotype</typeStatus>
, except for the date: “III:14:1959.”: (4) “ 
<typeStatus id="54C98839FF9DD530CE82F8D4D31BF8CA" box="[160,305,1878,1903]" pageId="11" pageNumber="462" type="paratype">PARATYPE</typeStatus>
/ 
<taxonomicName id="4C724D18FF9DD530CF1BF8D4D019F8CA" box="[313,563,1878,1903]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="brevitarsus">Hypotelus brevitarsus</taxonomicName>
/ 
<collectorName id="2687534DFF9DD530CC19F8D4D09AF8CA" box="[571,688,1878,1903]" pageId="11" pageNumber="462">Bortoluzzi</collectorName>
&amp; 
<collectorName id="2687534DFF9DD530CCC1F8D5D100F8CA" box="[739,810,1879,1903]" pageId="11" pageNumber="462">Caron</collectorName>
” [yellow label, printed in black]
</materialsCitation>
. 7 specimens, sex undetermined, with same three first labels of holotype: (4) “PARATYPE/ 
<taxonomicName id="4C724D18FF9DD530CA26F8F8D6D1F836" box="[1028,1275,1914,1939]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="brevitarsus">Hypotelus brevitarsus</taxonomicName>
/Bortoluzzi &amp; Caron” [yellow label, printed in black]; one specimen of them with additional label: (4) “ 
<taxonomicName id="4C724D18FF9DD530CAB6F81CD721F812" box="[1172,1291,1950,1975]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="genus">Hypotelus</taxonomicName>
/det. Newton 1994” [white label, first line, handwritten; second line, printed in black]; (5) “PARATYPE/ 
<taxonomicName id="4C724D18FF9DD530CA80F840D7B9F87E" box="[1186,1427,1986,2011]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="brevitarsus">Hypotelus brevitarsus</taxonomicName>
/ Bortoluzzi &amp; Caron” [yellow label, printed in black]. 1 specimen, sex undetermined, with same three first labels of holotype, except for the date: “III:14:1959.”: (4) “PARATYPE/ 
<taxonomicName id="4C724D18FF9AD537CD7EFF3ED664FF70" box="[860,1102,188,213]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="brevitarsus">Hypotelus brevitarsus</taxonomicName>
/Bortoluzzi &amp; Caron” [yellow label, printed in black].
</paragraph>
<paragraph id="8BCD369BFF9AD537CEE5FE86D050FEB9" blockId="12.[151,1437,151,1616]" box="[199,634,260,285]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FE86D393FEB8" bold="true" box="[199,441,260,285]" pageId="12" pageNumber="463">Additional material.</emphasis>
See Appendix 2.
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9AD537CEE5FEA5D67CFE75" pageId="12" pageNumber="463" type="diagnosis">
<paragraph id="8BCD369BFF9AD537CEE5FEA5D67CFE75" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FEA5D36BFEE5" bold="true" box="[199,321,295,320]" pageId="12" pageNumber="463">Diagnosis.</emphasis>
<taxonomicName id="4C724D18FF9AD537CF68FEA5D06BFEE5" box="[330,577,295,320]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="brevitarsus" status="sp. nov.">
<emphasis id="B906EA89FF9AD537CF68FEA5D06BFEE5" box="[330,577,295,320]" italics="true" pageId="12" pageNumber="463">Hypotelus brevitarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F2FF9AD537CC68FEA5D08FFEE5" box="[586,677,295,320]" pageId="12" pageNumber="463" rank="species">
<emphasis id="B906EA89FF9AD537CC68FEA5D08FFEE5" bold="true" box="[586,677,295,320]" pageId="12" pageNumber="463">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar to 
<taxonomicName id="4C724D18FF9AD537CD1DFEABD191FEE5" box="[831,955,295,320]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B906EA89FF9AD537CD1DFEABD191FEE5" box="[831,955,295,320]" italics="true" pageId="12" pageNumber="463">H. pusillus</emphasis>
</taxonomicName>
and differs by the antennal scape of male without prominent tooth on inner face; the antennae shorter, not reaching half-length of elytra; and tergite 10 with lateral sides at posterior margin emarginated in both sexes (
<figureCitation id="13492A1EFF9AD537CD0DFEEDD1A6FE2D" box="[815,908,367,392]" captionStart="FIGURES 48 – 58" captionStartId="16.[151,264,1844,1866]" captionTargetBox="[200,1385,250,1781]" captionTargetId="figure@16.[151,1436,193,1823]" captionTargetPageId="16" captionText="FIGURES 48 – 58. Tergites and sternites of males in dorsal and ventral view, respectively and right setae removed. 48, Hypotelus pusillus, tergite 8; 49, H. scheerpeltzi sp. nov., tergite 8; 50, H. castaneus sp. nov., tergite 8; 51, H. pusillus, sternite 8; 52, H. insulanus, sternite 8; 53, H. testaceus, sternite 8; 54, H. castaneus sp. nov., sternite 8; 55, H. pusillus, tergites 9 and 10; 56, H. insulanus, tergite 10; 57, H. castaneus sp. nov., tergite 10; 58, H. brevitarsus sp. nov., tergite 10. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803660/files/figure.png" pageId="12" pageNumber="463">Figs. 58</figureCitation>
, 
<figureCitation id="13492A1EFF9AD537CDB5FEF2D19CFE2D" box="[919,950,368,392]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="12" pageNumber="463">98</figureCitation>
), besides the genital structures (see below). 
<taxonomicName id="4C724D18FF9AD537CEB5FE17D31DFE08" box="[151,311,404,429]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="brevitarsus" status="sp. nov.">
<emphasis id="B906EA89FF9AD537CEB5FE17D31DFE08" box="[151,311,404,429]" italics="true" pageId="12" pageNumber="463">H. brevitarsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557F2FF9AD537CF1DFE16D3B0FE08" box="[319,410,404,429]" pageId="12" pageNumber="463" rank="species">
<emphasis id="B906EA89FF9AD537CF1DFE16D3B0FE08" bold="true" box="[319,410,404,429]" pageId="12" pageNumber="463">sp. nov.</emphasis>
</taxonomicNameLabel>
may be distinguished from other species of 
<taxonomicName id="4C724D18FF9AD537CDB6FE16D62CFE08" box="[916,1030,404,429]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA89FF9AD537CDB6FE16D62CFE08" box="[916,1030,404,429]" italics="true" pageId="12" pageNumber="463">Hypotelus</emphasis>
</taxonomicName>
by the metatarsomeres 5 somewhat shorter and transverse (
<figureCitation id="13492A1EFF9AD537CFBEFE35D3C5FE75" box="[412,495,439,464]" captionStart="FIGURES 36 – 47. 36 – 38" captionStartId="15.[151,264,1841,1863]" captionTargetBox="[213,1360,260,1787]" captionTargetId="figure@15.[176,1398,193,1850]" captionTargetPageId="15" captionText="FIGURES 36 – 47. 36 – 38, Hypotelus pusillus. 36, anterior leg, posterior view; 37, median leg, posterior view; 38, hind leg, posterior view. 39, H. brevitarsus sp. nov., metatarsus, lateral view. 40 – 45, H. pusillus. 40, tergites 1 – 3, dorsal view; 41, sternites 1 – 3, ventral view; 42, tergite 8, female, dorsal view, setae removed (drawn under microscope slides, plates were separated); 43, sternite 8, lateral view; 44, sternite 9, male, ventral view; 45, ovipositor, female, ventral view. 46 – 47, H. castaneus sp. nov. 46, abdominal segments 7 – 9, male, ventral view; 47, abdominal segments 7 – 8, female, ventral view. Scale bars = 0.125 mm, except Fig. 40 = 0.25 mm and Figs. 46 – 47 = 0.5 mm." httpUri="https://zenodo.org/record/803658/files/figure.png" pageId="12" pageNumber="463">Fig. 39</figureCitation>
), although in some specimens this is not so prominent.
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9AD537CEE5FE5ED3E5FA95" pageId="12" pageNumber="463" type="description">
<paragraph id="8BCD369BFF9AD537CEE5FE5ED05FFDFA" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FE5ED373FE50" bold="true" box="[199,345,476,501]" pageId="12" pageNumber="463">Description.</emphasis>
BL: 
<quantity id="4C8A9B7EFF9AD537CFB7FE5ED035FE51" box="[405,543,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="12" pageNumber="463" unit="mm" value="2.55" valueMax="2.9" valueMin="2.2">2.2–2.9 mm</quantity>
, BW: 
<quantity id="4C8A9B7EFF9AD537CC4BFE5ED0DFFE51" box="[617,757,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="12" pageNumber="463" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
. Body slightly convex; dorsal surface glossy; brown (elytra yellow); appendages lighter, except mandibles. Dorsal integument of head and pronotum with dispersed fine punctures and undulate microstriae only on margins; elytra with dispersed fine punctures and only one longitudinal finely punctate stria close to elytral suture.
</paragraph>
<paragraph id="8BCD369BFF9AD537CEE5FDEED61BFD68" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FDEED366FD20" box="[199,332,620,645]" italics="true" pageId="12" pageNumber="463">Male. Head</emphasis>
. Supra-antennal area slightly prominent. Antennae reaching humeral angle; antennomeres 5, 7, 9 and 11 with longest setae on inner face; antennomeres 2 and 3 of equal length, 5–11 gradually increasing in length toward antennal apex. Mandibles symmetrical. Mentum 1.6 times as wide as long.
</paragraph>
<paragraph id="8BCD369BFF9AD537CEE5FD55D32AFC25" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FD55D30AFD55" box="[199,288,727,752]" italics="true" pageId="12" pageNumber="463">Thorax.</emphasis>
Pronotum wider than long (PW/PL=1.3); anterior angles rounded and slightly prominent; apical half with somewhat parallel sides and basal half gradually narrowing toward the base; with complete internal midlongitudinal ridge and slight longitudinal median sulcus only on basal half. Elytra somewhat longer than wide (EL/ BW=1.1), covering partially or not abdominal tergite 3. Metatarsomeres 5 somewhat shorter and transverse (
<figureCitation id="13492A1EFF9AD537CEBDFCE5D2DEFC25" box="[159,244,871,896]" captionStart="FIGURES 36 – 47. 36 – 38" captionStartId="15.[151,264,1841,1863]" captionTargetBox="[213,1360,260,1787]" captionTargetId="figure@15.[176,1398,193,1850]" captionTargetPageId="15" captionText="FIGURES 36 – 47. 36 – 38, Hypotelus pusillus. 36, anterior leg, posterior view; 37, median leg, posterior view; 38, hind leg, posterior view. 39, H. brevitarsus sp. nov., metatarsus, lateral view. 40 – 45, H. pusillus. 40, tergites 1 – 3, dorsal view; 41, sternites 1 – 3, ventral view; 42, tergite 8, female, dorsal view, setae removed (drawn under microscope slides, plates were separated); 43, sternite 8, lateral view; 44, sternite 9, male, ventral view; 45, ovipositor, female, ventral view. 46 – 47, H. castaneus sp. nov. 46, abdominal segments 7 – 9, male, ventral view; 47, abdominal segments 7 – 8, female, ventral view. Scale bars = 0.125 mm, except Fig. 40 = 0.25 mm and Figs. 46 – 47 = 0.5 mm." httpUri="https://zenodo.org/record/803658/files/figure.png" pageId="12" pageNumber="463">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="8BCD369BFF9AD537CEE5FC0ED6C3FBB5" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FC0ED312FC00" box="[199,312,908,933]" italics="true" pageId="12" pageNumber="463">Abdomen.</emphasis>
Tergite 8 with posterior margin rounded; sternite 8 with posterior margin rounded; tergite 9 with short ventral struts; sternite 9 with posterior margin truncate and with two pairs of long setae; tergite 10 with posterior and lateral margins weakly pigmented, with short fringes and four setae on each side (
<figureCitation id="13492A1EFF9AD537CAFEFC56D71BFC48" box="[1244,1329,980,1005]" captionStart="FIGURES 48 – 58" captionStartId="16.[151,264,1844,1866]" captionTargetBox="[200,1385,250,1781]" captionTargetId="figure@16.[151,1436,193,1823]" captionTargetPageId="16" captionText="FIGURES 48 – 58. Tergites and sternites of males in dorsal and ventral view, respectively and right setae removed. 48, Hypotelus pusillus, tergite 8; 49, H. scheerpeltzi sp. nov., tergite 8; 50, H. castaneus sp. nov., tergite 8; 51, H. pusillus, sternite 8; 52, H. insulanus, sternite 8; 53, H. testaceus, sternite 8; 54, H. castaneus sp. nov., sternite 8; 55, H. pusillus, tergites 9 and 10; 56, H. insulanus, tergite 10; 57, H. castaneus sp. nov., tergite 10; 58, H. brevitarsus sp. nov., tergite 10. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803660/files/figure.png" pageId="12" pageNumber="463">Fig. 58</figureCitation>
). Median lobe of aedeagus with bulbous base in ventral view and curved shape in lateral view (
<figureCitation id="13492A1EFF9AD537CA71FC75D6F7FBB5" box="[1107,1245,1015,1040]" captionStart="FIGURES 59 – 74" captionStartId="17.[151,264,1924,1946]" captionTargetBox="[290,1268,229,1830]" captionTargetId="figure@17.[237,1336,193,1893]" captionTargetPageId="17" captionText="FIGURES 59 – 74. Aedeagus, for each species with lateral view and ventral view, respectively. 59 – 60, Hypotelus pusillus; 61 – 62, H. insulanus; 63 – 64, H. marginatus; 65 – 66, H. testaceus; 67 – 68, H. praecox; 69 – 70, H. brevitarsus sp. nov., parameres removed in ventral view; 71 – 72, H. castaneus sp. nov.; 73 – 74, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803662/files/figure.png" pageId="12" pageNumber="463">Figs. 69–70</figureCitation>
).
</paragraph>
<paragraph id="8BCD369BFF9AD537CEE5FB9ED769FB05" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FB9ED309FB90" box="[199,291,1052,1077]" italics="true" pageId="12" pageNumber="463">Female.</emphasis>
Similar to male except for: without very long setae on antennomeres 5, 7, 9 and 11; abdominal sternite 8 with posterior margin somewhat truncate and posterio-laterally slightly emarginated, with short setae (
<figureCitation id="13492A1EFF9AD537CB1BFBBDD7A5FBFD" box="[1337,1423,1087,1112]" captionStart="FIGURES 75 – 82" captionStartId="18.[151,264,1403,1425]" captionTargetBox="[194,1368,254,1322]" captionTargetId="figure@18.[151,1436,193,1381]" captionTargetPageId="18" captionText="FIGURES 75 – 82. Sternite 8 of females in ventral view, right setae removed. 75, Hypotelus pusillus; 76, H. insulanus; 77, H. marginatus; 78 H. testaceus; 79, H. melanodelta sp. nov.; 80, H. brevitarsus sp. nov.; 81, H. castaneus sp. nov.; 82, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803664/files/figure.png" pageId="12" pageNumber="463">Fig. 80</figureCitation>
); tergite 9 without ventral struts; bursa copulatrix as 
<taxonomicName id="4C724D18FF9AD537CCF0FBE7D161FBD8" box="[722,843,1124,1149]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B906EA89FF9AD537CCF0FBE7D161FBD8" box="[722,843,1124,1149]" italics="true" pageId="12" pageNumber="463">H. pusillus</emphasis>
</taxonomicName>
; ovipositor consisting of a pair of weakly pigmented hemisternites and a pair of more apical coxites, and with many long setae on apex; spermatheca as 
<figureCitation id="13492A1EFF9AD537CACEFB05D714FB05" box="[1260,1342,1159,1184]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="12" pageNumber="463">Fig. 99</figureCitation>
.
</paragraph>
<paragraph id="8BCD369BFF9AD537CEE5FB2ED10EFB61" blockId="12.[151,1437,151,1616]" box="[199,804,1196,1221]" pageId="12" pageNumber="463">
<materialsCitation id="3B1A3CC6FF9AD537CEE5FB2ED10EFB61" ID-GBIF-Occurrence="1563407744" box="[199,804,1196,1221]" country="Panama" location="Chiriqui" pageId="12" pageNumber="463" specimenCount="1" stateProvince="Chiriqui">
<emphasis id="B906EA89FF9AD537CEE5FB2ED3FAFB60" bold="true" box="[199,464,1196,1221]" pageId="12" pageNumber="463">Geographical records.</emphasis>
<collectingCountry id="F365760BFF9AD537CFF5FB2FD01AFB61" box="[471,560,1197,1220]" name="Panama" pageId="12" pageNumber="463">Panama</collectingCountry>
(
<collectingRegion id="49B6F879FF9AD537CC1CFB2ED0B5FB60" box="[574,671,1196,1221]" country="Panama" name="Chiriqui" pageId="12" pageNumber="463">Chiriqui</collectingRegion>
) (
<figureCitation id="13492A1EFF9AD537CC94FB2ED132FB61" box="[694,792,1196,1221]" captionStart="FIGURE 114" captionStartId="31.[151,250,1701,1723]" captionTargetBox="[190,1375,215,1655]" captionTargetId="figure@31.[150,1437,153,1679]" captionTargetPageId="31" captionText="FIGURE 114. Geographical records of Hypotelus. Hypotelus andinus, Colombia, Ecuador and Bolivia; H. insulanus, Caribbean islands; H. marginatus, Colombia, Venezuela and Guatemala; H. testaceus, Panama; H. brevitarsus sp. nov., Panama. Countries without dots but colored represent the occurrence but no locality given." httpUri="https://zenodo.org/record/803674/files/figure.png" pageId="12" pageNumber="463">Fig. 114</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCD369BFF9AD537CEE5FB4DD3E5FA95" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FB4DD3A6FB4D" bold="true" box="[199,396,1231,1256]" pageId="12" pageNumber="463">Biological notes.</emphasis>
This species was collected at elevations between 
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(
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) and 
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(
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), under a slab on a pile of cut chips and bark, in a split sapling, in scraping, in torn fibers of a wounded tree and under bark of a log on the ground.
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9AD537CEE5FABED32FF9AD" pageId="12" pageNumber="463" type="discussion">
<paragraph id="8BCD369BFF9AD537CEE5FABED15BFADD" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5FABED311FAF0" bold="true" box="[199,315,1340,1365]" pageId="12" pageNumber="463">Remarks.</emphasis>
Some specimens may have the metatarsomeres a little less transverse. The longest setae on inner face of antennomeres 5, 7, 9 and 11 are sometimes not so visible.
</paragraph>
<paragraph id="8BCD369BFF9AD537CEB5FA07D32FF9AD" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
In the material examined, there are five specimens collected from 
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(
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), which do not have the metatarsomeres 5 shorter and transverse, and we are unable to segregate these 
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into more than one species. The feature metatarsomere 5 shorter and transverse is only visible on the specimens collected above 
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(
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).
</paragraph>
</subSubSection>
<subSubSection id="C3686510FF9AD537CEE5F996D773F9F5" pageId="12" pageNumber="463" type="etymology">
<paragraph id="8BCD369BFF9AD537CEE5F996D773F9F5" blockId="12.[151,1437,151,1616]" pageId="12" pageNumber="463">
<emphasis id="B906EA89FF9AD537CEE5F996D364F988" bold="true" box="[199,334,1556,1581]" pageId="12" pageNumber="463">Etymology.</emphasis>
The specific name refers to the shape of the metatarsus and is a compound name: Latin adjective 
<emphasis id="B906EA89FF9AD537CEB5F9B5D2E2F9F5" box="[151,200,1591,1616]" italics="true" pageId="12" pageNumber="463">brev</emphasis>
- (short), the connective vowel 
<emphasis id="B906EA89FF9AD537CC07F9BBD007F9F5" box="[549,557,1593,1616]" italics="true" pageId="12" pageNumber="463">i</emphasis>
, and the Greek noun latinized to 
<emphasis id="B906EA89FF9AD537CD80F9BBD1CCF9F5" box="[930,998,1593,1616]" italics="true" pageId="12" pageNumber="463">tarsus</emphasis>
(foot). It is a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>