171 lines
12 KiB
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171 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/fr.25.83034" ID-Pensoft-Pub="2193-0074-1-187" ID-Pensoft-UUID="86764B0ACD9453CD965B288014B441CD" ID-ZooBank="6402F8F152294153823FCAEA106F90A1" ModsDocID="2193-0074-25-1-187" checkinTime="1667366246929" checkinUser="pensoft" docAuthor="Spasojevic, Tamara, Broad, Gavin R. & Klopfstein, Seraina" docDate="2022" docId="73457721C4B254DA859BE208EA623014" docLanguage="en" docName="FossilRecord 25(1): 187-212" docOrigin="Fossil Record 25 (1)" docPubDate="2022-06-07" docSource="http://dx.doi.org/10.3897/fr.25.83034" docTitle="Pimpla seyrigi Theobald 1937" docType="treatment" docVersion="1" id="86764B0ACD9453CD965B288014B441CD" lastPageNumber="187" masterDocId="86764B0ACD9453CD965B288014B441CD" masterDocTitle="Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology" masterLastPageNumber="212" masterPageNumber="187" pageNumber="187" updateTime="1667366246929" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Spasojevic, Tamara</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5301-5722</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, Basel, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, Bern, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Broad, Gavin R.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7223-5333</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum, Cromwell Road, London SW 7 6 BD, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Klopfstein, Seraina</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4025-975X</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, Basel, Switzerland & Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, Bern, Switzerland</mods:affiliation>
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<mods:nameIdentifier type="email">seraina.klopfstein@bs.ch</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Fossil Record</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>25</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>187</mods:start>
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<mods:end>212</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/fr.25.83034</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.25.83034</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2193-0074-1-187</mods:identifier>
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<mods:identifier type="ZooBank">6402F8F152294153823FCAEA106F90A1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">86764B0ACD9453CD965B288014B441CD</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:73457721C4B254DA859BE208EA623014" httpUri="http://treatment.plazi.org/id/73457721C4B254DA859BE208EA623014" lastPageNumber="187" pageId="0" pageNumber="187">
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<subSubSection pageId="0" pageNumber="187" type="nomenclature">
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<paragraph pageId="0" pageNumber="187">
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<taxonomicName LSID="73457721-C4B2-54DA-859B-E208EA623014" authority="Theobald, 1937" authorityName="Theobald" authorityYear="1937" class="Insecta" family="Ichneumonidae" genus="Pimpla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pimpla seyrigi" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="seyrigi">Pimpla? seyrigi Theobald, 1937</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="187" type="description">
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<paragraph pageId="0" pageNumber="187">
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pimpla? seyrigi (Theobald, 1937), comb. nov. A. Photograph of the part and B. Counterpart of the holotype of taken at the NMBA; C, D. Corresponding interpretative drawings. In the drawings, thicker lines indicate outlines of body structures, thinner lines show characters inside these outlines, and dotted lines represent uncertain interpretations." figureDoi="10.3897/fr.25.83034.figure13" httpUri="https://binary.pensoft.net/fig/693072" pageId="0" pageNumber="187">Fig. 13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="187" type="reference_group">
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<paragraph pageId="0" pageNumber="187">
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<taxonomicName authorityName="Theobald" authorityYear="1937" class="Insecta" family="Ichneumonidae" genus="Pimpla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pimpla seyrigi" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="seyrigi">* Pimpla seyrigi</taxonomicName>
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Theobald,1937
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="187" type="materials_examined">
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<paragraph pageId="0" pageNumber="187">Material.</paragraph>
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<paragraph pageId="0" pageNumber="187">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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(part F1322/1 and counterpart F1322/2) examined at the NMBA.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="187" type="stratum">
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<paragraph pageId="0" pageNumber="187">Stratum.</paragraph>
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<paragraph pageId="0" pageNumber="187">Kleinkembs, Pays de Bade, Haut-Rhin, France. Early Oligocene (Rupelian), 33.9-28.4 Ma.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="187" type="description">
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<paragraph pageId="0" pageNumber="187">Description.</paragraph>
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<paragraph pageId="0" pageNumber="187">Sex unknown. Dorsal aspect of head (part), fairly complete antennae, mesosoma, almost complete fore wings and one nearly complete and one partial hind wing, partial hind legs, and metasoma present. Body length ~9.3 mm.</paragraph>
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<paragraph pageId="0" pageNumber="187">Mainly black, dark brown on wing veins, antennae and most of tergites, those with light end margins, legs lighter brown.</paragraph>
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<paragraph pageId="0" pageNumber="187">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Head</emphasis>
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seemingly elongate with long gena, rather than representing front view, given that antennae insert at the anterior edge.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Antenna</emphasis>
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almost as long as body, 1.25
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than fore wing, with more than 34 flagellomeres, evenly tapered from middle to apex.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Mesosoma</emphasis>
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not well preserved, with no details discernible.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Fore wing</emphasis>
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6.6 mm; areolet closed, somewhat petiolate anteriorly and almost triangular, with vein 2m-cu meeting 3M almost at outer corner; 2m-cu bowed outwards, probably with two bullae; 1cu-a meeting M + Cu clearly distally of 1M, 3Cu clearly longer than 2cu-a; cell 2R1 2.5
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Hind wing</emphasis>
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with cell R conspicuously narrow, vein 1Rs a bit longer than rs-m; cell 1Cu broad with vein M + Cu bowed; 1Cu only about half as long as cu-a.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Legs</emphasis>
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not well preserved; hind legs rather elongate; hind tibia 6.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Metasoma</emphasis>
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with T1 parallel-sided, 1.7
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than broad; T2 and following tergites transverse; each tergite except for T1 with a light band apically which from T4 becomes narrower medially until it is cut into two lateral spots.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Ovipositor</emphasis>
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not discernible.
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</paragraph>
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<caption doi="10.3897/fr.25.83034.figure13" httpUri="https://binary.pensoft.net/fig/693072" pageId="0" pageNumber="187" start="Figure 13" startId="F13">
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<paragraph pageId="0" pageNumber="187">
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<emphasis bold="true" pageId="0" pageNumber="187">Figure 13.</emphasis>
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<taxonomicName authorityName="Theobald" authorityYear="1937" class="Insecta" family="Ichneumonidae" genus="Pimpla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pimpla seyrigi" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="seyrigi">Pimpla? seyrigi</taxonomicName>
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(Theobald, 1937), comb. nov.
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<emphasis bold="true" pageId="0" pageNumber="187">A.</emphasis>
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Photograph of the part and
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<emphasis bold="true" pageId="0" pageNumber="187">B.</emphasis>
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Counterpart of the holotype of taken at the NMBA;
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<emphasis bold="true" pageId="0" pageNumber="187">C, D.</emphasis>
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Corresponding interpretative drawings. In the drawings, thicker lines indicate outlines of body structures, thinner lines show characters inside these outlines, and dotted lines represent uncertain interpretations.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="187" type="interpretation">
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<paragraph pageId="0" pageNumber="187">Interpretation.</paragraph>
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<paragraph pageId="0" pageNumber="187">
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The taxonomic affinity of this fossil is very difficult to discern. The elongate, tapering antennae and shape of the metasoma point to
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<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Ctenopelmatinae">Ctenopelmatinae</taxonomicName>
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, but some
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<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
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cannot be ruled out. In addition, the wide quadrate areolet and outwards curved 2m-cu are rarely seen in
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<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Ctenopelmatinae">Ctenopelmatinae</taxonomicName>
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, but they are quite common in
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<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
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. Given the poor preservation of the mesosoma, many important characters cannot be observed. As we cannot decide on the subfamily placement, removing the fossil from
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<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
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and labelling it as incertae subfamiliae would require description of a new genus based on the insufficient character evidence. We thus take a conservative approach, leaving the fossil within the genus
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<taxonomicName class="Insecta" family="Ichneumonidae" genus="Pimpla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pimpla" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="187">Pimpla</emphasis>
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</taxonomicName>
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, but emphasizing the uncertainty in this placement by adding a question mark behind both the genus and subfamily name.
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</paragraph>
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<paragraph pageId="0" pageNumber="187">
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<emphasis bold="true" pageId="0" pageNumber="187">
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<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
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Wesmael, 1845
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</emphasis>
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</paragraph>
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<paragraph pageId="0" pageNumber="187">
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<emphasis bold="true" pageId="0" pageNumber="187">
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<taxonomicName authorityName="Gravenhorst" authorityYear="1829" class="Insecta" family="Ichneumonidae" genus="Polysphincta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polysphincta" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="187">Polysphincta</emphasis>
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</taxonomicName>
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Gravenhorst, 1829
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</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |