267 lines
40 KiB
XML
267 lines
40 KiB
XML
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<mods:title id="08DDC134D6887F98D6E65CB66F7FCCC9">Taxonomy of the freshwater crab Coccusa cristicervix Ng & Grinang, 2004 (Decapoda: Brachyura: Gecarcinucidae), with description of a new species of Thelphusula from Sarawak, Malaysia, Borneo</mods:title>
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<mods:namePart id="8A913EF0CCE6065BF3E362B71D882EF2">Grinang, Jongkar</mods:namePart>
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<mods:namePart id="552DE4CCAB80B82874F52BDF3DCD8F82">Ng, Peter K. L.</mods:namePart>
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<mods:title id="26400589DBC718C1D83D3E834757F911">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="FF5AC98279CD3B2C4278FCEA4282E5FE">2014</mods:date>
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<paragraph id="8BB8C71D7024FFB8FF66FB56C77F48A8" blockId="1.[163,702,1182,1269]" pageId="1" pageNumber="390">
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<taxonomicName id="4C07BC9E7024FFB8FF66FB56C7D548C8" ID-CoL="5699Y" authority="(Ng & Grinang, 2004)" authorityName="Grinang & Ng" authorityYear="2014" baseAuthorityName="Ng & Grinang" baseAuthorityYear="2004" box="[163,697,1182,1205]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="cristicervix" status="comb. nov.">
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<emphasis id="B9731B0F7024FFB8FF66FB56C4DC48C8" bold="true" box="[163,432,1182,1205]" italics="true" pageId="1" pageNumber="390">Thelphusula cristicervix</emphasis>
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(
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<bibRefCitation id="EF96BAEC7024FFB8FE05FB56C7DE48C8" author="Ng PKL & Grinang J" box="[448,690,1182,1205]" pageId="1" pageNumber="390" pagination="299 - 325" refId="ref4094" refString="Ng PKL & Grinang J (2004) Decapod crustaceans with descriptions of three new species. In: Yong HS Ng FSP & Yen EEL (eds.) Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, 59, No. 80 (New Series), Special Issue No. 6: 299 - 325." type="book chapter" year="2004">Ng & Grinang, 2004</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="A240A6747024FFB8FE8BFB76C77F48A8" box="[334,531,1214,1237]" pageId="1" pageNumber="390" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C31D94967024FFB8FE88FB16C7784888" box="[333,532,1246,1269]" pageId="1" pageNumber="390" type="description">
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<paragraph id="8BB8C71D7024FFB8FE88FB16C7784888" blockId="1.[163,702,1182,1269]" box="[333,532,1246,1269]" pageId="1" pageNumber="390">
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(
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<figureCitation id="133CDB987024FFB8FE90FB16C4CD4888" box="[341,417,1246,1269]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="1" pageNumber="390">Figs. 1</figureCitation>
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,
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<figureCitation id="133CDB987024FFB8FE75FB16C49C4888" box="[432,496,1246,1269]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="1" pageNumber="390">3A–E</figureCitation>
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,
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<figureCitation id="133CDB987024FFB8FE3BFB16C7604888" box="[510,524,1246,1269]" captionStart="Fig" captionStartId="5.[113,148,1245,1266]" captionTargetBox="[115,752,520,1221]" captionTargetId="figure-13@5.[113,752,518,1221]" captionTargetPageId="5" captionText="Fig. 4. Colours in life, Thelphusula cristicervix (Ng & Grinang, 2004). A, male (ca. 19.0 mm carapace width), in situ in forest, specimen not collected; B, female (19.3 × 14.2 mm) (ZRC) [Photograph by: Tan HH]. Both specimens from Sama Jaya Nature Reserve, Kuching Division, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5353788" httpUri="https://zenodo.org/record/5353788/files/figure.png" pageId="1" pageNumber="390">4</figureCitation>
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)
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</paragraph>
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<subSubSection id="C31D94967024FFB8FFB4FAD5C4214916" pageId="1" pageNumber="390" type="reference_group">
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<paragraph id="8BB8C71D7024FFB8FFB4FAD5C4214916" blockId="1.[113,752,1309,1387]" pageId="1" pageNumber="390">
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<treatmentCitationGroup id="AB17E0337024FFB8FFB4FAD5C4214916" pageId="1" pageNumber="390">
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<taxonomicName id="4C07BC9E7024FFB8FFB4FAD5C77C494F" ID-CoL="WMS2" authority="Ng & Grinang, 2004" authorityName="Ng & Grinang" authorityYear="2004" box="[113,528,1309,1330]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="cristicervix">
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<emphasis id="B9731B0F7024FFB8FFB4FAD5C454494F" box="[113,312,1309,1330]" italics="true" pageId="1" pageNumber="390">Coccusa cristicervix</emphasis>
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<bibRefCitation id="EF96BAEC7024FFB8FEFAFAD5C77C494F" author="Ng PKL & Grinang J" box="[319,528,1309,1330]" pageId="1" pageNumber="390" pagination="299 - 325" refId="ref4094" refString="Ng PKL & Grinang J (2004) Decapod crustaceans with descriptions of three new species. In: Yong HS Ng FSP & Yen EEL (eds.) Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, 59, No. 80 (New Series), Special Issue No. 6: 299 - 325." type="book chapter" year="2004">Ng & Grinang, 2004</bibRefCitation>
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</taxonomicName>
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: fig. 5; Ng, 2004: 321;
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<treatmentCitation id="0AA6E10C7024FFB8FF52FAF1C4354933" author="Ng PKL & Yeo DCJ" box="[151,345,1337,1358]" page="9" pageId="1" pageNumber="390" year="2007">
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<bibRefCitation id="EF96BAEC7024FFB8FF52FAF1C4354933" author="Ng PKL & Yeo DCJ" box="[151,345,1337,1358]" pageId="1" pageNumber="390" pagination="95 - 120" refId="ref4256" refString="Ng PKL & Yeo DCJ (2007) Malaysian freshwater crabs: conservation prospects and challenges. In: Chua L (ed.) Proceedings of the Seminar on the Status of Biological Diversity in Malaysia and Threat Assessment of Plant Species in Malaysia, 28 - 30 June 2005. Forest Research Institute Malaysia, Kepong. Pp. 95 - 120." type="book chapter" year="2007">Ng & Yeo, 2007: 9</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA6E10C7024FFB8FEA3FAF1C74B4933" author="Ng PKL & Guinot D & Davie PJF" box="[358,551,1337,1358]" page="69" pageId="1" pageNumber="390" year="2008">
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<bibRefCitation id="EF96BAEC7024FFB8FEA3FAF1C74B4933" author="Ng PKL & Guinot D & Davie PJF" box="[358,551,1337,1358]" pageId="1" pageNumber="390" pagination="1 - 286" refId="ref4055" refString="Ng PKL, Guinot D & Davie PJF (2008) Systema Brachyurorum Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 69</bibRefCitation>
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</treatmentCitation>
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;
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||
<bibRefCitation id="EF96BAEC7024FFB8FDF6FAF1C5A74916" author="Cumberlidge N & Ng PKL & Yeo DCJ & Magalhaes C & Campos MR & Alvarez F & Naruse T & Daniels SR & Esser LJ & Attipoe FYK & Clotilde-Ba F-L & Darwall W & Mcivor A & Collen B & Ram M" pageId="1" pageNumber="390" pagination="1665 - 1673" refId="ref3511" refString="Cumberlidge N, Ng PKL, Yeo DCJ, Magalhaes C, Campos MR, Alvarez F, Naruse T, Daniels SR, Esser LJ, Attipoe FYK, Clotilde-Ba F-L, Darwall W, Mcivor A, Collen B & Ram M (2009) Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation, 142: 1665 - 1673." type="journal article" year="2009">Cumberlidge et al., 2009</bibRefCitation>
|
||
: Appendix 1.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D94967024FFB8FFB4FA58C77C4AD5" pageId="1" pageNumber="390" type="materials_examined">
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<emphasis id="B9731B0F7024FFB8FFB4FA58C43849D8" bold="true" box="[113,340,1424,1445]" pageId="1" pageNumber="390">Material examined.</emphasis>
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<materialsCitation id="3B6FCD407024FFB8FE9AFA58C77C4AD5" collectingDate="2002-10-02" collectingDateMax="2013-09-13" collectingDateMin="2002-10-02" collectionCode="SBC, ZRC" collectorName="A. Denis & J. Grinang & Kuching Division & P. K. L. Ng" country="Malaysia" county="Sama Jaya Nature Reserve" elevation="16" location="Gunung Doya" municipality="Sama Jaya Nature Reserve" pageId="1" pageNumber="390" specimenCount="10" specimenCount-female="4" specimenCount-male="6" typeStatus="holotype">
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<typeStatus id="54BC79BF7024FFB8FE9AFA58C4AB49D8" box="[351,455,1424,1445]" pageId="1" pageNumber="390" type="holotype">Holotype</typeStatus>
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: female (19.9 ×
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) (SBC.C.00130),
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, limestone outcrop,
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<location id="8ED891C67024FFB8FDB4FA65C78049BF" LSID="urn:lsid:plazi:treatment:03AE760B7024FFB8FF66FB56C6A24AA8:8ED891C67024FFB8FDB4FA65C78049BF" box="[625,748,1453,1474]" country="Malaysia" county="Sama Jaya Nature Reserve" municipality="Sama Jaya Nature Reserve" name="Bau District" pageId="1" pageNumber="390">Bau District</location>
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,
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<collectingCountry id="F310878D7024FFB8FFB4FA01C5AA49A3" box="[113,198,1481,1502]" name="Malaysia" pageId="1" pageNumber="390">Sarawak</collectingCountry>
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, coll.
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<collectorName id="26F2A2CB7024FFB8FEC0FA01C43249A3" box="[261,350,1481,1502]" pageId="1" pageNumber="390">A. Denis</collectorName>
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et al.,
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.
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<location id="8ED891C67024FFB8FD96FA01C7FB49A3" LSID="urn:lsid:plazi:treatment:03AE760B7024FFB8FF66FB56C6A24AA8:8ED891C67024FFB8FD96FA01C7FB49A3" box="[595,663,1481,1502]" country="Malaysia" county="Sama Jaya Nature Reserve" municipality="Sama Jaya Nature Reserve" name="Others" pageId="1" pageNumber="390">Others</location>
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:
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(20.0 ×
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, 21.8 ×
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) (
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<collectionCode id="ED165FD87024FFB8FE35FA2EC74F4986" box="[496,547,1510,1531]" country="USA" lsid="urn:lsid:biocol.org:col:12519" name="University of California" pageId="1" pageNumber="390" type="Herbarium">SBC</collectionCode>
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),
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<collectingMunicipality id="6BDC5D677024FFB8FDF2FA2EC5A94A65" pageId="1" pageNumber="390">Sama Jaya Nature Reserve</collectingMunicipality>
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, lowland heath swamp forest,
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above sea level
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</elevation>
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, Kuching Division,
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, coll.
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<collectorName id="26F2A2CB7024FFB8FE7EF9E8C74D4A48" box="[443,545,1568,1589]" pageId="1" pageNumber="390">J. Grinang</collectorName>
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et al.,
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<date id="FFB9E1DD7024FFB8FDA1F9E8C7B54A48" box="[612,729,1568,1589]" pageId="1" pageNumber="390" value="2013-07-03">
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<collectingDate id="EFFD18357024FFB8FDA1F9E8C7B54A48" box="[612,729,1568,1589]" pageId="1" pageNumber="390" value="2013-07-03">3 July 2013</collectingDate>
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</date>
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;
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<specimenCount id="9D010C947024FFB8FD21F9E8C5C54A2F" pageId="1" pageNumber="390" type="male">4 males</specimenCount>
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(
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|
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<quantity id="4CFF6AF87024FFB8FE99F9F5C4864A2F" box="[348,490,1597,1618]" metricMagnitude="-2" metricUnit="m" metricValue="1.305" metricValueMax="1.48" metricValueMin="1.13" pageId="1" pageNumber="390" unit="mm" value="13.05" valueMax="14.8" valueMin="11.3">11.3–14.8 mm</quantity>
|
||
),
|
||
<specimenCount id="9D010C947024FFB8FE3FF9F5C73B4A2F" box="[506,599,1597,1618]" pageId="1" pageNumber="390" type="female">4 females</specimenCount>
|
||
(
|
||
<quantity id="4CFF6AF87024FFB8FDA6F9F5C79C4A2F" box="[611,752,1597,1618]" metricMagnitude="-2" metricUnit="m" metricValue="2.195" metricValueMax="2.48" metricValueMin="1.91" pageId="1" pageNumber="390" unit="mm" value="21.95" valueMax="24.8" valueMin="19.1">19.1–24.8 mm</quantity>
|
||
<emphasis id="B9731B0F7024FFB8FFB4F991C5134A13" bold="true" box="[113,127,1625,1646]" pageId="1" pageNumber="390">×</emphasis>
|
||
<quantity id="4CFF6AF87024FFB8FF41F991C4634A13" box="[132,271,1625,1646]" metricMagnitude="-2" metricUnit="m" metricValue="1.6649999999999998" metricValueMax="1.8699999999999999" metricValueMin="1.46" pageId="1" pageNumber="390" unit="mm" value="16.65" valueMax="18.7" valueMin="14.6">14.6–18.7 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED165FD87024FFB8FEE4F991C4384A13" box="[289,340,1625,1646]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="390">ZRC</collectionCode>
|
||
),
|
||
<collectingCounty id="62D9BF917024FFB8FEA6F991C70F4A13" box="[355,611,1625,1646]" pageId="1" pageNumber="390">Sama Jaya Nature Reserve</collectingCounty>
|
||
, lowland heath swamp forest,
|
||
<elevation id="002A202E7024FFB8FEC4F9BEC4BE4AF6" box="[257,466,1654,1675]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="390" unit="m" value="16.0">
|
||
<quantity id="4CFF6AF87024FFB8FEC4F9BEC45F4AF6" box="[257,307,1654,1675]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="390" unit="m" value="16.0">16 m</quantity>
|
||
above sea level
|
||
</elevation>
|
||
,
|
||
<collectorName id="26F2A2CB7024FFB8FE18F9BEC7E04AF6" box="[477,652,1654,1675]" pageId="1" pageNumber="390">Kuching Division</collectorName>
|
||
,
|
||
<collectingCountry id="F310878D7024FFB8FD52F9BEC7804AF6" box="[663,748,1654,1675]" name="Malaysia" pageId="1" pageNumber="390">Sarawak</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26F2A2CB7024FFB8FF66F95BC4694AD5" box="[163,261,1683,1704]" pageId="1" pageNumber="390">P.K.L. Ng</collectorName>
|
||
et al.,
|
||
<date id="FFB9E1DD7024FFB8FE8EF95BC7604AD5" box="[331,524,1683,1704]" pageId="1" pageNumber="390" value="2013-09-13">
|
||
<collectingDate id="EFFD18357024FFB8FE8EF95BC7604AD5" box="[331,524,1683,1704]" pageId="1" pageNumber="390" value="2013-09-13">13 September 2013</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D94967024FFB8FFB4F919C6D64DA8" pageId="1" pageNumber="390" type="diagnosis">
|
||
<paragraph id="8BB8C71D7024FFB8FFB4F919C0EC4D48" blockId="1.[113,753,1745,2088]" lastBlockId="1.[813,1452,190,309]" pageId="1" pageNumber="390">
|
||
<emphasis id="B9731B0F7024FFB8FFB4F919C5854A95" bold="true" box="[113,233,1745,1768]" pageId="1" pageNumber="390">Diagnosis.</emphasis>
|
||
Carapace gently ovate; cervical groove deep, prominent, with distinct rows of cristae running along length; postorbital cristae low, rounded, not sharp, separated from low, rounded epigastric cristae low by shallow depressions; anterolateral margin entire, epibranchial teeth not discernible; H-shaped depression relatively shallow (
|
||
<figureCitation id="133CDB987024FFB8FD99F8B9C78E4BF5" box="[604,738,1905,1928]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="1" pageNumber="390">Fig. 1A, B</figureCitation>
|
||
). Frontal margin deflexed; frontal median triangle not clearly discernible, area marked by depression (
|
||
<figureCitation id="133CDB987024FFB8FDF4F879C7EB4BB5" box="[561,647,1969,1992]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="1" pageNumber="390">Fig. 1C</figureCitation>
|
||
). Median tooth on posterior margin of epistome subtruncate (
|
||
<figureCitation id="133CDB987024FFB8FD55F819C78F4B95" box="[656,739,2001,2024]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="1" pageNumber="390">Fig. 1C</figureCitation>
|
||
). Ambulatory legs relatively long (
|
||
<figureCitation id="133CDB987024FFB8FE08F839C74C4475" box="[461,544,2033,2056]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="1" pageNumber="390">Fig. 1A</figureCitation>
|
||
). Major chela stout, with gap at base of closed fingers (
|
||
<figureCitation id="133CDB987024FFB8FE22F7D9C7504455" box="[487,572,2065,2088]" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="1" pageNumber="390">Fig. 1G</figureCitation>
|
||
). Male abdomen with somite 6 elongate, lateral margins gently concave (
|
||
<figureCitation id="133CDB987024FFB8FA45FF76C6234C88" captionStart="Fig" captionStartId="2.[136,171,2034,2055]" captionTargetBox="[142,1467,188,2010]" captionTargetId="figure-69@2.[142,1467,188,2010]" captionTargetPageId="2" captionText="Fig. 1. Thelphusula cristicervix (Ng & Grinang, 2004). Male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right chela; G, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.5353782" httpUri="https://zenodo.org/record/5353782/files/figure.png" pageId="1" pageNumber="390">Fig. 1D</figureCitation>
|
||
). G1 terminal segment subconical, tip sharply tapering, about 2 times length of subterminal segment (
|
||
<figureCitation id="133CDB987024FFB8FACAFF36C0CC4D68" box="[1295,1440,254,277]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="1" pageNumber="390">Fig. 3A, C–E</figureCitation>
|
||
). G2 with distal segment very short, tip sharp (
|
||
<figureCitation id="133CDB987024FFB8FADBFED6C01F4D48" box="[1310,1395,286,309]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="1" pageNumber="390">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8C71D7024FFB8FCE8FE96C6D64DA8" blockId="1.[813,1452,350,469]" pageId="1" pageNumber="390">
|
||
<emphasis id="B9731B0F7024FFB8FCE8FE96C6ED4D08" bold="true" box="[813,897,350,373]" pageId="1" pageNumber="390">Colour.</emphasis>
|
||
In life, the species is red to bright red overall all the dorsal surfaces and pereopods (adult male chelae usually more orangish-red), with the ventral surfaces yellowish-white (
|
||
<figureCitation id="133CDB987024FFB8FCF0FE76C6C04DA8" box="[821,940,446,469]" captionStart="Fig" captionStartId="5.[113,148,1245,1266]" captionTargetBox="[115,752,520,1221]" captionTargetId="figure-13@5.[113,752,518,1221]" captionTargetPageId="5" captionText="Fig. 4. Colours in life, Thelphusula cristicervix (Ng & Grinang, 2004). A, male (ca. 19.0 mm carapace width), in situ in forest, specimen not collected; B, female (19.3 × 14.2 mm) (ZRC) [Photograph by: Tan HH]. Both specimens from Sama Jaya Nature Reserve, Kuching Division, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5353788" httpUri="https://zenodo.org/record/5353788/files/figure.png" pageId="1" pageNumber="390">Fig. 4A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D94967024FFB8FCE8FE36C6A24AA8" pageId="1" pageNumber="390" type="discussion">
|
||
<paragraph id="8BB8C71D7024FFB8FCE8FE36C0C84EA8" blockId="1.[813,1452,510,725]" pageId="1" pageNumber="390">
|
||
<emphasis id="B9731B0F7024FFB8FCE8FE36C6F74E68" bold="true" box="[813,923,510,533]" pageId="1" pageNumber="390">Remarks.</emphasis>
|
||
The recent series of specimens of
|
||
<taxonomicName id="4C07BC9E7024FFB8FADDFE36C0C74E68" authorityName="Ng & Grinang" authorityYear="2004" box="[1304,1451,510,533]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7024FFB8FADDFE36C0C74E68" box="[1304,1451,510,533]" italics="true" pageId="1" pageNumber="390">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
from Sama Jaya Nature Reserve just outside Kuching city is about
|
||
<quantity id="4CFF6AF87024FFB8FC54FDF6C6B44E28" box="[913,984,574,597]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="390" unit="km" value="40.0">40 km</quantity>
|
||
from the type locality in Bau, but agree extremely well with the
|
||
<typeStatus id="54BC79BF7024FFB8FBF8FD96C1F64E08" box="[1085,1178,606,629]" pageId="1" pageNumber="390" type="holotype">holotype</typeStatus>
|
||
specimen in all aspects; including the structure of the carapace and proportions of the ambulatory legs. The good series, which includes small and large specimens, leaves little doubt to their conspecificity.
|
||
</paragraph>
|
||
<paragraph id="8BB8C71D7024FFB8FCE8FD36C0EB4848" blockId="1.[813,1452,766,1077]" pageId="1" pageNumber="390">
|
||
The morphology of the G1 of this species agrees well with what has been diagnosed and figured for
|
||
<taxonomicName id="4C07BC9E7024FFB8FB2EFCD6C6084F28" authority="Bott, 1969" authorityName="Bott" authorityYear="1969" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9731B0F7024FFB8FB2EFCD6C0034F48" box="[1259,1391,798,821]" italics="true" pageId="1" pageNumber="390">Thelphusula</emphasis>
|
||
<bibRefCitation id="EF96BAEC7024FFB8FAB2FCD6C6084F28" author="Bott R" pageId="1" pageNumber="390" pagination="359 - 366" refId="ref3387" refString="Bott R (1969) Flusskrabben aus Asien und ihre Klassifikation (Crustacea, Decapoda). Senckenbergiana Biologica, 50 (5 - 6): 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
, by
|
||
<bibRefCitation id="EF96BAEC7024FFB8FC49FCF6C1534F28" author="Tan SH & Ng PKL" box="[908,1087,830,853]" pageId="1" pageNumber="390" pagination="808 - 822" refId="ref4350" refString="Tan SH & Ng PKL (1998) Revision of the genus Thelphusula (Decapoda: Brachyura: Gecarcinucidae) with description of a new genus and three new species. Journal of Crustacean Biology, 18 (4): 808 - 822." type="journal article" year="1998">Tan & Ng (1998)</bibRefCitation>
|
||
, with the terminal segment slender, cylindrical and distinctly shorter than the subterminal segment (
|
||
<figureCitation id="133CDB987024FFB8FCF0FCB6C6C04FE8" box="[821,940,894,917]" captionStart="Fig" captionStartId="4.[136,171,2004,2025]" captionTargetBox="[136,1485,266,1950]" captionTargetId="figure-7@4.[108,1500,238,1953]" captionTargetPageId="4" captionText="Fig. 3. A–E, Thelphusula cristicervix (Ng & Grinang, 2004), male (19.4 × 14.8 mm) (ZRC), Sama Jaya Nature Reserve, Kuching Division, Sarawak; F–J, Thelphusula pueh, new species, male (16.2 × 12.1 mm) (ZRC), Kampung Sebako, Pueh, Lundu District, Sarawak. A, F, ventral view of left G1; B, G, ventral view of left G2; C, H, dorsal view of left G1; D, I, ventral view of terminal segment of left G1; E, J, dorsal view of terminal segment of left G1. Scale bars: A–C, F–H = 1.0 mm; D, E, I, J = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353786" httpUri="https://zenodo.org/record/5353786/files/figure.png" pageId="1" pageNumber="390">Fig. 3A, C</figureCitation>
|
||
). The carapace shape, as well as granulation of
|
||
<taxonomicName id="4C07BC9E7024FFB8FCE8FC56C6D24FC8" authorityName="Ng & Grinang" authorityYear="2004" box="[813,958,926,949]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7024FFB8FCE8FC56C6D24FC8" box="[813,958,926,949]" italics="true" pageId="1" pageNumber="390">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
, while relatively more ovate than other known species of
|
||
<taxonomicName id="4C07BC9E7024FFB8FC6DFC76C15C4FA8" authorityName="Bott" authorityYear="1969" box="[936,1072,958,981]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9731B0F7024FFB8FC6DFC76C15C4FA8" box="[936,1072,958,981]" italics="true" pageId="1" pageNumber="390">Thelphusula</emphasis>
|
||
</taxonomicName>
|
||
which have more quadrate ones, approaches the condition in more species like
|
||
<taxonomicName id="4C07BC9E7024FFB8FACBFC16C0C74F88" authorityName="Ng & Stuebing" authorityYear="1990" box="[1294,1451,990,1013]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="dicerophilus">
|
||
<emphasis id="B9731B0F7024FFB8FACBFC16C0C74F88" box="[1294,1451,990,1013]" italics="true" pageId="1" pageNumber="390">T. dicerophilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C07BC9E7024FFB8FC9DFC36C6A44868" authorityName="Holthuis" authorityYear="1979" box="[856,968,1022,1045]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="granosa">
|
||
<emphasis id="B9731B0F7024FFB8FC9DFC36C6A44868" box="[856,968,1022,1045]" italics="true" pageId="1" pageNumber="390">T. granosa</emphasis>
|
||
</taxonomicName>
|
||
which have relatively more inflated carapaces (cf.
|
||
<bibRefCitation id="EF96BAEC7024FFB8FC9CFBD6C6924848" author="Holthuis LB" box="[857,1022,1054,1077]" pageId="1" pageNumber="390" pagination="1 - 47" refId="ref3584" refString="Holthuis LB (1979) Cavernicolous and terrestrial decapod crustacean from northern Sarawak, Borneo. Zoologische Verhandelingen, 171: 1 - 47, pls. 1 - 8." type="journal article" year="1979">Holthuis, 1979</bibRefCitation>
|
||
: pl. 5;
|
||
<bibRefCitation id="EF96BAEC7024FFB8FB89FBD6C0554848" author="Ng PKL & Stuebing R" box="[1100,1337,1054,1077]" pageId="1" pageNumber="390" pagination="45 - 51" refId="ref4153" refString="Ng PKL & Stuebing R (1990) Thelphusula dicerophilus sp. nov., a new species of freshwater crab (Crustacea: Decapoda: Brachyura: Gecarcinucidae) found in mud wallows of the Sumatran Rhinoceros from Sabah, Borneo. IndoMalayan Zoology, (1989) 6 (1): 45 - 51, pl. 1." type="journal article" year="1990">Ng & Stuebing, 1990</bibRefCitation>
|
||
: pl. 1).
|
||
</paragraph>
|
||
<paragraph id="8BB8C71D7024FFB8FCE8FB96C1834908" blockId="1.[813,1452,1118,1397]" pageId="1" pageNumber="390">
|
||
In contrast to the moist karst forest where the
|
||
<typeStatus id="54BC79BF7024FFB8FAEBFB96C0E74808" box="[1326,1419,1118,1141]" pageId="1" pageNumber="390" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C07BC9E7024FFB8FCE8FBB6C6AD48E8" authorityName="Ng & Grinang" authorityYear="2004" box="[813,961,1150,1173]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7024FFB8FCE8FBB6C6AD48E8" box="[813,961,1150,1173]" italics="true" pageId="1" pageNumber="390">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
was collected, the recent specimens were all collected from a swampy part of a well-shaded kerangas forest. The soil is wet and loamy, usually with a dense root system permeating it. The crabs dig relatively deep burrows, up to perhaps a metre in depth in places and come out only at night. The presence of the species in both limestone and kerangas forests indicates that its distribution is delimited by soil conditions rather than forest type.
|
||
</paragraph>
|
||
<paragraph id="8BB8C71D7024FFB8FCE8FA56C6A24AA8" blockId="1.[813,1452,1438,1749]" pageId="1" pageNumber="390">
|
||
<bibRefCitation id="EF96BAEC7024FFB8FCE8FA56C16F49C8" author="Ng PKL & Yeo DCJ" box="[813,1027,1438,1461]" pageId="1" pageNumber="390" pagination="95 - 120" refId="ref4256" refString="Ng PKL & Yeo DCJ (2007) Malaysian freshwater crabs: conservation prospects and challenges. In: Chua L (ed.) Proceedings of the Seminar on the Status of Biological Diversity in Malaysia and Threat Assessment of Plant Species in Malaysia, 28 - 30 June 2005. Forest Research Institute Malaysia, Kepong. Pp. 95 - 120." type="book chapter" year="2007">Ng & Yeo (2007: 9)</bibRefCitation>
|
||
noted that the species was endangered, commenting that “the Bau area [
|
||
<typeStatus id="54BC79BF7024FFB8FB46FA76C1D849A8" box="[1155,1204,1470,1493]" pageId="1" pageNumber="390">type</typeStatus>
|
||
locality] does not have long-term protection and the species occurs in low-lying areas, it is very vulnerable to anthropogenic and associated effects.” (see also
|
||
<bibRefCitation id="EF96BAEC7024FFB8FC3CF9D6C0634A48" author="Cumberlidge N & Ng PKL & Yeo DCJ & Magalhaes C & Campos MR & Alvarez F & Naruse T & Daniels SR & Esser LJ & Attipoe FYK & Clotilde-Ba F-L & Darwall W & Mcivor A & Collen B & Ram M" box="[1017,1295,1566,1589]" pageId="1" pageNumber="390" pagination="1665 - 1673" refId="ref3511" refString="Cumberlidge N, Ng PKL, Yeo DCJ, Magalhaes C, Campos MR, Alvarez F, Naruse T, Daniels SR, Esser LJ, Attipoe FYK, Clotilde-Ba F-L, Darwall W, Mcivor A, Collen B & Ram M (2009) Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation, 142: 1665 - 1673." type="journal article" year="2009">Cumberlidge et al., 2009</bibRefCitation>
|
||
: Appendix 1). The discovery of
|
||
<taxonomicName id="4C07BC9E7024FFB8FC3DF9F6C1FF4A28" authorityName="Ng & Grinang" authorityYear="2004" box="[1016,1171,1598,1621]" class="Malacostraca" family="Gecarcinucidae" genus="Coccusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B9731B0F7024FFB8FC3DF9F6C1FF4A28" box="[1016,1171,1598,1621]" italics="true" pageId="1" pageNumber="390">T. cristicervix</emphasis>
|
||
</taxonomicName>
|
||
in a nature reserve near Kuching city does not change its status as the protected area is open to the public and the species is known mainly from one small patch of swamp only several hundred square metres in area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |