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<document id="7F3C7153DF30C310E811A8F06BFFC38A" ID-CLB-Dataset="24000" ID-DOI="10.11646/zootaxa.4735.1.1" ID-GBIF-Dataset="ffaf3b73-8bfe-499d-9073-0ce8c104e078" ID-ISSN="1175-5326" ID-Zenodo-Dep="3670762" ID-ZooBank="BD52DF91-3A7E-46FB-8975-38A67BFBBD61" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581952963911" checkinUser="plazi" docAuthor="Lonsdale, Owen" docDate="2020" docId="BD15296C6A6FFF93FF1AFA50DA1EA662" docLanguage="en" docName="zootaxa.4735.1.1.pdf.imf" docOrigin="Zootaxa 4735 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Gobryidae McAlpine 1997" docType="treatment" docVersion="10" lastPageNumber="22" masterDocId="412C51146A7CFF86FF8DFFD5D855A559" masterDocTitle="Family groups of Diopsoidea and Nerioidea (Diptera: Schizophora) - Definition, history and relationships" masterLastPageNumber="177" masterPageNumber="1" pageNumber="20" updateTime="1698783921049" updateUser="ExternalLinkService">
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<mods:title id="0997E14569AD7A25BCFCA2D969A5498E">Family groups of Diopsoidea and Nerioidea (Diptera: Schizophora) - Definition, history and relationships</mods:title>
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<mods:namePart id="1B02CA49E149EC3179F5EB1812DD79FE">Lonsdale, Owen</mods:namePart>
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<treatment id="BD15296C6A6FFF93FF1AFA50DA1EA662" ID-DOI="http://doi.org/10.5281/zenodo.3679580" ID-GBIF-Taxon="162147168" ID-Zenodo-Dep="3679580" LSID="urn:lsid:plazi:treatment:BD15296C6A6FFF93FF1AFA50DA1EA662" httpUri="http://treatment.plazi.org/id/BD15296C6A6FFF93FF1AFA50DA1EA662" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="7DA6CBF16A6FFF95FF1AFA50D9BFA0C6" box="[151,490,1413,1439]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="3503987A6A6FFF95FF1AFA50D9BFA0C6" blockId="19.[151,490,1413,1475]" box="[151,490,1413,1439]" pageId="19" pageNumber="20">
<heading id="6E4B2F166A6FFF95FF1AFA50D9BFA0C6" bold="true" box="[151,490,1413,1439]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="F2BCE3F96A6FFF95FF1AFA50D9BFA0C6" ID-CoL="623LJ" authority="McAlpine, 1997" authorityName="McAlpine" authorityYear="1997" box="[151,490,1413,1439]" class="Insecta" family="Gobryidae" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">
<emphasis id="07C844686A6FFF95FF1AFA50D9BFA0C6" bold="true" box="[151,490,1413,1439]" pageId="19" pageNumber="20">Gobryidae McAlpine, 1997</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7DA6CBF16A6FFF95FF1AFA7CD9E6A09A" box="[151,435,1449,1475]" pageId="19" pageNumber="20" type="description">
<paragraph id="3503987A6A6FFF95FF1AFA7CD9E6A09A" blockId="19.[151,490,1413,1475]" box="[151,435,1449,1475]" pageId="19" pageNumber="20">
(
<figureCitation id="AD8784FF6A6FFF95FF12FA7CD975A09A" box="[159,288,1449,1475]" captionStart="FIGURES 62–70" captionStartId="101.[151,264,1796,1821]" captionTargetBox="[158,1423,179,1765]" captionTargetId="figure@101.[151,1436,174,1777]" captionTargetPageId="101" captionText="FIGURES 62–70. Figs 62–64: Nothybus biguttatus Wulp (Nothybidae), female abdomen; 62: segment 7 to cercus, ventral; 63: same, dorsal; 64: internal genitalia. Figs 65–66: Somatia aestiva (Fabricius), female abdomen; 65: segment 7 to cerci; 66: internal genitalia. Figs 67–70: Gobrya sp., female abdomen; 67: segment 6 to cerci, dorsal; 68: same, right lateral; 69: same, ventral; 70: spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3670780" httpUri="https://zenodo.org/record/3670780/files/figure.png" pageId="19" pageNumber="20">Figs 67–70</figureCitation>
,
<figureCitation id="AD8784FF6A6FFF95FEA1FA7CD920A09A" box="[300,373,1449,1475]" captionStart-0="FIGURES 82–88" captionStart-1="FIGURES 89–97" captionStartId-0="104.[151,264,1799,1824]" captionStartId-1="105.[151,264,1799,1824]" captionTargetBox-0="[151,1436,174,1780]" captionTargetBox-1="[160,1428,182,1769]" captionTargetId-0="figure@104.[151,1436,174,1780]" captionTargetId-1="figure@105.[151,1436,174,1780]" captionTargetPageId-0="104" captionTargetPageId-1="105" captionText-0="FIGURES 82–88. Gobrya sp. (Gobryidae); 82: female, dorsal; 83: same, lateral; 84: male, ventral; 85: female head, anterior; 86: same, dorsal; 87: male, Seram, Indonesia; 88: male, Sarawak, Malaysia." captionText-1="FIGURES 89–97. Gobrya sp. (Gobryidae), male abdomen; 89: right lateral; 90: ventral (right surstylus shaded); 91: terminalia, left lateral; 92: external genitalia, anterior (right surstylus and subepandrial sclerite shaded); 93: enlarged detail of subepandrial sclerite; 94: ejaculatory apodeme; 95: internal genitalia, right lateral; 96: same, ventral; 97: same, left lateral." figureDoi-0="http://doi.org/10.5281/zenodo.3670786" figureDoi-1="http://doi.org/10.5281/zenodo.3670788" httpUri-0="https://zenodo.org/record/3670786/files/figure.png" httpUri-1="https://zenodo.org/record/3670788/files/figure.png" pageId="19" pageNumber="20">82–97</figureCitation>
,
<figureCitation id="AD8784FF6A6FFF95FE0DFA7CD9F9A09A" box="[384,428,1449,1475]" captionStart="FIGURES 395–402" captionStartId="146.[151,264,1508,1533]" captionTargetBox="[151,1436,174,1479]" captionTargetId="figure@146.[151,1436,174,1489]" captionTargetPageId="146" captionText="FIGURES 395–402. Wings; 395: Belobackenbardia cornicula Shatalkin (Psilidae), female paratype; 396: Chyliza notata Loew (Psilidae); 397: Psila hennigi (Thompson &amp; Pont) (Psilidae); 398: Loxocera cylindrica Say (Psilidae); 399: Nothybus longicollis (Walker) (Nothybidae); 400: N. biguttatus Wulp; 401: Somatia aestiva (Fabricius) (Somatiidae); 402: Gobrya sp. (Gobryidae)." figureDoi="http://doi.org/10.5281/zenodo.3670874" httpUri="https://zenodo.org/record/3670874/files/figure.png" pageId="19" pageNumber="20">402</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7DA6CBF16A6FFF95FF1AFA38D9D3A37D" pageId="19" pageNumber="20" type="type_taxon">
<paragraph id="3503987A6A6FFF95FF1AFA38D9D3A37D" blockId="19.[151,1437,1517,1572]" pageId="19" pageNumber="20">
<emphasis id="07C844686A6FFF95FF1AFA38D94CA35C" bold="true" box="[151,281,1517,1541]" pageId="19" pageNumber="20">
<typeStatus id="EA0726D86A6FFF95FF1AFA38D899A35C" box="[151,204,1517,1541]" pageId="19" pageNumber="20">Type</typeStatus>
genus:
</emphasis>
<taxonomicName id="F2BCE3F96A6FFF95FE92FA38DA7CA35F" ID-CoL="4PQP" authority="Walker 1860: 166" authorityName="Walker" authorityPageNumber="166" authorityYear="1860" box="[287,553,1517,1542]" class="Insecta" family="Gobryidae" genus="Gobrya" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="07C844686A6FFF95FE92FA38D939A35C" box="[287,364,1517,1541]" italics="true" pageId="19" pageNumber="20">Gobrya</emphasis>
<bibRefCitation id="512DE58B6A6FFF95FEFFFA38DA7CA35F" author="Walker, F." box="[370,553,1517,1542]" pageId="19" pageNumber="20" pagination="144 - 168" refId="ref98710" refString="Walker, F. (1860) Catalogue of the dipterous insects collected at Makessar, in Celebes, by Mr. A. R. Wallace, with descriptions of new species. Journal and Proceedings of the Linnean Society of London. Zoology, 5 (Supplement), 144 - 168. https: // doi. org / 10.1111 / j. 1096 - 3642.1860. tb 01023. x" type="journal article" year="1860">Walker 1860: 166</bibRefCitation>
</taxonomicName>
, by
<bibRefCitation id="512DE58B6A6FFF95FDDEFA38DB10A35C" author="McAlpine, D. K." box="[595,837,1517,1542]" pageId="19" pageNumber="20" pagination="465 - 475" refId="ref88714" refString="McAlpine, D. K. (1997 a) Relationships of the Megamerinidae. Beitrage zur Entomologie, 47 (2), 465 - 475." type="journal article" year="1997">McAlpine (1997a: 182)</bibRefCitation>
.
<emphasis id="07C844686A6FFF95FCC2FA38DC68A35C" bold="true" box="[847,1085,1517,1542]" pageId="19" pageNumber="20">
<typeStatus id="EA0726D86A6FFF95FCC2FA38DBD1A35C" box="[847,900,1517,1541]" pageId="19" pageNumber="20">Type</typeStatus>
species of genus:
</emphasis>
<taxonomicName id="F2BCE3F96A6FFF95FBCEFA38D8BAA37D" ID-CoL="6KP42" authority="Walker, 1860: 167" authorityName="Walker" authorityPageNumber="167" authorityYear="1860" class="Insecta" family="Gobryidae" genus="Gobrya" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bacchoides">
<emphasis id="07C844686A6FFF95FBCEFA38DD5DA35C" box="[1091,1288,1517,1541]" italics="true" pageId="19" pageNumber="20">Gobrya bacchoides</emphasis>
<bibRefCitation id="512DE58B6A6FFF95FA83FA38D8BAA37D" author="Walker, F." pageId="19" pageNumber="20" pagination="144 - 168" refId="ref98710" refString="Walker, F. (1860) Catalogue of the dipterous insects collected at Makessar, in Celebes, by Mr. A. R. Wallace, with descriptions of new species. Journal and Proceedings of the Linnean Society of London. Zoology, 5 (Supplement), 144 - 168. https: // doi. org / 10.1111 / j. 1096 - 3642.1860. tb 01023. x" type="journal article" year="1860">Walker, 1860: 167</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="7DA6CBF16A6FFF95FF1AF987DC85A279" pageId="19" pageNumber="20" type="discussion">
<paragraph id="3503987A6A6FFF95FF1AF987DC85A279" blockId="19.[151,1437,1618,2041]" pageId="19" pageNumber="20">
The family
<taxonomicName id="F2BCE3F96A6FFF95FE93F987D9C1A335" authorityName="McAlpine" authorityYear="1997" box="[286,404,1618,1644]" class="Insecta" family="Gobryidae" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Gobryidae</taxonomicName>
is one of the most recently described families of flies. They were termed “hinge flies” by
<bibRefCitation id="512DE58B6A6FFF95FF1AF9A3D93DA3C8" author="McAlpine, D. K." box="[151,360,1654,1681]" pageId="19" pageNumber="20" pagination="167 - 194" refId="ref88742" refString="McAlpine, D. K. (1997 b) Gobryidae, a new family of acalyptrate flies (Diptera: Diopsoidea), and a discussion of relationships of the diopsoid families. Records of the Australian Museum, 49, 167 - 194. https: // doi. org / 10.3853 / j. 0067 - 1975.49.1997.1264" type="journal article" year="1997">McAlpine (1997b)</bibRefCitation>
because of the characteristic articulation of the narrow, straight abdomen with the thorax, which is also seen to a lesser degree in the
<taxonomicName id="F2BCE3F96A6FFF95FDAEF94FDA83A3ED" authorityName="Hendel" authorityYear="1913" box="[547,726,1690,1716]" class="Insecta" family="Megamerinidae" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Megamerinidae</taxonomicName>
, the family in which there were once placed.
<taxonomicName id="F2BCE3F96A6FFF95FB4AF94FDD68A3ED" authorityName="McAlpine" authorityYear="1997" box="[1223,1341,1690,1716]" class="Insecta" family="Gobryidae" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Gobryidae</taxonomicName>
consists of at least eight species in the single genus
<taxonomicName id="F2BCE3F96A6FFF95FD09F96BDBEBA380" authority="(McAlpine, 1997 b)" baseAuthorityName="McAlpine" baseAuthorityYear="1997" box="[644,958,1726,1753]" class="Insecta" family="Gobryidae" genus="Gobrya" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="07C844686A6FFF95FD09F96BDA8FA381" box="[644,730,1726,1752]" italics="true" pageId="19" pageNumber="20">Gobrya</emphasis>
(
<bibRefCitation id="512DE58B6A6FFF95FD64F96BDBE3A380" author="McAlpine, D. K." box="[745,950,1726,1753]" pageId="19" pageNumber="20" pagination="167 - 194" refId="ref88742" refString="McAlpine, D. K. (1997 b) Gobryidae, a new family of acalyptrate flies (Diptera: Diopsoidea), and a discussion of relationships of the diopsoid families. Records of the Australian Museum, 49, 167 - 194. https: // doi. org / 10.3853 / j. 0067 - 1975.49.1997.1264" type="journal article" year="1997">McAlpine, 1997b</bibRefCitation>
)
</taxonomicName>
, although only five are described. Species occur in the
<collectingRegion id="F77856986A6FFF95FEAFF937D983A3A5" box="[290,470,1762,1788]" country="Democratic Republic of the Congo" name="Orientale" pageId="19" pageNumber="20">Oriental Region</collectingRegion>
and adjoining parts of the
<collectingRegion id="F77856986A6FFF95FC8DF937DB85A3A5" box="[768,976,1762,1788]" inferredBySuffix="true" pageId="19" pageNumber="20">Australian Region</collectingRegion>
. There is no key to species. Species were catalogued as
<taxonomicName id="F2BCE3F96A6FFF95FEBBF8D3DA93A279" authority="in Steyskal (1977 e)" authorityName="in Steyskal" authorityYear="1977" box="[310,710,1798,1824]" class="Insecta" family="Megamerinidae" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">
Megamerinidae in
<bibRefCitation id="512DE58B6A6FFF95FD84F8D3DA93A279" author="Steyskal, G. C." box="[521,710,1798,1824]" pageId="19" pageNumber="20" pagination="28 - 29" refId="ref97067" refString="Steyskal, G. C. (1977 e) Family Megamerinidae. In: Delfinado, M. C. &amp; Hardy, D. E. (Eds.), A Catalog of the Diptera of the Oriental Region 3: Suborder Cyclorrhapha (Excluding Division Aschiza). University Press of Hawaii, Honolulu, pp. 28 - 29." type="book chapter" year="1977">Steyskal (1977e)</bibRefCitation>
</taxonomicName>
[
<collectingRegion id="F77856986A6FFF95FD59F8D0DB63A279" box="[724,822,1797,1824]" country="Democratic Republic of the Congo" name="Orientale" pageId="19" pageNumber="20">Oriental</collectingRegion>
] and
<bibRefCitation id="512DE58B6A6FFF95FCFFF8D3DC6FA278" author="Evenhuis, N. L." box="[882,1082,1798,1825]" pageId="19" pageNumber="20" pagination="471" refId="ref79992" refString="Evenhuis, N. L. (1989 b) Family Megamerinidae. In: Evenhuis, N. L. (Ed.), Catalog of the Diptera of the Australasian and Oceanian Regions. Bishop Museum Press and E. J. Brill, Honolulu, pp. 471." type="book chapter" year="1989">Evenhuis (1989b)</bibRefCitation>
[Australian].
</paragraph>
</subSubSection>
<subSubSection id="7DA6CBF16A6FFF95FF4AF8FCDA46A2D5" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="3503987A6A6FFF95FF4AF8FCDB52A230" blockId="19.[151,1437,1618,2041]" pageId="19" pageNumber="20">
<emphasis id="07C844686A6FFF95FF4AF8FCD972A21D" bold="true" box="[199,295,1833,1860]" pageId="19" pageNumber="20">Biology.</emphasis>
The biology of
<taxonomicName id="F2BCE3F96A6FFF95FE6FF8FFDA6DA21D" baseAuthorityName="McAlpine" baseAuthorityYear="1997" box="[482,568,1834,1860]" class="Insecta" family="Gobryidae" genus="Gobrya" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="07C844686A6FFF95FE6FF8FFDA6DA21D" box="[482,568,1834,1860]" italics="true" pageId="19" pageNumber="20">Gobrya</emphasis>
</taxonomicName>
is unknown, although specimens have been collected on foliage or near the margin of lowland rainforests (D.K.
<bibRefCitation id="512DE58B6A6FFF95FDBDF89BDAAFA230" author="McAlpine, D. K." box="[560,762,1870,1897]" pageId="19" pageNumber="20" pagination="167 - 194" refId="ref88742" refString="McAlpine, D. K. (1997 b) Gobryidae, a new family of acalyptrate flies (Diptera: Diopsoidea), and a discussion of relationships of the diopsoid families. Records of the Australian Museum, 49, 167 - 194. https: // doi. org / 10.3853 / j. 0067 - 1975.49.1997.1264" type="journal article" year="1997">McAlpine, 1997b</bibRefCitation>
).
</paragraph>
<paragraph id="3503987A6A6FFF95FF4AF8A7DA46A2D5" blockId="19.[151,1437,1618,2041]" box="[199,531,1906,1932]" pageId="19" pageNumber="20">
<emphasis id="07C844686A6FFF95FF4AF8A7D9C3A2D5" bold="true" box="[199,406,1906,1932]" pageId="19" pageNumber="20">Immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="7DA6CBF16A6FFF93FF4AF840DA1EA662" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="3503987A6A6FFF92FF4AF840DC83A4F6" blockId="19.[151,1437,1618,2041]" lastBlockId="20.[151,1437,153,2015]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="07C844686A6FFF95FF4AF840D9D8A2E9" bold="true" box="[199,397,1941,1968]" pageId="19" pageNumber="20">Adult Diagnosis.</emphasis>
Colour dark brown (including halter knob), slightly iridescent, with antenna, mouthparts and parts of legs white to yellow; body surface mostly glossy (
<figureCitation id="AD8784FF6A6FFF95FCADF86FDBF6A28D" box="[800,931,1978,2004]" captionStart="FIGURES 82–88" captionStartId="104.[151,264,1799,1824]" captionTargetBox="[151,1436,174,1780]" captionTargetId="figure@104.[151,1436,174,1780]" captionTargetPageId="104" captionText="FIGURES 82–88. Gobrya sp. (Gobryidae); 82: female, dorsal; 83: same, lateral; 84: male, ventral; 85: female head, anterior; 86: same, dorsal; 87: male, Seram, Indonesia; 88: male, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3670786" httpUri="https://zenodo.org/record/3670786/files/figure.png" pageId="19" pageNumber="20">Figs 82–88</figureCitation>
). Body length 3.5–8.0mm. Antenna elbowed, first flagellomere elongate; pedicel with dorsal suture; arista sparsely long plumose, with rays becoming shorter apically and sparser ventrally to base. Head short and broad, nearly twice width of thorax; face convex and well sclerotized with mouthparts ventral and receded; frons and dorsum of ptilinal suture narrow (lateral arms nearly absent). Cephalic chaetotaxy reduced to two verticals, with vibrissa, fronto-orbitals, postocellar and ocellar setae absent. Thoracic chaetotaxy reduced to notopleural (one), two supra-alars, and apical scutellar (on slight tubercle). Broad precoxal and postmetacoxal bridges present, although postmetacoxal bridge sometimes membranous medially. Hind femur long and stout with two ventral rows of spines. Abdomen petiolate, straight and narrow with apex slightly widened. Wing with anal lobe and alula strongly reduced; costa unbroken; subcosta complete; R
<subScript id="A9389A3F6A68FF92FAA0FE57DD12A4C9" attach="left" box="[1325,1351,386,400]" fontSize="6" pageId="20" pageNumber="21">4+5</subScript>
and M
<subScript id="A9389A3F6A68FF92FA19FE57DDC9A4C9" attach="left" box="[1428,1436,386,400]" fontSize="6" pageId="20" pageNumber="21">1</subScript>
slightly convergent (
<figureCitation id="AD8784FF6A68FF92FEF0FE40D9B4A4F6" box="[381,481,405,431]" captionStart="FIGURES 395–402" captionStartId="146.[151,264,1508,1533]" captionTargetBox="[151,1436,174,1479]" captionTargetId="figure@146.[151,1436,174,1489]" captionTargetPageId="146" captionText="FIGURES 395–402. Wings; 395: Belobackenbardia cornicula Shatalkin (Psilidae), female paratype; 396: Chyliza notata Loew (Psilidae); 397: Psila hennigi (Thompson &amp; Pont) (Psilidae); 398: Loxocera cylindrica Say (Psilidae); 399: Nothybus longicollis (Walker) (Nothybidae); 400: N. biguttatus Wulp; 401: Somatia aestiva (Fabricius) (Somatiidae); 402: Gobrya sp. (Gobryidae)." figureDoi="http://doi.org/10.5281/zenodo.3670874" httpUri="https://zenodo.org/record/3670874/files/figure.png" pageId="20" pageNumber="21">Fig. 402</figureCitation>
). Male external terminalia globular and asymmetrical (
<figureCitation id="AD8784FF6A68FF92FBCAFE40DC9FA4F6" box="[1095,1226,405,431]" captionStart="FIGURES 89–97" captionStartId="105.[151,264,1799,1824]" captionTargetBox="[160,1428,182,1769]" captionTargetId="figure@105.[151,1436,174,1780]" captionTargetPageId="105" captionText="FIGURES 89–97. Gobrya sp. (Gobryidae), male abdomen; 89: right lateral; 90: ventral (right surstylus shaded); 91: terminalia, left lateral; 92: external genitalia, anterior (right surstylus and subepandrial sclerite shaded); 93: enlarged detail of subepandrial sclerite; 94: ejaculatory apodeme; 95: internal genitalia, right lateral; 96: same, ventral; 97: same, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3670788" httpUri="https://zenodo.org/record/3670788/files/figure.png" pageId="20" pageNumber="21">Figs 89–97</figureCitation>
).
</paragraph>
<paragraph id="3503987A6A68FF92FF4AFE6DD93EA742" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="21">
<emphasis id="07C844686A68FF92FF4AFE6DD9D8A48A" bold="true" box="[199,397,440,467]" pageId="20" pageNumber="21">Adult Definition</emphasis>
. Colour dark brown (including halter knob and sometimes base), slightly bluish iridescent on darkly pigmented sections, and with antenna, mouthparts and parts of legs white to yellow (
<figureCitation id="AD8784FF6A68FF92FB45FE08DD19A4AE" box="[1224,1356,477,503]" captionStart="FIGURES 82–88" captionStartId="104.[151,264,1799,1824]" captionTargetBox="[151,1436,174,1780]" captionTargetId="figure@104.[151,1436,174,1780]" captionTargetPageId="104" captionText="FIGURES 82–88. Gobrya sp. (Gobryidae); 82: female, dorsal; 83: same, lateral; 84: male, ventral; 85: female head, anterior; 86: same, dorsal; 87: male, Seram, Indonesia; 88: male, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3670786" httpUri="https://zenodo.org/record/3670786/files/figure.png" pageId="20" pageNumber="21">Figs 82–88</figureCitation>
). Body length 3.5–8.0mm.
</paragraph>
<paragraph id="3503987A6A68FF92FF4AFDF0DB8BA6DA" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="21">
<emphasis id="07C844686A68FF92FF4AFDF0D912A766" box="[199,327,549,575]" italics="true" pageId="20" pageNumber="21">Chaetotaxy</emphasis>
: 1 inner vertical; 1 outer vertical; 0 fronto-orbital; 0 ocellar; 0 postocellar; vibrissa absent. Face and gena irregularly and sparsely setulose. Postoccipital setae present dorsolaterally. 0 presutural intra-alar; 0 postpronotal; 0–1 anterior notopleural, 1 posterior notopleural; 0–1 posterior supra-alar; 0 posterior intra-alar; 0 dorsocentrals; 0 acrostichals; 1 apical scutellar (on small tubercle); 0 proepisternal; 0 anepisternal; 0 katepisternal (sometimes with dense setulae ventrally). Body mostly glossy and bare with very short microtomentum present on head, notum, limited portions of abdomen, and pre- and postmetacoxal bridges; setulose on prosternum (posteriorly and laterally), posterior margin of anepisternum, behind posterior thoracic spiracle, katepisternum, notum and gena; parafacial and lateral margin of frons with single row of erect setulae; pedicel with several ventromedial and one dorsal seta. Mid tibia with ventroapical seta. Hind tibia sometimes with one or two distinct anterodistal setae. Hind basitarsomere with longer basal hairs and bare elongate anterobasal recess.
</paragraph>
<paragraph id="3503987A6A68FF92FF4AFC58D928A06A" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="21">
<emphasis id="07C844686A68FF92FF4AFC58D951A6FE" box="[199,260,909,935]" italics="true" pageId="20" pageNumber="21">Head</emphasis>
. Antenna elbowed, bases approximate; pedicel rounded, with wide, deep seam to dorsum, dividing small- er outer surface from broader, convex inner surface; first flagellomere elongate, narrower than pedicel; arista unsegmented and sparsely long plumose, with rays becoming shorter apically and sparser ventrally to base (
<figureCitation id="AD8784FF6A68FF92FAB6FC00DDDAA6B6" box="[1339,1423,980,1007]" captionStart="FIGURES 82–88" captionStartId="104.[151,264,1799,1824]" captionTargetBox="[151,1436,174,1780]" captionTargetId="figure@104.[151,1436,174,1780]" captionTargetPageId="104" captionText="FIGURES 82–88. Gobrya sp. (Gobryidae); 82: female, dorsal; 83: same, lateral; 84: male, ventral; 85: female head, anterior; 86: same, dorsal; 87: male, Seram, Indonesia; 88: male, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3670786" httpUri="https://zenodo.org/record/3670786/files/figure.png" pageId="20" pageNumber="21">Fig. 85</figureCitation>
). Head short and broad, nearly twice width of thorax; frons narrow (slightly more than half width of eye), smooth, flat, horizontal and slightly constricted posteriorly, with ocellar triangle not evident; orbital plate not delimited, but evident as short, pale posteromedial region that tapers at both ends (
<figureCitation id="AD8784FF6A68FF92FC2CFB94DBA0A105" box="[929,1013,1089,1116]" captionStart="FIGURES 82–88" captionStartId="104.[151,264,1799,1824]" captionTargetBox="[151,1436,174,1780]" captionTargetId="figure@104.[151,1436,174,1780]" captionTargetPageId="104" captionText="FIGURES 82–88. Gobrya sp. (Gobryidae); 82: female, dorsal; 83: same, lateral; 84: male, ventral; 85: female head, anterior; 86: same, dorsal; 87: male, Seram, Indonesia; 88: male, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3670786" httpUri="https://zenodo.org/record/3670786/files/figure.png" pageId="20" pageNumber="21">Fig. 86</figureCitation>
); ocelli at vertex; ptilinal suture very short, without long lateral arms. Back of head concave dorsomedially and with setulose bulge above foramen. Lunule absent. Face strongly convex ventrally (similar to
<taxonomicName id="F2BCE3F96A68FF92FD76FB5CDB01A1FA" authorityName="Macquart" authorityYear="1835" box="[763,852,1161,1187]" class="Insecta" family="Psilidae" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Psilidae</taxonomicName>
but broadly rounded) and fused to parafacial (suture sometimes evident in part); face and parafacial sometimes strongly swollen laterally, forming medial groove; gena broad, not distinct from face, and both it and venter of postgena mostly sunken beneath head and facing ventrally; buccal cavity ovate, relatively small, sharply delimited; clypeus longer than wide, rounded with apex pointed; mouthparts reduced.
</paragraph>
<paragraph id="3503987A6A68FF92FF4AFAE8DB12A32E" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="21">
<emphasis id="07C844686A68FF92FF4AFAE8D94DA00E" box="[199,280,1341,1367]" italics="true" pageId="20" pageNumber="21">Thorax</emphasis>
. Thorax elongate and narrow, without pronotal “neck”, but with shallow medial and lateral carinae displacing cervical sclerites. Precoxal bridge present; bulging laterally, with medial groove, posterior surface raised (
<figureCitation id="AD8784FF6A68FF92FF2DFA50D8A6A0C6" box="[160,243,1413,1439]" captionStart="FIGURES 82–88" captionStartId="104.[151,264,1799,1824]" captionTargetBox="[151,1436,174,1780]" captionTargetId="figure@104.[151,1436,174,1780]" captionTargetPageId="104" captionText="FIGURES 82–88. Gobrya sp. (Gobryidae); 82: female, dorsal; 83: same, lateral; 84: male, ventral; 85: female head, anterior; 86: same, dorsal; 87: male, Seram, Indonesia; 88: male, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3670786" httpUri="https://zenodo.org/record/3670786/files/figure.png" pageId="20" pageNumber="21">Fig. 84</figureCitation>
); prosternum velvety, relatively broad, setose laterally; postmetacoxal bridge high if present. Proepisternum broad, bulging anterodorsally, sometimes fused to postpronotum. Suture between katepisternum and anepisternum only present for a short distance posteriorly, partially evident anteriorly as darker line. Suture between katatergite and anepimeron sunken; posteromedial margin of mediotergite extended as lobe that meets longer band-like anteromedial extension of T1. Greater ampulla present. Anterior thoracic spiracle laterally compressed, ventrally with groove terminating in small, deep pit (similar to some
<taxonomicName id="F2BCE3F96A68FF92FC84F9ECDB32A30D" authorityName="Macquart" authorityYear="1835" box="[777,871,1593,1620]" class="Insecta" family="Psilidae" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Psilidae</taxonomicName>
); tubercle present below posterior spiracle. Broad membranous space separating anepisternum and anepimeron.
</paragraph>
<paragraph id="3503987A6A68FF92FF4AF957DD32A25E" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="21">
<emphasis id="07C844686A68FF92FF4AF957D954A3C2" box="[199,257,1666,1691]" italics="true" pageId="20" pageNumber="21">Wing</emphasis>
. (
<figureCitation id="AD8784FF6A68FF92FE95F954D92CA3C2" box="[280,377,1665,1691]" captionStart="FIGURES 395–402" captionStartId="146.[151,264,1508,1533]" captionTargetBox="[151,1436,174,1479]" captionTargetId="figure@146.[151,1436,174,1489]" captionTargetPageId="146" captionText="FIGURES 395–402. Wings; 395: Belobackenbardia cornicula Shatalkin (Psilidae), female paratype; 396: Chyliza notata Loew (Psilidae); 397: Psila hennigi (Thompson &amp; Pont) (Psilidae); 398: Loxocera cylindrica Say (Psilidae); 399: Nothybus longicollis (Walker) (Nothybidae); 400: N. biguttatus Wulp; 401: Somatia aestiva (Fabricius) (Somatiidae); 402: Gobrya sp. (Gobryidae)." figureDoi="http://doi.org/10.5281/zenodo.3670874" httpUri="https://zenodo.org/record/3670874/files/figure.png" pageId="20" pageNumber="21">Fig. 402</figureCitation>
) Wing slender; anal lobe and alula strongly reduced; apically infuscated. Calypter linear, with several long hairs. Cells bm and
<emphasis id="07C844686A68FF92FD80F973DA72A3E6" box="[525,551,1702,1727]" italics="true" pageId="20" pageNumber="21">cu</emphasis>
a (slightly shorter) ending approximately at apex of Sc; cells
<emphasis id="07C844686A68FF92FB6CF973DCAEA3E6" box="[1249,1275,1702,1727]" italics="true" pageId="20" pageNumber="21">cu</emphasis>
a and bm, and adjoining wing margin, without microsetulae. Cell dm narrow, slightly wider apically.
<collectingRegion id="F77856986A68FF92FBF5F91CDCEDA3BA" box="[1144,1208,1737,1763]" country="Tanzania" name="Pwani" pageId="20" pageNumber="21">Costa</collectingRegion>
unbroken; subcosta complete; R
<subScript id="A9389A3F6A68FF92FE92F92BD96CA255" attach="left" box="[287,313,1790,1804]" fontSize="6" pageId="20" pageNumber="21">4+5</subScript>
and M
<subScript id="A9389A3F6A68FF92FE08F92BD9D8A255" attach="left" box="[389,397,1790,1804]" fontSize="6" pageId="20" pageNumber="21">1</subScript>
slightly convergent. M
<subScript id="A9389A3F6A68FF92FD02F92BDAC2A255" attach="left" box="[655,663,1790,1804]" fontSize="6" pageId="20" pageNumber="21">4</subScript>
ending before wing margin, CuA+CuP ending closer to margin.
</paragraph>
<paragraph id="3503987A6A68FF92FF4AF8C7DBE2A2CE" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="21">
<emphasis id="07C844686A68FF92FF4AF8C7D8A9A272" box="[199,252,1810,1835]" italics="true" pageId="20" pageNumber="21">Legs</emphasis>
. Hind femur long, slightly swollen (
<figureCitation id="AD8784FF6A68FF92FD13F8C4DAA6A272" box="[670,755,1809,1835]" captionStart="FIGURES 82–88" captionStartId="104.[151,264,1799,1824]" captionTargetBox="[151,1436,174,1780]" captionTargetId="figure@104.[151,1436,174,1780]" captionTargetPageId="104" captionText="FIGURES 82–88. Gobrya sp. (Gobryidae); 82: female, dorsal; 83: same, lateral; 84: male, ventral; 85: female head, anterior; 86: same, dorsal; 87: male, Seram, Indonesia; 88: male, Sarawak, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.3670786" httpUri="https://zenodo.org/record/3670786/files/figure.png" pageId="20" pageNumber="21">Fig. 83</figureCitation>
). Femora ventrally with paired rows of thickened setae that may appear spine-like (sometimes reduced to base); spines darker and larger on the more posterior legs; fore and mid femora with rows of usually 4–6 spinose setae (smaller and fewer on posterior row), and hind femur with 3–6 spinulose setae on anterior row and 8–12 on tubercles on posterior row.
</paragraph>
<paragraph id="3503987A6A68FF93FF4AF874DD0DA5A2" blockId="20.[151,1437,153,2015]" lastBlockId="21.[151,1437,153,827]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="07C844686A68FF92FF4AF874D964A2E2" box="[199,305,1953,1979]" italics="true" pageId="20" pageNumber="21">Abdomen</emphasis>
. Abdomen straight, narrow (narrowest medially on segment 2), petiolate with apex slightly clavate. T1 and S1 broadly emarginated anteriorly to provide membranous space that folds to meet thorax when abdomen angled. T1 completely fused to T2, lateral margins evident (
<figureCitation id="AD8784FF6A69FF93FCBEFF4CDBD3A5EA" box="[819,902,153,179]" captionStart="FIGURES 89–97" captionStartId="105.[151,264,1799,1824]" captionTargetBox="[160,1428,182,1769]" captionTargetId="figure@105.[151,1436,174,1780]" captionTargetPageId="105" captionText="FIGURES 89–97. Gobrya sp. (Gobryidae), male abdomen; 89: right lateral; 90: ventral (right surstylus shaded); 91: terminalia, left lateral; 92: external genitalia, anterior (right surstylus and subepandrial sclerite shaded); 93: enlarged detail of subepandrial sclerite; 94: ejaculatory apodeme; 95: internal genitalia, right lateral; 96: same, ventral; 97: same, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3670788" httpUri="https://zenodo.org/record/3670788/files/figure.png" pageId="21" pageNumber="22">Fig. 89</figureCitation>
); S1 with reduced setation; T1+2 nearly as long as remining pregenitalic segments; T3 and S3 slightly longer than wide, tergites and sternites 4–6 wider than long, becoming progressively shorter posterioly. Spiracles
<quantity id="F244359F6A69FF93FD64FF35DB65A5A2" box="[745,816,224,251]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="21" pageNumber="22" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1–5 in</quantity>
membrane, spiracles 6 and 7 enclosed by tergite.
</paragraph>
<paragraph id="3503987A6A69FF93FF4AFED0D925A4D2" blockId="21.[151,1437,153,827]" pageId="21" pageNumber="22">
<emphasis id="07C844686A69FF93FF4AFED0D9D3A446" box="[199,390,261,287]" italics="true" pageId="21" pageNumber="22">Female genitalia</emphasis>
. (
<figureCitation id="AD8784FF6A69FF93FE11FED0DA48A479" box="[412,541,261,288]" captionStart="FIGURES 62–70" captionStartId="101.[151,264,1796,1821]" captionTargetBox="[158,1423,179,1765]" captionTargetId="figure@101.[151,1436,174,1777]" captionTargetPageId="101" captionText="FIGURES 62–70. Figs 62–64: Nothybus biguttatus Wulp (Nothybidae), female abdomen; 62: segment 7 to cercus, ventral; 63: same, dorsal; 64: internal genitalia. Figs 65–66: Somatia aestiva (Fabricius), female abdomen; 65: segment 7 to cerci; 66: internal genitalia. Figs 67–70: Gobrya sp., female abdomen; 67: segment 6 to cerci, dorsal; 68: same, right lateral; 69: same, ventral; 70: spermathecae." figureDoi="http://doi.org/10.5281/zenodo.3670780" httpUri="https://zenodo.org/record/3670780/files/figure.png" pageId="21" pageNumber="22">Figs 67–70</figureCitation>
) T6 and S6 fused with suture not evident. Segments 7, 8 and 10 very short with sclerites broad; S8 bare in material examined; T10 larger than S10. Cerci very short and broad. Spermathecae 1 + 2, with paired spermathecae on short stalks; spermathecae and apical section of ducts pigmented; spermathecae with sparse, minute pits.
</paragraph>
<paragraph id="3503987A6A69FF93FF4AFE40DA1EA662" blockId="21.[151,1437,153,827]" pageId="21" pageNumber="22">
<emphasis id="07C844686A69FF93FF4AFE40D938A4F6" box="[199,365,405,431]" italics="true" pageId="21" pageNumber="22">Male genitalia</emphasis>
. (
<figureCitation id="AD8784FF6A69FF93FE09FE40DA50A4F6" box="[388,517,405,431]" captionStart="FIGURES 89–97" captionStartId="105.[151,264,1799,1824]" captionTargetBox="[160,1428,182,1769]" captionTargetId="figure@105.[151,1436,174,1780]" captionTargetPageId="105" captionText="FIGURES 89–97. Gobrya sp. (Gobryidae), male abdomen; 89: right lateral; 90: ventral (right surstylus shaded); 91: terminalia, left lateral; 92: external genitalia, anterior (right surstylus and subepandrial sclerite shaded); 93: enlarged detail of subepandrial sclerite; 94: ejaculatory apodeme; 95: internal genitalia, right lateral; 96: same, ventral; 97: same, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3670788" httpUri="https://zenodo.org/record/3670788/files/figure.png" pageId="21" pageNumber="22">Figs 89–97</figureCitation>
) T6 fused to S8, which is nearly symmetrical; S6 and S7 absent, with pregenital sclerites membranous ventrally. Terminalia past T8 strongly asymmetrical; components of internal genitalia of tentative homology. Phallapodeme present (not absent, as stated in McAlpine (1997)). Epandrium subspherical, atrophied on left side, oblique; right anteroventral corner produced. Left surstylus small, pale, subrectangular; right surstylus larger, darker, bulbous. Subepandrial sclerite small, near ventral margin of epandrium and with one pair of small separate sclerites. Hypandrium bare, with medial bridge weakly sclerotized, flat and minutely setose; arms meeting dorsally; approximately U-shaped with right side strongly elaborated into several different posterior lobes and thin anterior carina. Phallapodeme short, carinate. Postgonite absent. Right pregonite absent; left pregonite (perhaps further elaboration of hypandrium?) flat, short and with more sclerotized margin. Epiphallus absent. Distiphal- lus short, stout, narrowest at base, and with cylindrical subbasal process with apical hairs; basiphallus produced as long left lateral lobe. Ejaculatory apodeme with broad, pale blade, and short thin stalk with narrow base; sperm pump cup-shaped with dark basal ring.
</paragraph>
</subSubSection>
</treatment>
</document>
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