treatments-xml/data/37/06/C9/3706C9276D14FFBAFFD4FE45FD93FEB7.xml

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<mods:title id="74939656CA32A644C3E4D5B111B067C7">Four new species of Pleroma (Melastomataceae) from campos rupestres and vegetation on granitic inselbergs in Eastern Minas Gerais, Brazil</mods:title>
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<mods:namePart id="0994510B002BAA8D6BB9310E06EC3EA7">Goldenberg, R.</mods:namePart>
<mods:affiliation id="77CE4EE38B03F6E6638FF6C0609CC45A">Departamento de Botânica, Universidade Federal do Paraná, Caixa Postal 19031, 81531 - 970 Curitiba - PR, Brazil. E-mail: renato. goldenberg @ gmail. com.</mods:affiliation>
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<mods:namePart id="01F77A5ED0256EFCE666444926354079">Gonella, P. M.</mods:namePart>
<mods:affiliation id="7DCDDF9A867D4DACD4F0C1860B79D98B">Universidade Federal de São João del-Rei, Campus Sete Lagoas, 35701 - 970 Sete Lagoas - MG, Brazil.</mods:affiliation>
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<mods:namePart id="501FDA294E7B337EE17263B2B3B55D7B">Meyer, F. S.</mods:namePart>
<mods:affiliation id="755E4E0F61C771928ABB1F177B3B8779">Herbarium UPCB and Pós-Graduação em Botânica, Universidade Federal do Paraná, Caixa Postal 19031, 81531 - 970 - Curitiba, PR, Brazil.</mods:affiliation>
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<mods:title id="E0FFBBDB11BE82848E25C75F1A983EC7">Edinburgh Journal of Botany</mods:title>
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<mods:date id="65797A790A53C63A564B3B0F92E7D7FF">2022</mods:date>
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<paragraph id="BF1078316D14FFB0FFD4FE45FCB0FE30" blockId="2.[121,771,438,463]" box="[121,771,438,463]" pageId="2" pageNumber="3">
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<emphasis id="8DDBA4236D14FFB0FFD4FE45FF3DFE30" bold="true" box="[121,142,438,463]" pageId="2" pageNumber="3">1.</emphasis>
<taxonomicName id="78AF03B26D14FFB0FF38FE45FD10FE30" ID-CoL="9C7RD" authority="F.S.Mey. &amp; R.Goldenb." authorityName="F. S. Mey. &amp; R. Goldenb." authorityYear="2022" box="[149,675,438,463]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="brevicomosum" status="sp. nov.">
<emphasis id="8DDBA4236D14FFB0FF38FE45FE17FE30" bold="true" box="[149,420,438,463]" pageId="2" pageNumber="3">Pleroma brevicomosum</emphasis>
F.S.Mey. &amp; R.Goldenb.
</taxonomicName>
, 
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<taxonomicNameLabel id="96E819586D14FFB0FD01FE45FCB0FE30" box="[684,771,438,463]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
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<taxonomicName id="78AF03B26D14FFB0FF3AFE28FE2DFE0B" box="[151,414,475,500]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D14FFB0FF3AFE28FE2DFE0B" box="[151,414,475,500]" italics="true" pageId="2" pageNumber="3">Pleroma brevicomosum</emphasis>
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differs from 
<taxonomicName id="78AF03B26D14FFB0FD9FFE28FBF7FE0B" authority="(Cogn.) P. J. F. Guim. &amp; Michelang." authorityName="P. J. F. Guim. &amp; Michelang." baseAuthorityName="Cogn." box="[562,1092,475,500]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="divaricatum">
<emphasis id="8DDBA4236D14FFB0FD9FFE28FD7EFE0B" box="[562,717,475,500]" italics="true" pageId="2" pageNumber="3">P. divaricatum</emphasis>
(Cogn.) P.J.F.Guim. &amp; Michelang.
</taxonomicName>
by its ovate leaves (versus oblong-lanceolate in 
<taxonomicName id="78AF03B26D14FFB0FD3FFDF2FC9FFDE5" box="[658,812,513,538]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="divaricatum">
<emphasis id="8DDBA4236D14FFB0FD3FFDF2FC9FFDE5" box="[658,812,513,538]" italics="true" pageId="2" pageNumber="3">P. divaricatum</emphasis>
</taxonomicName>
), these being chartaceous (versus membranaceous) and strigose on the adaxial surface (versus sericeous); slightly dimorphic stamens (versus pronouncedly dimorphic); elongate style, 
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long (versus short, c. 
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long); and ecostate capsules (versus costate).
</paragraph>
</subSubSection>
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– Type: 
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, 
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, 
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, 
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, 
<location id="BA702EEA6D14FFB0FCBFFD65FBBBFD50" LSID="urn:lsid:plazi:treatment:3706C9276D14FFBAFFD4FE45FD93FEB7:BA702EEA6D14FFB0FCBFFD65FBBBFD50" box="[786,1032,662,687]" country="Brazil" county="Type" latitude="-19.318068" longLatPrecision="1" longitude="-41.578403" municipality="Conselheiro Pena" name="Pico do Padre Angelo" pageId="2" pageNumber="3" stateProvince="Minas Gerais">Pico do Padre Ângelo</location>
, 
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, 
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, 
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, 
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, 
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<collectingDate id="DB55A7196D14FFB0FD70FD48FCE9FD2B" box="[733,858,699,724]" pageId="2" pageNumber="3" value="2020-07-11">11 vii 2020</collectingDate>
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, 
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<collectorName id="125A1DE76D14FFB0FCCEFD48FC5EFD2B" box="[867,1005,699,724]" pageId="2" pageNumber="3">P.M. Gonella</collectorName>
, 
<collectorName id="125A1DE76D14FFB0FC55FD48FBCBFD2B" box="[1016,1144,699,724]" pageId="2" pageNumber="3">R.S. Ribeiro</collectorName>
, 
<collectorName id="125A1DE76D14FFB0FF35FD12FE97FD05" box="[152,292,737,762]" pageId="2" pageNumber="3">G.A. da Silva</collectorName>
, 
<collectorName id="125A1DE76D14FFB0FE82FD12FE15FD05" box="[303,422,737,762]" pageId="2" pageNumber="3">A.P. Araújo</collectorName>
&amp; 
<collectorName id="125A1DE76D14FFB0FE6EFD12FDF1FD05" box="[451,578,737,762]" pageId="2" pageNumber="3">J.C. Ribeiro</collectorName>
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1425 (
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,
</materialsCitation>
<materialsCitation id="0FC7726C6D14FFB0FCE8FD12FB31FD05" box="[837,1154,737,762]" collectingDate="2020-07-11" collectionCode="MBML" collectorName="Gonella &amp; Ribeiro &amp; da Silva &amp; Araujo" country="Brazil" county="Type" elevation="1480" latitude="-19.318068" location="Serra do Padre Angelo" longLatPrecision="1" longitude="-41.578403" municipality="Conselheiro Pena" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Minas Gerais" typeStatus="isotype">
<typeStatus id="6014C6936D14FFB0FCE8FD12FC24FD05" box="[837,919,737,762]" pageId="2" pageNumber="3" type="isotype">isotype</typeStatus>
<collectionCode id="D9BEE0F46D14FFB0FC33FD12FC5DFD05" box="[926,1006,737,762]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15042" lsid="urn:lsid:biocol.org:col:15042" name="Museu de Biologia Mello Leitão" pageId="2" pageNumber="3" type="Herbarium">MBML</collectionCode>
). 
<emphasis id="8DDBA4236D14FFB0FC53FD12FB31FD05" bold="true" box="[1022,1154,737,762]" pageId="2" pageNumber="3">
<figureCitation id="279464B46D14FFB0FC53FD12FBD4FD05" box="[1022,1127,737,762]" captionStart="Figure 1" captionStartId="3.[121,185,1422,1444]" captionTargetBox="[223,1062,265,1399]" captionTargetId="figure-150@3.[218,1067,262,1401]" captionTargetPageId="3" captionText="Figure 1. Pleroma brevicomosum F.S.Mey. &amp; R.Goldenb., sp. nov. A, Branch with inflorescence; B, surface of a young branch; C, leaf (adaxial surface); D, leaf (abaxial surface); E, detail of the leaf indument (adaxial surface); F, detail of the leaf indumentum (abaxial surface); G, bracteole (abaxial surface); H, flower bud and bracteoles; I, detail of the indument on the hypanthium (abaxial surface); J, antesepalous stamen with a glabrous filament; K, antesepalous stamen with a pair of trichomes on the filament; L, antepetalous stamen; M, detail of the two trichomes shown in K; N, detail of the connective and appendages shown in L; O, detail of the connective and appendages shown in J; P, ovary, detached from the hypanthium; Q, ovary apex; R, immature capsule without the deciduous sepals. All from the holotype, Gonella et al. 1425 (UPCB). Photographs: F. S. Meyer." figureDoi="http://doi.org/10.5281/zenodo.10523468" httpUri="https://zenodo.org/record/10523468/files/figure.png" pageId="2" pageNumber="3">Figures 1</figureCitation>
, 
<figureCitation id="279464B46D14FFB0FBD9FD12FB31FD05" box="[1140,1154,737,762]" captionStart="Figure 2" captionStartId="4.[121,184,1653,1675]" captionTargetBox="[182,1103,265,1635]" captionTargetId="figure-6@4.[182,1103,265,1635]" captionTargetPageId="4" captionText="Figure 2. Pleroma brevicomosum in the field. A, Plant growing on quarZitic substrate; B, leaf pair (adaxial surfaces); C, inflorescence; D, flower (lateral view); E, immature capsules. Photographs: P. M. Gonella." figureDoi="http://doi.org/10.5281/zenodo.10523470" httpUri="https://zenodo.org/record/10523470/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
</emphasis>
</materialsCitation>
<emphasis id="8DDBA4236D14FFB0FB2FFD12FB3AFD05" bold="true" box="[1154,1161,737,762]" pageId="2" pageNumber="3">.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F7B52BBA6D14FFB7FFD4FCE6FC47FC3C" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
<paragraph id="BF1078316D14FFB7FFD4FCE6FC47FC3C" blockId="2.[121,1162,789,1710]" lastBlockId="5.[121,1151,266,963]" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
Erect to decumbent shrubs 
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tall, with sympodial growth, poorly branched. 
<emphasis id="8DDBA4236D14FFB0FFD7FCC9FEF2FCAC" box="[122,321,826,851]" italics="true" pageId="2" pageNumber="3">Younger branches</emphasis>
terete, not winged, moderately strigose, trichomes 
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long, unbranched, eglandular, appressed, the base linear, not immersed, not forked; 
<emphasis id="8DDBA4236D14FFB0FC4BFC93FB39FC86" box="[998,1162,864,889]" italics="true" pageId="2" pageNumber="3">older branches</emphasis>
terete, not winged, with indument similar to younger branches, but deciduous, and basally decorticating; 
<emphasis id="8DDBA4236D14FFB0FEB7FC59FEEDFC3C" box="[282,350,938,963]" italics="true" pageId="2" pageNumber="3">nodes</emphasis>
slender. 
<emphasis id="8DDBA4236D14FFB0FE69FC59FDA1FC3C" box="[452,530,938,963]" italics="true" pageId="2" pageNumber="3">Leaves</emphasis>
opposite; chartaceous, with petioles very short, 
<quantity id="7857D5D46D14FFB0FFD7FC23FF5DFC16" box="[122,238,976,1001]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="2" pageNumber="3" unit="mm" value="1.45" valueMax="2.0" valueMin="0.9">0.9–2 mm</quantity>
long; 
<emphasis id="8DDBA4236D14FFB0FE9EFC23FECFFC16" box="[307,380,976,1001]" italics="true" pageId="2" pageNumber="3">blades</emphasis>
2.2–4.3 × 
<quantity id="7857D5D46D14FFB0FE58FC23FDC4FC16" box="[501,631,976,1001]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.9" metricValueMin="1.9" pageId="2" pageNumber="3" unit="cm" value="2.4" valueMax="2.9" valueMin="1.9">1.9–2.9 cm</quantity>
, chartaceous, slightly discolorous, ovate, lacking domatia on the abaxial surface, base cordate, apex acute, margins crenulate, 5 acrodromous nerves, the marginals tenuous, adaxial surface flat, brown in dry specimens, bright green in fresh material, moderately strigose, trichomes 
<quantity id="7857D5D46D14FFB0FC82FBB3FC0AFBA6" box="[815,953,1088,1113]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="2" pageNumber="3" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
long, unbranched, eglandular, appressed, the base linear, not immersed, not forked, abaxial surface flat, light brown or pale green in dry specimens, pale green in fresh material, moderately strigose to setulose on the primary and secondary veins, trichomes 
<quantity id="7857D5D46D14FFB0FD5BFB43FC33FB36" box="[758,896,1200,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="2" pageNumber="3" unit="mm" value="1.3" valueMax="1.8" valueMin="0.8">0.8–1.8 mm</quantity>
long, unbranched, eglandular, appressed or curved, the base slightly broadened, not immersed, not forked, but moderately strigose to setulose on the surface, tertiary and quaternary veins, trichomes 
<quantity id="7857D5D46D14FFB0FFD7FAD3FEB0FAC6" box="[122,259,1312,1337]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="2" pageNumber="3" unit="mm" value="0.35" valueMax="0.6" valueMin="0.1">0.1–0.6 mm</quantity>
long, unbranched, eglandular, appressed, the base linear, not immersed, not forked. 
<emphasis id="8DDBA4236D14FFB0FF62FAB6FE88FAA1" box="[207,315,1349,1374]" italics="true" pageId="2" pageNumber="3">Thyrsoids</emphasis>
9–26.5 × 
<quantity id="7857D5D46D14FFB0FE00FAB6FDA6FAA1" box="[429,533,1349,1374]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="2" pageNumber="3" unit="cm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 cm</quantity>
, terminal, c.15–25 flowers, axis terete, with the same indumentum as the younger branches, reddish; 
<emphasis id="8DDBA4236D14FFB0FD3DFA99FD66FA7C" box="[656,725,1386,1411]" italics="true" pageId="2" pageNumber="3">bracts</emphasis>
late deciduous, leafy, petioles short, 
<quantity id="7857D5D46D14FFB0FFD7FA63FEB0FA56" box="[122,259,1424,1449]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="2" pageNumber="3" unit="mm" value="0.85" valueMax="1.1" valueMin="0.6">0.6–1.1 mm</quantity>
long, blade 21.4–24.9 × 
<quantity id="7857D5D46D14FFB0FDB6FA63FD70FA56" box="[539,707,1424,1449]" metricMagnitude="-2" metricUnit="m" metricValue="1.9549999999999996" metricValueMax="2.19" metricValueMin="1.72" pageId="2" pageNumber="3" unit="mm" value="19.549999999999997" valueMax="21.9" valueMin="17.2">17.2–21.9 mm</quantity>
, ovate, indumentum the same as on the leaves; 
<emphasis id="8DDBA4236D14FFB0FF60FA46FEF2FA31" box="[205,321,1461,1486]" italics="true" pageId="2" pageNumber="3">bracteoles</emphasis>
early deciduous, 5.5–6 × 
<quantity id="7857D5D46D14FFB0FDCFFA46FD5DFA31" box="[610,750,1461,1486]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="2" pageNumber="3" unit="mm" value="3.5" valueMax="3.7" valueMin="3.3">3.3–3.7 mm</quantity>
, elliptic or ovate, apex rounded, not covering the apex of the flower bud, margins entire, ciliate, adaxial surface glabrous, abaxial surface moderately strigose or moderately setulose, with indument uniformly arranged along the entire abaxial surface, trichomes 
<quantity id="7857D5D46D14FFB0FDCCF9D6FD59F9C1" box="[609,746,1573,1598]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="0.6" valueMax="0.9" valueMin="0.3">0.3–0.9 mm</quantity>
long, unbranched, eglandular, appressed or curved, the base linear or slightly broadened, not immersed, not forked. 
<emphasis id="8DDBA4236D14FFB0FFD7F983FF62F976" box="[122,209,1648,1673]" italics="true" pageId="2" pageNumber="3">Flowers</emphasis>
5-merous, pedicels 
<quantity id="7857D5D46D14FFB0FE1FF983FD94F976" box="[434,551,1648,1673]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="2" pageNumber="3" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.7–1 mm</quantity>
long; 
<emphasis id="8DDBA4236D14FFB0FDC1F983FD5DF976" box="[620,750,1648,1673]" italics="true" pageId="2" pageNumber="3">hypanthium</emphasis>
5.4–5.8 × 
<quantity id="7857D5D46D14FFB0FCCBF983FC6EF976" box="[870,989,1648,1673]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="2" pageNumber="3" unit="mm" value="5.15" valueMax="5.3" valueMin="5.0">5–5.3 mm</quantity>
, obovate, not costate, moderately setulose, trichomes 
<quantity id="7857D5D46D14FFB0FDEEF966FD7EF951" box="[579,717,1685,1710]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="13.0" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="0.85" valueMax="1.3" valueMin="0.4">0.4–1.3 mm</quantity>
long, unbranched, eglandular, curved, the base broadened, not immersed, not forked; 
<emphasis id="8DDBA4236D13FFB7FD23FEF9FD65FEDC" box="[654,726,266,291]" italics="true" pageId="5" pageNumber="6">sepals</emphasis>
early deciduous, 4–4.3 × 
<quantity id="7857D5D46D13FFB7FC55FEF9FBDDFEDC" box="[1016,1134,266,291]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="5" pageNumber="6" unit="mm" value="2.15" valueMax="2.3" valueMin="2.0">2–2.3 mm</quantity>
, triangular, margins ciliate, apex acute, adaxial surface glabrous, abaxial surface with the same trichomes as the hypanthium, but restricted to its central portion; 
<emphasis id="8DDBA4236D13FFB7FC0FFEA7FC55FE92" box="[930,998,340,365]" italics="true" pageId="5" pageNumber="6">petals</emphasis>
purple with a white base (during anthesis) or purple with a red base (in senescent flowers), 22.7–25.8 × 
<quantity id="7857D5D46D13FFB7FFD4FE6CFEBFFE47" box="[121,268,415,440]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" metricValueMax="2.12" metricValueMin="1.9" pageId="5" pageNumber="6" unit="mm" value="20.1" valueMax="21.2" valueMin="19.0">19–21.2 mm</quantity>
, obovate, apex cuspidate or mucronate, ciliate; 
<emphasis id="8DDBA4236D13FFB7FC8EFE6CFC30FE47" box="[803,899,415,440]" italics="true" pageId="5" pageNumber="6">stamens</emphasis>
10, slightly dimorphic, antesepalous with the filaments white (during anthesis) or purple to reddish (in senescent flowers), 
<quantity id="7857D5D46D13FFB7FF4CFE19FED9FDFC" box="[225,362,490,515]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="8.3" metricValueMin="7.3" pageId="5" pageNumber="6" unit="mm" value="7.8" valueMax="8.3" valueMin="7.3">7.3–8.3 mm</quantity>
long, glabrous or sparsely setulose on its central portion, trichomes 
<quantity id="7857D5D46D13FFB7FFD4FDFCFEB0FDD7" box="[121,259,527,552]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="6" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
long, unbranched, eglandular, curved to erect, the base linear, not immersed, not forked, pedoconnective purple, 
<quantity id="7857D5D46D13FFB7FDA8FDC7FD3CFDB2" box="[517,655,564,589]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="5" pageNumber="6" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6–0.9 mm</quantity>
prolonged below the thecae, glabrous, ventral appendages bilobed, apex obtuse, c. 
<quantity id="7857D5D46D13FFB7FDCEFDA9FD0FFD8C" box="[611,700,602,627]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="6" unit="mm" value="0.4">0.4 mm</quantity>
long, glabrous, thecae 7.1–7.6 × 
<quantity id="7857D5D46D13FFB7FFD7FD8CFF43FD67" box="[122,240,639,664]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="5" pageNumber="6" unit="mm" value="1.05" valueMax="1.1" valueMin="1.0">1–1.1 mm</quantity>
, falcate, purple, antepetalous with filaments white (during anthesis) or purple to reddish (in senescent flowers), 
<quantity id="7857D5D46D13FFB7FE5AFD57FD32FD42" box="[503,641,676,701]" metricMagnitude="-3" metricUnit="m" metricValue="6.85" metricValueMax="7.1" metricValueMin="6.6" pageId="5" pageNumber="6" unit="mm" value="6.85" valueMax="7.1" valueMin="6.6">6.6–7.1 mm</quantity>
long, glabrous, pedoconnective purple, 
<quantity id="7857D5D46D13FFB7FFD4FD39FEB0FD1C" box="[121,259,714,739]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
prolonged below the thecae, glabrous, ventral appendages bilobed, apex obtuse, c. 
<quantity id="7857D5D46D13FFB7FF4EFD1CFE8EFCF7" box="[227,317,751,776]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
long, glabrous, thecae 6.4–6.7 × 
<quantity id="7857D5D46D13FFB7FD19FD1CFD44FCF7" box="[692,759,751,776]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="mm" value="1.0">1 mm</quantity>
, falcate, purple; 
<emphasis id="8DDBA4236D13FFB7FC00FD1CFC5AFCF7" box="[941,1001,751,776]" italics="true" pageId="5" pageNumber="6">ovary</emphasis>
5.5–6.1 × 
<quantity id="7857D5D46D13FFB7FFD7FCE7FEB6FCD2" box="[122,261,788,813]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="5" pageNumber="6" unit="mm" value="5.05" valueMax="5.2" valueMin="4.9">4.9–5.2 mm</quantity>
, 5-locular, apex densely sericeous, trichomes 
<quantity id="7857D5D46D13FFB7FCAAFCE7FC22FCD2" box="[775,913,788,813]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="13.0" metricValueMin="5.0" pageId="5" pageNumber="6" unit="mm" value="0.9" valueMax="1.3" valueMin="0.5">0.5–1.3 mm</quantity>
long, unbranched, eglandular, appressed, the base linear, not immersed, not forked; 
<emphasis id="8DDBA4236D13FFB7FCF8FCC9FC3AFCAC" box="[853,905,826,851]" italics="true" pageId="5" pageNumber="6">style</emphasis>
purple in the basal three-quarters and white in its apical portion (both during anthesis and in senescent flowers), 
<quantity id="7857D5D46D13FFB7FF4CFC77FE34FC62" box="[225,391,900,925]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" metricValueMax="1.41" metricValueMin="1.33" pageId="5" pageNumber="6" unit="mm" value="13.7" valueMax="14.1" valueMin="13.3">13.3–14.1 mm</quantity>
long, apex curved, glabrous, stigma truncate. 
<emphasis id="8DDBA4236D13FFB7FC20FC77FB83FC62" box="[909,1072,900,925]" italics="true" pageId="5" pageNumber="6">Capsular fruits</emphasis>
6.3–8 × 
<quantity id="7857D5D46D13FFB7FF23FC59FEB6FC3C" box="[142,261,938,963]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="8.1" metricValueMin="6.0" pageId="5" pageNumber="6" unit="mm" value="7.05" valueMax="8.1" valueMin="6.0">6–8.1 mm</quantity>
, sepals early deciduous, epicarp undivided when mature, ecostate.
</paragraph>
</subSubSection>
<caption id="EBD028B96D15FFB1FFD4FA7DFC0FF950" ID-DOI="http://doi.org/10.5281/zenodo.10523468" ID-Zenodo-Dep="10523468" httpUri="https://zenodo.org/record/10523468/files/figure.png" pageId="3" pageNumber="4" startId="3.[121,185,1422,1444]" targetBox="[223,1062,265,1399]" targetPageId="3" targetType="figure">
<paragraph id="BF1078316D15FFB1FFD4FA7DFC0FF950" blockId="3.[121,1131,1422,1711]" pageId="3" pageNumber="4">
<emphasis id="8DDBA4236D15FFB1FFD4FA7DFF61FA5B" bold="true" box="[121,210,1422,1444]" pageId="3" pageNumber="4">Figure 1.</emphasis>
<taxonomicName id="78AF03B26D15FFB1FF74FA7DFD1BFA5B" authority="F.S.Mey. &amp; R.Goldenb." authorityName="F. S. Mey. &amp; R. Goldenb." authorityYear="2022" box="[217,680,1422,1444]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="brevicomosum" status="sp. nov.">
<emphasis id="8DDBA4236D15FFB1FF74FA7DFE77FA5B" box="[217,452,1422,1444]" italics="true" pageId="3" pageNumber="4">Pleroma brevicomosum</emphasis>
F.S.Mey. &amp; R.Goldenb.
</taxonomicName>
, 
<taxonomicNameLabel id="96E819586D15FFB1FD1DFA7DFD4CFA5B" box="[688,767,1422,1444]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
A, Branch with inflorescence; B, surface of a young branch; C, leaf (adaxial surface); D, leaf (abaxial surface); E, detail of the leaf indument (adaxial surface); F, detail of the leaf indumentum (abaxial surface); G, bracteole (abaxial surface); H, flower bud and bracteoles; I, detail of the indument on the hypanthium (abaxial surface); J, antesepalous stamen with a glabrous filament; K, antesepalous stamen with a pair of trichomes on the filament; L, antepetalous stamen; M, detail of the two trichomes shown in K; N, detail of the connective and appendages shown in L; O, detail of the connective and appendages shown in J; P, ovary, detached from the hypanthium; Q, ovary apex; R, immature capsule without the deciduous sepals. All from the holotype, 
<emphasis id="8DDBA4236D15FFB1FE08F96AFE43F950" box="[421,496,1689,1711]" italics="true" pageId="3" pageNumber="4">Gonella</emphasis>
et al. 1425 (UPCB). Photographs: F. S. Meyer.
</paragraph>
</caption>
<caption id="EBD028B96D12FFB6FFD4F986FBDCF952" ID-DOI="http://doi.org/10.5281/zenodo.10523470" ID-Zenodo-Dep="10523470" httpUri="https://zenodo.org/record/10523470/files/figure.png" pageId="4" pageNumber="5" startId="4.[121,184,1653,1675]" targetBox="[182,1103,265,1635]" targetPageId="4" targetType="figure">
<paragraph id="BF1078316D12FFB6FFD4F986FBDCF952" blockId="4.[121,1163,1653,1709]" pageId="4" pageNumber="5">
<emphasis id="8DDBA4236D12FFB6FFD4F986FF62F974" bold="true" box="[121,209,1653,1675]" pageId="4" pageNumber="5">Figure 2.</emphasis>
<taxonomicName id="78AF03B26D12FFB6FF75F986FE71F974" box="[216,450,1653,1675]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D12FFB6FF75F986FE71F974" box="[216,450,1653,1675]" italics="true" pageId="4" pageNumber="5">Pleroma brevicomosum</emphasis>
</taxonomicName>
in the field. A, Plant growing on quarZitic substrate; B, leaf pair (adaxial surfaces); C, inflorescence; D, flower (lateral view); E, immature capsules. Photographs: P. M. Gonella.
</paragraph>
</caption>
<subSubSection id="F7B52BBA6D13FFB7FFD4FC2DFDB5FA6D" pageId="5" pageNumber="6" type="distribution">
<paragraph id="BF1078316D13FFB7FFD4FC2DFDB5FA6D" blockId="5.[121,1156,990,1426]" pageId="5" pageNumber="6">
<emphasis id="8DDBA4236D13FFB7FFD4FC2DFECFFC08" box="[121,380,990,1015]" italics="true" pageId="5" pageNumber="6">Distribution and habitat</emphasis>
. 
<taxonomicName id="78AF03B26D13FFB7FE27FC2DFD22FC08" box="[394,657,990,1015]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D13FFB7FE27FC2DFD22FC08" box="[394,657,990,1015]" italics="true" pageId="5" pageNumber="6">Pleroma brevicomosum</emphasis>
</taxonomicName>
was collected around the higher peaks of the João Pinto Formation, east 
<collectingRegion id="7D6BB6D36D13FFB7FE57FBF0FD23FBE3" box="[506,656,1027,1052]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="6">Minas Gerais</collectingRegion>
, Southeast 
<collectingCountry id="C7B838A16D13FFB7FCBEFBF0FCE1FBE3" box="[787,850,1027,1052]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
(
<figureCitation id="279464B46D13FFB7FCCDFBF0FC0FFBE3" box="[864,956,1027,1052]" captionStart="Figure 3" captionStartId="6.[121,185,1007,1029]" captionTargetBox="[121,1170,255,985]" captionTargetId="figure-240@6.[121,1170,255,985]" captionTargetPageId="6" captionText="Figure 3. Distribution in Minas Gerais (MG), Brazil, of the four new species described in this article. BA, Bahia; ES, Espírito Santo; RJ, Rio de Janeiro." figureDoi="http://doi.org/10.5281/zenodo.10523473" httpUri="https://zenodo.org/record/10523473/files/figure.png" pageId="5" pageNumber="6">
<emphasis id="8DDBA4236D13FFB7FCCDFBF0FC0FFBE3" bold="true" box="[864,956,1027,1052]" pageId="5" pageNumber="6">Figure 3</emphasis>
</figureCitation>
), in campos rupestres on sandy soil and quartzitic rock outcrops, as both terrestrial and rupicolous. The species was found at Serra do Padre Ângelo (municipality of Conselheiro Pena), in the Pico do Padre Ângelo, around 
<quantity id="7857D5D46D13FFB7FE62FB80FDC3FB73" box="[463,624,1139,1164]" metricMagnitude="3" metricUnit="m" metricValue="1.51" metricValueMax="1.54" metricValueMin="1.48" pageId="5" pageNumber="6" unit="m" value="1510.0" valueMax="1540.0" valueMin="1480.0">1480–1540 m</quantity>
, and at the Pico do Sossego, around 
<quantity id="7857D5D46D13FFB7FBBFFB80FBD8FB73" box="[1042,1131,1139,1164]" metricMagnitude="3" metricUnit="m" metricValue="1.275" pageId="5" pageNumber="6" unit="m" value="1275.0">1275 m</quantity>
. It was also collected at the Pico da Aliança (municipality of Alvarenga), around 
<quantity id="7857D5D46D13FFB7FC5AFB6AFBFCFB4D" box="[1015,1103,1177,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="6" unit="m" value="1400.0">1400 m</quantity>
. The species was found syntopic with other endemic taxa of the region, such as 
<taxonomicName id="78AF03B26D13FFB7FC51FB4DFE78FB03" authority="Rivadavia &amp; Gonella" authorityName="Rivadavia &amp; Gonella" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis id="8DDBA4236D13FFB7FC51FB4DFF55FB03" italics="true" pageId="5" pageNumber="6">Drosera magnifica</emphasis>
Rivadavia &amp; Gonella
</taxonomicName>
(
<taxonomicName id="78AF03B26D13FFB7FE75FB10FDD8FB03" box="[472,619,1251,1276]" class="Magnoliopsida" family="Droseraceae" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Droseraceae</taxonomicName>
; 
<bibRefCitation id="DB3E05C06D13FFB7FDD8FB10FCFFFB03" author="Gonella PM &amp; Rivadavia F &amp; Fleischmann A." box="[629,844,1251,1276]" pageId="5" pageNumber="6" refId="ref13105" refString="Gonella PM, Rivadavia F, Fleischmann A. 2015. Drosera magnifica (Droseraceae): the largest New" type="book" year="2015">
Gonella 
<emphasis id="8DDBA4236D13FFB7FD7CFB10FCBBFB03" box="[721,776,1251,1276]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 2015
</bibRefCitation>
), 
<taxonomicName id="78AF03B26D13FFB7FCCDFB10FE9DFADD" authority="Loeuille &amp; Pirani" authorityName="Loeuille &amp; Pirani" class="Magnoliopsida" family="Asteraceae" genus="Eremanthus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="ovatifolius">
<emphasis id="8DDBA4236D13FFB7FCCDFB10FBECFB03" box="[864,1119,1251,1276]" italics="true" pageId="5" pageNumber="6">Eremanthus ovatifolius</emphasis>
Loeuille &amp; Pirani
</taxonomicName>
(
<taxonomicName id="78AF03B26D13FFB7FE96FAFAFE0CFADD" box="[315,447,1289,1314]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
; 
<bibRefCitation id="DB3E05C06D13FFB7FE64FAFAFD71FADD" author="Loeuille B &amp; Pirani JR" box="[457,706,1289,1314]" pageId="5" pageNumber="6" pagination="128 - 136" refId="ref13630" refString="Loeuille B, Pirani JR. 2016. Two new syncephalous species of Eremanthus (Asteraceae: Vernonieae) from southeastern Brazil. Phytotaxa. 243 (2): 128 - 136. https: // doi. org / 10.11646 / phytotaxa. 243.2.2" type="journal article" year="2016">Loeuille &amp; Pirani, 2016</bibRefCitation>
), 
<taxonomicName id="78AF03B26D13FFB7FD7AFAFAFF62FAB8" authority="Antar &amp; Loeuille" authorityName="Antar &amp; Loeuille" class="Magnoliopsida" family="Asteraceae" genus="Lepidaploa" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="campirupestris">
<emphasis id="8DDBA4236D13FFB7FD7AFAFAFC4DFADD" box="[727,1022,1289,1314]" italics="true" pageId="5" pageNumber="6">Lepidaploa campirupestris</emphasis>
Antar &amp; Loeuille
</taxonomicName>
(
<taxonomicName id="78AF03B26D13FFB7FF73FADDFED1FAB8" box="[222,354,1326,1351]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
; 
<bibRefCitation id="DB3E05C06D13FFB7FEC1FADDFD8AFAB8" author="Antar GM &amp; Siniscalchi CM &amp; Gonella PM &amp; Monge M &amp; Loeuille B." box="[364,569,1326,1351]" pageId="5" pageNumber="6" pagination="121 - 136" refId="ref12439" refString="Antar GM, Siniscalchi CM, Gonella PM, Monge M, Loeuille B. 2021 b. Novelties in Lepidaploinae (Asteraceae, Vernonieae) from the easternmost campos rupestres of Minas Gerais, Brazil: two new species and a range expansion. Plant Ecology and Evolution. 154: 121 - 136. https: // doi. org / 10.5091 / plecevo. 2021.1792" type="journal article" year="2021">
Antar 
<emphasis id="8DDBA4236D13FFB7FE1DFADDFE55FAB8" box="[432,486,1326,1351]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 2021b
</bibRefCitation>
) and 
<taxonomicName id="78AF03B26D13FFB7FDD7FADDFBEEFAB8" authority="Andrino &amp; Gonella" authorityName="Andrino &amp; Gonella" box="[634,1117,1326,1351]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="oreodoxus">
<emphasis id="8DDBA4236D13FFB7FDD7FADDFC3FFAB8" box="[634,908,1326,1351]" italics="true" pageId="5" pageNumber="6">Paepalanthus oreodoxus</emphasis>
Andrino &amp; Gonella
</taxonomicName>
(
<taxonomicName id="78AF03B26D13FFB7FFD2FAA0FE91FA93" box="[127,290,1363,1388]" class="Liliopsida" family="Eriocaulaceae" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Eriocaulaceae</taxonomicName>
; 
<bibRefCitation id="DB3E05C06D13FFB7FE80FAA0FD88FA93" author="Andrino CO &amp; Gonella PM" box="[301,571,1363,1388]" pageId="5" pageNumber="6" pagination="137 - 149" refId="ref12300" refString="Andrino CO, Gonella PM. 2021. An escape from the Espinhaco Range: a new species of Paepalanthus subg. Xeractis (Eriocaulaceae) from the campos rupestres of Serra do Padre Angelo, Minas Gerais, Brazil. Plant Ecology and Evolution. 154 (1): 137 - 149. https: // doi. org / 10.5091 / plecevo. 2021.1770" type="journal article" year="2021">Andrino &amp; Gonella, 2021</bibRefCitation>
). At the three localities, the species is syntopic with 
<taxonomicName id="78AF03B26D13FFB7FFD7FA8AFE8FFA6D" box="[122,316,1401,1426]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="caetanoi">
<emphasis id="8DDBA4236D13FFB7FFD7FA8AFE8FFA6D" box="[122,316,1401,1426]" italics="true" pageId="5" pageNumber="6">Pleroma caetanoi</emphasis>
</taxonomicName>
, described below.
</paragraph>
</subSubSection>
<subSubSection id="F7B52BBA6D13FFB7FFD4FA5EFC96FA39" box="[121,805,1453,1478]" pageId="5" pageNumber="6" type="biology_ecology">
<paragraph id="BF1078316D13FFB7FFD4FA5EFC96FA39" blockId="5.[121,805,1453,1478]" box="[121,805,1453,1478]" pageId="5" pageNumber="6">
<emphasis id="8DDBA4236D13FFB7FFD4FA5EFF5EFA39" box="[121,237,1453,1478]" italics="true" pageId="5" pageNumber="6">Phenology</emphasis>
. Collected with flowers and fruits in May and July.
</paragraph>
</subSubSection>
<subSubSection id="F7B52BBA6D13FFB4FFD4FA11FBE1FA95" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="conservation">
<paragraph id="BF1078316D13FFB4FFD4FA11FBE1FA95" blockId="5.[121,1150,1506,1717]" lastBlockId="6.[121,1163,1137,1386]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="8DDBA4236D13FFB7FFD4FA11FDACFA04" box="[121,543,1506,1531]" italics="true" pageId="5" pageNumber="6">Proposed IUCN conservation category</emphasis>
. Critically Endangered: CR B1ab(iii). With estimates of EOO = 
<quantity id="7857D5D46D13FFB7FF48F9F4FEE8F9DF" box="[229,347,1543,1568]" metricMagnitude="4" metricUnit="m" metricValue="4.3378" pageId="5" pageNumber="6" unit="km" value="43.378">43.378 km</quantity>
<superScript id="48DAD5796D13FFB7FEF1F9F5FED7F9EA" attach="left" box="[348,356,1542,1557]" fontSize="6" pageId="5" pageNumber="6">2</superScript>
and AOO = 
<quantity id="7857D5D46D13FFB7FE46F9F4FD82F9DF" box="[491,561,1543,1568]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="km" value="12.0">12 km</quantity>
<superScript id="48DAD5796D13FFB7FD9CF9F5FD8AF9EA" attach="left" box="[561,569,1542,1557]" fontSize="6" pageId="5" pageNumber="6">2</superScript>
, 
<taxonomicName id="78AF03B26D13FFB7FDE8F9F4FCFEF9DF" box="[581,845,1543,1568]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D13FFB7FDE8F9F4FCFEF9DF" box="[581,845,1543,1568]" italics="true" pageId="5" pageNumber="6">Pleroma brevicomosum</emphasis>
</taxonomicName>
is isolated at the higher areas of the João Pinto Formation, and has a severely fragmented distribution, which hinders pollen and seed dispersal among the areas. The three areas where the species was found are under active invasion by African grasses, most remarkably 
<taxonomicName id="78AF03B26D13FFB7FC29F984FF64F94A" authority="P. Beauv." authorityName="P. Beauv." class="Liliopsida" family="Poaceae" genus="Melinis" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="minutiflora">
<emphasis id="8DDBA4236D13FFB7FC29F984FBE1F96F" box="[900,1106,1655,1680]" italics="true" pageId="5" pageNumber="6">Melinis minutiflora</emphasis>
P.Beauv.
</taxonomicName>
(
<taxonomicName id="78AF03B26D13FFB7FF4EF96FFEFFF94A" box="[227,332,1692,1717]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
), which is found even in pristine areas that are not in direct contact with pastures, including the habitat of this species. The subpopulation at Pico do Padre Ângelo was directly affected by a catastrophic anthropogenic fire in early 
<date id="CB115EF16D10FFB4FCCDFB65FC4FFB50" box="[864,1020,1174,1199]" pageId="6" pageNumber="7" value="2020-10">October 2020</date>
, which also affected other endemics (
<bibRefCitation id="DB3E05C06D10FFB4FE7DFB48FD51FB2B" author="Andrino CO &amp; Gonella PM" box="[464,738,1211,1236]" pageId="6" pageNumber="7" pagination="137 - 149" refId="ref12300" refString="Andrino CO, Gonella PM. 2021. An escape from the Espinhaco Range: a new species of Paepalanthus subg. Xeractis (Eriocaulaceae) from the campos rupestres of Serra do Padre Angelo, Minas Gerais, Brazil. Plant Ecology and Evolution. 154 (1): 137 - 149. https: // doi. org / 10.5091 / plecevo. 2021.1770" type="journal article" year="2021">Andrino &amp; Gonella, 2021</bibRefCitation>
; 
<bibRefCitation id="DB3E05C06D10FFB4FD42FB48FC0FFB2B" author="Antar GM &amp; Siniscalchi CM &amp; Gonella PM &amp; Monge M &amp; Loeuille B." box="[751,956,1211,1236]" pageId="6" pageNumber="7" pagination="121 - 136" refId="ref12439" refString="Antar GM, Siniscalchi CM, Gonella PM, Monge M, Loeuille B. 2021 b. Novelties in Lepidaploinae (Asteraceae, Vernonieae) from the easternmost campos rupestres of Minas Gerais, Brazil: two new species and a range expansion. Plant Ecology and Evolution. 154: 121 - 136. https: // doi. org / 10.5091 / plecevo. 2021.1792" type="journal article" year="2021">
Antar 
<emphasis id="8DDBA4236D10FFB4FC9EFB48FCDAFB2B" box="[819,873,1211,1236]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 2021b
</bibRefCitation>
; 
<bibRefCitation id="DB3E05C06D10FFB4FC65FB48FEA0FB05" author="Kollmann LJC &amp; Gonella PM" pageId="6" pageNumber="7" refId="ref13509" refString="Kollmann LJC, Gonella PM. 2021. Novelties in Begonia (Begoniaceae) from the campos rupestres" type="book" year="2021">Kollmann &amp; Gonella, 2021</bibRefCitation>
); post-fire resprout has not been monitored. At all sites, the populations of 
<taxonomicName id="78AF03B26D10FFB4FFD4FAF5FE33FAE0" box="[121,384,1286,1311]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D10FFB4FFD4FAF5FE33FAE0" box="[121,384,1286,1311]" italics="true" pageId="6" pageNumber="7">Pleroma brevicomosum</emphasis>
</taxonomicName>
are small and scattered, with fewer than 10 flowering individuals observed at each site during fieldwork. Based on the available data, the species is, therefore, preliminarily assessed as Critically Endangered according to IUCN criteria (
<bibRefCitation id="DB3E05C06D10FFB4FC6FFAA2FBF0FA95" author="IUCN" box="[962,1091,1361,1386]" pageId="6" pageNumber="7" refId="ref13362" refString="IUCN. 2012. IUCN Red List Categories and Criteria, version 3.1, 2 nd edition. IUCN Species Survival Commission. Gland, Switzerland, and Cambridge: International Union for Conservation of Nature." type="book" year="2012">IUCN, 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="EBD028B96D10FFB4FFD4FC1CFD84FBD8" ID-DOI="http://doi.org/10.5281/zenodo.10523473" ID-Zenodo-Dep="10523473" httpUri="https://zenodo.org/record/10523473/files/figure.png" pageId="6" pageNumber="7" startId="6.[121,185,1007,1029]" targetBox="[121,1170,255,985]" targetPageId="6" targetType="figure">
<paragraph id="BF1078316D10FFB4FFD4FC1CFD84FBD8" blockId="6.[121,1155,1007,1063]" pageId="6" pageNumber="7">
<emphasis id="8DDBA4236D10FFB4FFD4FC1CFF61FBFA" bold="true" box="[121,210,1007,1029]" pageId="6" pageNumber="7">Figure 3.</emphasis>
Distribution in Minas Gerais (MG), Brazil, of the four new species described in this article. BA, Bahia; ES, Espírito Santo; RJ, Rio de Janeiro.
</paragraph>
</caption>
<subSubSection id="F7B52BBA6D10FFB4FFD4FA76FD44FA3B" pageId="6" pageNumber="7" type="etymology">
<paragraph id="BF1078316D10FFB4FFD4FA76FD44FA3B" blockId="6.[121,1068,1413,1476]" pageId="6" pageNumber="7">
<emphasis id="8DDBA4236D10FFB4FFD4FA76FF5EFA61" box="[121,237,1413,1438]" italics="true" pageId="6" pageNumber="7">Etymology</emphasis>
. The specific epithet, 
<taxonomicName id="78AF03B26D10FFB4FE4EFA76FD34FA61" box="[483,647,1413,1438]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D10FFB4FE4EFA76FD34FA61" box="[483,647,1413,1438]" italics="true" pageId="6" pageNumber="7">brevicomosum</emphasis>
</taxonomicName>
, is related to the indument with short trichomes on the hypanthium and leaves of this species.
</paragraph>
</subSubSection>
<subSubSection id="F7B52BBA6D10FFB4FFD4FA2EFCABF9A8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="BF1078316D10FFB4FFD4FA2EFCABF9A8" blockId="6.[121,1135,1501,1623]" pageId="6" pageNumber="7">
<emphasis id="8DDBA4236D10FFB4FFD4FA2EFE06FA0C" box="[121,437,1501,1523]" italics="true" pageId="6" pageNumber="7">Additional specimens examined</emphasis>
. 
<materialsCitation id="0FC7726C6D10FFB4FE6CFA2EFC5DF9EB" collectingDate="2021-05-09" collectionCode="P, M, MBML, UPCB" collectorName="da Alianca" country="Brazil" elevation="1400" latitude="-19.395807" location="Pico da Alianca" longLatPrecision="1" longitude="-41.67042" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Minas Gerais">
<collectingCountry id="C7B838A16D10FFB4FE6CFA2EFDB4FA0C" box="[449,519,1501,1523]" name="Brazil" pageId="6" pageNumber="7">BRAZIL</collectingCountry>
. 
<emphasis id="8DDBA4236D10FFB4FDBDFA2EFD26FA0C" bold="true" box="[528,661,1501,1523]" pageId="6" pageNumber="7">
<collectingRegion id="7D6BB6D36D10FFB4FDBDFA2EFD26FA0C" box="[528,661,1501,1523]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="7">Minas Gerais</collectingRegion>
</emphasis>
: Alvarenga, 
<location id="BA702EEA6D10FFB4FCBDFA2EFC1CFA0C" LSID="urn:lsid:plazi:treatment:3706C9276D14FFBAFFD4FE45FD93FEB7:BA702EEA6D10FFB4FCBDFA2EFC1CFA0C" box="[784,943,1501,1523]" country="Brazil" latitude="-19.395807" longLatPrecision="1" longitude="-41.67042" name="Pico da Alianca" pageId="6" pageNumber="7" stateProvince="Minas Gerais">
Pico 
<collectorName id="125A1DE76D10FFB4FCEEFA2EFC1CFA0C" box="[835,943,1501,1523]" pageId="6" pageNumber="7">da Aliança</collectorName>
</location>
, no topo do pico, 
<geoCoordinate id="DA9B1EF66D10FFB4FFD4FA0DFEB4F9EB" box="[121,263,1534,1556]" degrees="19" direction="south" minutes="23" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="44.91" value="-19.395807">19°23′44.91′′S</geoCoordinate>
, 
<geoCoordinate id="DA9B1EF66D10FFB4FEBDFA0DFE10F9EB" box="[272,419,1534,1556]" degrees="41" direction="west" minutes="40" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="13.52" value="-41.67042">41°40′13.52′′W</geoCoordinate>
, 
<quantity id="7857D5D46D10FFB4FE01FA0DFE49F9EB" box="[428,506,1534,1556]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="7" unit="m" value="1400.0">
<elevation id="34829F026D10FFB4FE01FA0DFE49F9EB" box="[428,506,1534,1556]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="7" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, 
<date id="CB115EF16D10FFB4FDA9FA0DFDEEF9EB" box="[516,605,1534,1556]" pageId="6" pageNumber="7" value="2021-05-09">
<collectingDate id="DB55A7196D10FFB4FDA9FA0DFDEEF9EB" box="[516,605,1534,1556]" pageId="6" pageNumber="7" value="2021-05-09">9 v 2021</collectingDate>
</date>
, 
<emphasis id="8DDBA4236D10FFB4FDC8FA0DFD53F9EB" box="[613,736,1534,1556]" italics="true" pageId="6" pageNumber="7">
<collectionCode id="D9BEE0F46D10FFB4FDC8FA0DFDC7F9EB" box="[613,628,1534,1556]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="6" pageNumber="7" type="Herbarium">P</collectionCode>
. 
<collectionCode id="D9BEE0F46D10FFB4FDD5FA0DFD3FF9EB" box="[632,652,1534,1556]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="6" pageNumber="7" type="Herbarium">M</collectionCode>
. Gonella
</emphasis>
et al. 2866 (
<collectionCode id="D9BEE0F46D10FFB4FCF1FA0DFC11F9EB" box="[860,930,1534,1556]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15042" lsid="urn:lsid:biocol.org:col:15042" name="Museu de Biologia Mello Leitão" pageId="6" pageNumber="7" type="Herbarium">MBML</collectionCode>
, 
<collectionCode id="D9BEE0F46D10FFB4FC06FA0DFC59F9EB" box="[939,1002,1534,1556]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15175" lsid="urn:lsid:biocol.org:col:15175" name="Universidade Federal do Paraná" pageId="6" pageNumber="7" type="Herbarium">UPCB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0FC7726C6D10FFB4FC5AFA0DFCA6F9A8" collectingDate="2021-05-02" collectionCode="P, M, MBML" country="Brazil" county="Serra do Padre Angelo" elevation="1275" latitude="-19.239815" location="Pico do Sossego" longLatPrecision="1" longitude="-41.581253" municipality="Serra do Pinhao" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Minas Gerais">
Conselheiro Pena, 
<collectingCounty id="567100BD6D10FFB4FF1AF9D3FE29F9C9" box="[183,410,1568,1590]" pageId="6" pageNumber="7">Serra do Padre Ângelo</collectingCounty>
, 
<collectingMunicipality id="5F74E24B6D10FFB4FE0EF9D3FDF5F9C9" box="[419,582,1568,1590]" pageId="6" pageNumber="7">Serra do Pinhão</collectingMunicipality>
, subida para o 
<location id="BA702EEA6D10FFB4FD73F9D3FC39F9C9" LSID="urn:lsid:plazi:treatment:3706C9276D14FFBAFFD4FE45FD93FEB7:BA702EEA6D10FFB4FD73F9D3FC39F9C9" box="[734,906,1568,1590]" country="Brazil" county="Serra do Padre Angelo" latitude="-19.239815" longLatPrecision="1" longitude="-41.581253" municipality="Serra do Pinhao" name="Pico do Sossego" pageId="6" pageNumber="7" stateProvince="Minas Gerais">Pico do Sossego</location>
, 
<geoCoordinate id="DA9B1EF66D10FFB4FC3EF9D3FB92F9C9" box="[915,1057,1568,1590]" degrees="19" direction="south" minutes="14" orientation="latitude" pageId="6" pageNumber="7" precision="1" seconds="23.33" value="-19.239815">19°14′23.33′′S</geoCoordinate>
, 
<geoCoordinate id="DA9B1EF66D10FFB4FFD4F9B2FEBFF9A8" box="[121,268,1601,1623]" degrees="41" direction="west" minutes="34" orientation="longitude" pageId="6" pageNumber="7" precision="1" seconds="52.51" value="-41.581253">41°34′52.51′′W</geoCoordinate>
, 
<quantity id="7857D5D46D10FFB4FEB8F9B2FED0F9A8" box="[277,355,1601,1623]" metricMagnitude="3" metricUnit="m" metricValue="1.275" pageId="6" pageNumber="7" unit="m" value="1275.0">
<elevation id="34829F026D10FFB4FEB8F9B2FED0F9A8" box="[277,355,1601,1623]" metricMagnitude="3" metricUnit="m" metricValue="1.275" pageId="6" pageNumber="7" unit="m" value="1275.0">1275 m</elevation>
</quantity>
, 
<date id="CB115EF16D10FFB4FEC0F9B2FE75F9A8" box="[365,454,1601,1623]" pageId="6" pageNumber="7" value="2021-05-02">
<collectingDate id="DB55A7196D10FFB4FEC0F9B2FE75F9A8" box="[365,454,1601,1623]" pageId="6" pageNumber="7" value="2021-05-02">2 v 2021</collectingDate>
</date>
, 
<emphasis id="8DDBA4236D10FFB4FE63F9B2FDFAF9A8" box="[462,585,1601,1623]" italics="true" pageId="6" pageNumber="7">
<collectionCode id="D9BEE0F46D10FFB4FE63F9B2FE6EF9A8" box="[462,477,1601,1623]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="6" pageNumber="7" type="Herbarium">P</collectionCode>
. 
<collectionCode id="D9BEE0F46D10FFB4FE4CF9B2FE46F9A8" box="[481,501,1601,1623]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="6" pageNumber="7" type="Herbarium">M</collectionCode>
. Gonella
</emphasis>
et al. 2524 (
<collectionCode id="D9BEE0F46D10FFB4FD68F9B2FCBDF9A8" box="[709,782,1601,1623]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15042" lsid="urn:lsid:biocol.org:col:15042" name="Museu de Biologia Mello Leitão" pageId="6" pageNumber="7" type="Herbarium">MBML</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7B52BBA6D10FFBAFFD4F980FD93FEB7" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="discussion">
<paragraph id="BF1078316D10FFB5FFD4F980FE90FE6C" blockId="6.[121,1095,1651,1714]" lastBlockId="7.[121,1165,266,1709]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<taxonomicName id="78AF03B26D10FFB4FFD4F980FE33F973" box="[121,384,1651,1676]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D10FFB4FFD4F980FE33F973" box="[121,384,1651,1676]" italics="true" pageId="6" pageNumber="7">Pleroma brevicomosum</emphasis>
</taxonomicName>
is morphologically closely related to 
<taxonomicName id="78AF03B26D10FFB4FC89F980FC0CF973" box="[804,959,1651,1676]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="divaricatum">
<emphasis id="8DDBA4236D10FFB4FC89F980FC0CF973" box="[804,959,1651,1676]" italics="true" pageId="6" pageNumber="7">P. divaricatum</emphasis>
</taxonomicName>
; both occur in the state of 
<collectingRegion id="7D6BB6D36D10FFB4FEB1F96AFE03F94D" box="[284,432,1689,1714]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="7">Minas Gerais</collectingRegion>
and are erect shrubs with sessile or almost sessile leaves, thyrsoid inflorescences, and flowers with setulose hypanthia and purple petals. 
<taxonomicName id="78AF03B26D11FFB5FC55FEF9FEAEFEB7" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D11FFB5FC55FEF9FEAEFEB7" italics="true" pageId="7" pageNumber="8">Pleroma brevicomosum</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="78AF03B26D11FFB5FE1CFEDCFDFFFEB7" box="[433,588,303,328]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="divaricatum">
<emphasis id="8DDBA4236D11FFB5FE1CFEDCFDFFFEB7" box="[433,588,303,328]" italics="true" pageId="7" pageNumber="8">P. divaricatum</emphasis>
</taxonomicName>
by the characters described in the diagnosis, and also by its larger hypanthia, 5.4–5.8 × 
<quantity id="7857D5D46D11FFB5FDF5FEA7FD7EFE92" box="[600,717,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="7" pageNumber="8" unit="mm" value="5.15" valueMax="5.3" valueMin="5.0">5–5.3 mm</quantity>
(versus 2–2.3 × 
<quantity id="7857D5D46D11FFB5FC26FEA7FBB3FE92" box="[907,1024,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="7" pageNumber="8" unit="mm" value="2.2" valueMax="2.4" valueMin="2.0">2–2.4 mm</quantity>
in 
<taxonomicName id="78AF03B26D11FFB5FFD7FE89FEA7FE6C" box="[122,276,378,403]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="divaricatum">
<emphasis id="8DDBA4236D11FFB5FFD7FE89FEA7FE6C" box="[122,276,378,403]" italics="true" pageId="7" pageNumber="8">P. divaricatum</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="BF1078316D11FFB5FF34FE6CFC6FFDB2" blockId="7.[121,1165,266,1709]" pageId="7" pageNumber="8">
<taxonomicName id="78AF03B26D11FFB5FF34FE6CFE13FE47" ID-CoL="9C7RD" box="[153,416,415,440]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D11FFB5FF34FE6CFE13FE47" box="[153,416,415,440]" italics="true" pageId="7" pageNumber="8">Pleroma brevicomosum</emphasis>
</taxonomicName>
also resembles some species of 
<taxonomicName id="78AF03B26D11FFB5FCB1FE6CFCCBFE47" ID-CoL="6SJD" box="[796,888,415,440]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="genus">
<emphasis id="8DDBA4236D11FFB5FCB1FE6CFCCBFE47" box="[796,888,415,440]" italics="true" pageId="7" pageNumber="8">Pleroma</emphasis>
</taxonomicName>
formerly recognised in 
<taxonomicName id="78AF03B26D11FFB5FFD7FE37FDF6FE22" authority="Triana" authorityName="Triana" box="[122,581,452,477]" class="Magnoliopsida" family="Melastomataceae" genus="Tibouchina" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="section" section="Diotanthera">
<emphasis id="8DDBA4236D11FFB5FFD7FE37FF47FE22" box="[122,244,452,477]" italics="true" pageId="7" pageNumber="8">Tibouchina</emphasis>
Aubl. sect. 
<emphasis id="8DDBA4236D11FFB5FED4FE37FE4AFE22" box="[377,505,452,477]" italics="true" pageId="7" pageNumber="8">Diotanthera</emphasis>
Triana
</taxonomicName>
(
<emphasis id="8DDBA4236D11FFB5FDF8FE37FD24FE22" box="[597,663,452,477]" italics="true" pageId="7" pageNumber="8">sensu</emphasis>
<bibRefCitation id="DB3E05C06D11FFB5FD33FE37FCFFFE22" author="Cogniaux AC" box="[670,844,452,477]" pageId="7" pageNumber="8" pagination="288 - 418" refId="ref12605" refString="Cogniaux AC. 1885. Tibouchina. In: Martius CFP, Eichler AG, Urban I, editors. Flora Brasiliensis, volume 14, part 3. Munich: Frid. Fleischer. pp. 288 - 418." type="book chapter" year="1885">Cogniaux, 1885</bibRefCitation>
, 
<bibRefCitation id="DB3E05C06D11FFB5FCF5FE37FC22FE22" author="Cogniaux AC" box="[856,913,452,477]" pageId="7" pageNumber="8" pagination="197 - 278" refId="ref12646" refString="Cogniaux AC. 1891. Melastomacees. In: Candolle A, Candolle C, editors. Monographiae Phanerogamarum, volume 7. Paris: G. Masson. pp. 197 - 278." type="book chapter" year="1891">1891</bibRefCitation>
), such as 
<taxonomicName id="78AF03B26D11FFB5FBA8FE37FF70FDFC" ID-CoL="4K92Y" authority="Triana" authorityName="Triana" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="australe">
<emphasis id="8DDBA4236D11FFB5FBA8FE37FBC4FE22" box="[1029,1143,452,477]" italics="true" pageId="7" pageNumber="8">P. australe</emphasis>
Triana
</taxonomicName>
, 
<taxonomicName id="78AF03B26D11FFB5FF66FE19FD6AFDFC" ID-CoL="4K945" authority="(Cogn.) P. J. F. Guim. &amp; Michelang." authorityName="P. J. F. Guim. &amp; Michelang." baseAuthorityName="Cogn." box="[203,729,490,515]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="cordifolium">
<emphasis id="8DDBA4236D11FFB5FF66FE19FED1FDFC" box="[203,354,490,515]" italics="true" pageId="7" pageNumber="8">P. cordifolium</emphasis>
(Cogn.) P.J.F.Guim. &amp; Michelang.
</taxonomicName>
and 
<taxonomicName id="78AF03B26D11FFB5FCBDFE19FF4BFDD7" ID-CoL="4K975" authority="(Cogn.) P. J. F. Guim. &amp; Michelang." authorityName="P. J. F. Guim. &amp; Michelang." baseAuthorityName="Cogn." class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="mosenii">
<emphasis id="8DDBA4236D11FFB5FCBDFE19FC32FDFC" box="[784,897,490,515]" italics="true" pageId="7" pageNumber="8">P. mosenii</emphasis>
(Cogn.) P.J.F.Guim. &amp; Michelang.
</taxonomicName>
, by its ovate leaves, thyrsoid inflorescences, setulose hypanthia, stamens with glabrous filaments and pedoconnectives, glabrous styles and ecostate fruits.
</paragraph>
<paragraph id="BF1078316D11FFB5FF34FDA9FC91FC62" blockId="7.[121,1165,266,1709]" pageId="7" pageNumber="8">
<taxonomicName id="78AF03B26D11FFB5FF34FDA9FE13FD8C" ID-CoL="9C7RD" box="[153,416,602,627]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D11FFB5FF34FDA9FE13FD8C" box="[153,416,602,627]" italics="true" pageId="7" pageNumber="8">Pleroma brevicomosum</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="78AF03B26D11FFB5FD99FDA9FD15FD8C" ID-CoL="4K92Y" box="[564,678,602,627]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="australe">
<emphasis id="8DDBA4236D11FFB5FD99FDA9FD15FD8C" box="[564,678,602,627]" italics="true" pageId="7" pageNumber="8">P. australe</emphasis>
</taxonomicName>
by its terete branches (versus quadrangular in 
<taxonomicName id="78AF03B26D11FFB5FE9DFD8CFE12FD67" ID-CoL="4K92Y" box="[304,417,639,664]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="australe">
<emphasis id="8DDBA4236D11FFB5FE9DFD8CFE12FD67" box="[304,417,639,664]" italics="true" pageId="7" pageNumber="8">P. australe</emphasis>
</taxonomicName>
) and its smaller leaves, 2.2–4.3 × 
<quantity id="7857D5D46D11FFB5FC88FD8CFC16FD67" box="[805,933,639,664]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.9" metricValueMin="1.9" pageId="7" pageNumber="8" unit="cm" value="2.4" valueMax="2.9" valueMin="1.9">1.9–2.9 cm</quantity>
(versus 9.1–10.8 × 
<quantity id="7857D5D46D11FFB5FFD7FD57FF4DFD42" box="[122,254,676,701]" metricMagnitude="-2" metricUnit="m" metricValue="6.05" metricValueMax="7.5" metricValueMin="4.6" pageId="7" pageNumber="8" unit="cm" value="6.05" valueMax="7.5" valueMin="4.6">4.6–7.5 cm</quantity>
), with crenulate margins (versus serrated), only 5 nerves (versus 7–9), and short petioles, 
<quantity id="7857D5D46D11FFB5FF72FD39FEE0FD1C" box="[223,339,714,739]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="7" pageNumber="8" unit="mm" value="1.45" valueMax="2.0" valueMin="0.9">0.9–2 mm</quantity>
long (versus elongate, 
<quantity id="7857D5D46D11FFB5FDF7FD39FD41FD1C" box="[602,754,714,739]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="13.1" metricValueMin="4.9" pageId="7" pageNumber="8" unit="mm" value="9.0" valueMax="13.1" valueMin="4.9">4.9–13.1 mm</quantity>
long). It also differs from 
<taxonomicName id="78AF03B26D11FFB5FBB4FD39FF61FCF7" ID-CoL="4K92Y" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="australe">
<emphasis id="8DDBA4236D11FFB5FBB4FD39FF61FCF7" italics="true" pageId="7" pageNumber="8">Pleroma australe</emphasis>
</taxonomicName>
by its smaller hypanthia, 5.4–5.8 × 
<quantity id="7857D5D46D11FFB5FDCFFD1CFD64FCF7" box="[610,727,751,776]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="7" pageNumber="8" unit="mm" value="5.15" valueMax="5.3" valueMin="5.0">5–5.3 mm</quantity>
(versus 7.5–8.6 × 
<quantity id="7857D5D46D11FFB5FC07FD1CFB8BFCF7" box="[938,1080,751,776]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="7" pageNumber="8" unit="mm" value="5.4" valueMax="5.5" valueMin="5.3">5.3–5.5 mm</quantity>
); triangular, caducous (i.e. falling off immediately after anthesis) sepals, 
<quantity id="7857D5D46D11FFB5FC34FCE7FBBDFCD2" box="[921,1038,788,813]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="7" pageNumber="8" unit="mm" value="4.15" valueMax="4.3" valueMin="4.0">4–4.3 mm</quantity>
long (versus oblong, 
<quantity id="7857D5D46D11FFB5FE80FCC9FE05FCAC" box="[301,438,826,851]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="7.1" metricValueMin="5.6" pageId="7" pageNumber="8" unit="mm" value="6.35" valueMax="7.1" valueMin="5.6">5.6–7.1 mm</quantity>
long sepals persisting on the hypanthium until the fruits are almost mature, then falling off when they are mature); and antesepalous stamens with a short pedoconnective, 
<quantity id="7857D5D46D11FFB5FED7FC77FDB7FC62" box="[378,516,900,925]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="7" pageNumber="8" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6–0.9 mm</quantity>
(versus 
<quantity id="7857D5D46D11FFB5FDC8FC77FD69FC62" box="[613,730,900,925]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="7" pageNumber="8" unit="mm" value="4.15" valueMax="4.3" valueMin="4.0">4–4.3 mm</quantity>
long).
</paragraph>
<paragraph id="BF1078316D11FFB5FF34FC59FC39FB5C" blockId="7.[121,1165,266,1709]" pageId="7" pageNumber="8">
<taxonomicName id="78AF03B26D11FFB5FF34FC59FE13FC3C" box="[153,416,938,963]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D11FFB5FF34FC59FE13FC3C" box="[153,416,938,963]" italics="true" pageId="7" pageNumber="8">Pleroma brevicomosum</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="78AF03B26D11FFB5FD99FC59FD78FC3C" box="[564,715,938,963]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="cordifolium">
<emphasis id="8DDBA4236D11FFB5FD99FC59FD78FC3C" box="[564,715,938,963]" italics="true" pageId="7" pageNumber="8">P. cordifolium</emphasis>
</taxonomicName>
by its terete branches (versus quadrangular in 
<taxonomicName id="78AF03B26D11FFB5FE9DFC3CFE75FC17" box="[304,454,975,1000]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="cordifolium">
<emphasis id="8DDBA4236D11FFB5FE9DFC3CFE75FC17" box="[304,454,975,1000]" italics="true" pageId="7" pageNumber="8">P. cordifolium</emphasis>
</taxonomicName>
) and its smaller leaves, 2.2–4.3 × 
<quantity id="7857D5D46D11FFB5FCE7FC3CFC79FC17" box="[842,970,975,1000]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.9" metricValueMin="1.9" pageId="7" pageNumber="8" unit="cm" value="2.4" valueMax="2.9" valueMin="1.9">1.9–2.9 cm</quantity>
(versus 9–11.3 × 
<quantity id="7857D5D46D11FFB5FF23FC07FEA0FBF2" box="[142,275,1012,1037]" metricMagnitude="-2" metricUnit="m" metricValue="8.55" metricValueMax="9.7" metricValueMin="7.4" pageId="7" pageNumber="8" unit="cm" value="8.55" valueMax="9.7" valueMin="7.4">7.4–9.7 cm</quantity>
), with crenulate margins (versus serrated), only 5 nerves (versus 9), and shorter petioles, 
<quantity id="7857D5D46D11FFB5FE9BFBE9FE18FBCC" box="[310,427,1050,1075]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="7" pageNumber="8" unit="mm" value="1.45" valueMax="2.0" valueMin="0.9">0.9–2 mm</quantity>
long (versus 
<quantity id="7857D5D46D11FFB5FDE9FBE9FD66FBCC" box="[580,725,1050,1075]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" metricValueMax="1.98" metricValueMin="1.2" pageId="7" pageNumber="8" unit="mm" value="15.9" valueMax="19.8" valueMin="12.0">12–19.8 mm</quantity>
long). It also differs from 
<taxonomicName id="78AF03B26D11FFB5FC50FBE9FF44FBA7" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="cordifolium">
<emphasis id="8DDBA4236D11FFB5FC50FBE9FF44FBA7" italics="true" pageId="7" pageNumber="8">Pleroma cordifolium</emphasis>
</taxonomicName>
by its smaller hypanthia, 5.4–5.8 × 
<quantity id="7857D5D46D11FFB5FD2AFBCCFD4FFBA7" box="[647,764,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="7" pageNumber="8" unit="mm" value="5.15" valueMax="5.3" valueMin="5.0">5–5.3 mm</quantity>
(versus 7.7–7.9 × 
<quantity id="7857D5D46D11FFB5FC62FBCCFBEEFBA7" box="[975,1117,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.9" metricValueMin="4.6" pageId="7" pageNumber="8" unit="mm" value="5.25" valueMax="5.9" valueMin="4.6">4.6–5.9 mm</quantity>
); antesepalous stamens with a short pedoconnective, 
<quantity id="7857D5D46D11FFB5FD61FB97FCE5FB82" box="[716,854,1124,1149]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="7" pageNumber="8" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6–0.9 mm</quantity>
(versus 
<quantity id="7857D5D46D11FFB5FC1AFB97FBF2FB82" box="[951,1089,1124,1149]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.4" metricValueMin="5.7" pageId="7" pageNumber="8" unit="mm" value="6.05" valueMax="6.4" valueMin="5.7">5.7–6.4 mm</quantity>
long); and ovary apex covered with eglandular trichomes (versus glandular).
</paragraph>
<paragraph id="BF1078316D11FFB5FF34FB5CFCF5FA57" blockId="7.[121,1165,266,1709]" pageId="7" pageNumber="8">
<taxonomicName id="78AF03B26D11FFB5FF34FB5CFE13FB37" box="[153,416,1199,1224]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D11FFB5FF34FB5CFE13FB37" box="[153,416,1199,1224]" italics="true" pageId="7" pageNumber="8">Pleroma brevicomosum</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="78AF03B26D11FFB5FD99FB5CFD16FB37" box="[564,677,1199,1224]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="mosenii">
<emphasis id="8DDBA4236D11FFB5FD99FB5CFD16FB37" box="[564,677,1199,1224]" italics="true" pageId="7" pageNumber="8">P. mosenii</emphasis>
</taxonomicName>
by its terete branches (versus quadrangular in 
<taxonomicName id="78AF03B26D11FFB5FE9DFB27FE13FB12" box="[304,416,1236,1261]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="mosenii">
<emphasis id="8DDBA4236D11FFB5FE9DFB27FE13FB12" box="[304,416,1236,1261]" italics="true" pageId="7" pageNumber="8">P. mosenii</emphasis>
</taxonomicName>
) and its smaller leaves, 2.2–4.3 × 
<quantity id="7857D5D46D11FFB5FC8EFB27FC17FB12" box="[803,932,1236,1261]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.9" metricValueMin="1.9" pageId="7" pageNumber="8" unit="cm" value="2.4" valueMax="2.9" valueMin="1.9">1.9–2.9 cm</quantity>
(versus 7.5–11.6 × 
<quantity id="7857D5D46D11FFB5FF23FB09FEA0FAEC" box="[142,275,1274,1299]" metricMagnitude="-2" metricUnit="m" metricValue="6.6499999999999995" metricValueMax="8.7" metricValueMin="4.6" pageId="7" pageNumber="8" unit="cm" value="6.6499999999999995" valueMax="8.7" valueMin="4.6">4.6–8.7 cm</quantity>
), with only 5 nerves (versus 9–11), and shorter petioles, 
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long (versus 
<quantity id="7857D5D46D11FFB5FF7EFAECFEC9FAC7" box="[211,378,1311,1336]" metricMagnitude="-2" metricUnit="m" metricValue="2.3049999999999997" metricValueMax="3.53" metricValueMin="1.08" pageId="7" pageNumber="8" unit="mm" value="23.049999999999997" valueMax="35.3" valueMin="10.8">10.8–35.3 mm</quantity>
long); antesepalous stamens with a short pedoconnective, 
<quantity id="7857D5D46D11FFB5FFD7FAB7FEB0FAA2" box="[122,259,1348,1373]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="7" pageNumber="8" unit="mm" value="0.75" valueMax="0.9" valueMin="0.6">0.6–0.9 mm</quantity>
(versus 
<quantity id="7857D5D46D11FFB5FEC9FAB7FE6AFAA2" box="[356,473,1348,1373]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="7" pageNumber="8" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
long); and smaller fruits, 6.3–8 × 
<quantity id="7857D5D46D11FFB5FCFBFAB7FC79FAA2" box="[854,970,1348,1373]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="8.1" metricValueMin="6.0" pageId="7" pageNumber="8" unit="mm" value="7.05" valueMax="8.1" valueMin="6.0">6–8.1 mm</quantity>
(versus 7.3–9.5 × 
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), with caducous sepals (versus sepals persisting on the hypanthium until the fruits are almost mature, then falling off when they are mature).
</paragraph>
<paragraph id="BF1078316D11FFBAFF34FA47FD93FEB7" blockId="7.[121,1165,266,1709]" lastBlockId="8.[121,1135,266,328]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<taxonomicName id="78AF03B26D11FFB5FF34FA47FE13FA32" box="[153,416,1460,1485]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="brevicomosum">
<emphasis id="8DDBA4236D11FFB5FF34FA47FE13FA32" box="[153,416,1460,1485]" italics="true" pageId="7" pageNumber="8">Pleroma brevicomosum</emphasis>
</taxonomicName>
is related to 
<taxonomicName id="78AF03B26D11FFB5FD99FA47FBF9FA32" authority="(Cogn.) P. J. F. Guim. &amp; Michelang." authorityName="P. J. F. Guim. &amp; Michelang." baseAuthorityName="Cogn." box="[564,1098,1460,1485]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="ackermannii">
<emphasis id="8DDBA4236D11FFB5FD99FA47FD67FA32" box="[564,724,1460,1485]" italics="true" pageId="7" pageNumber="8">P. ackermannii</emphasis>
(Cogn.) P.J.F.Guim. &amp; Michelang.
</taxonomicName>
by its strigose branches; its leaves with a short petiole (
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long in 
<taxonomicName id="78AF03B26D11FFB5FC20FA29FB9FFA0C" box="[909,1068,1498,1523]" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="ackermannii">
<emphasis id="8DDBA4236D11FFB5FC20FA29FB9FFA0C" box="[909,1068,1498,1523]" italics="true" pageId="7" pageNumber="8">P. ackermannii</emphasis>
</taxonomicName>
), with a brown (in dried specimens) and moderately strigose adaxial surface, and with 5 nerves; and also by its elliptic or ovate bracteoles, and its glabrous style. It differs from 
<taxonomicName id="78AF03B26D11FFB5FC56F9D7FEB3F99C" class="Magnoliopsida" family="Melastomataceae" genus="Pleroma" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="ackermannii">
<emphasis id="8DDBA4236D11FFB5FC56F9D7FEB3F99C" italics="true" pageId="7" pageNumber="8">Pleroma ackermannii</emphasis>
</taxonomicName>
by its ovate leaves (versus elliptic), with a cordate base (versus obtuse), these being smaller, 2.2–4.3 × 
<quantity id="7857D5D46D11FFB5FE23F99CFDBCF977" box="[398,527,1647,1672]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.9" metricValueMin="1.9" pageId="7" pageNumber="8" unit="cm" value="2.4" valueMax="2.9" valueMin="1.9">1.9–2.9 cm</quantity>
(versus 3.8–5 × 
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); its bracteoles with the indument arranged along the entire abaxial surface (versus on only the central portion); its setulose hypanthia (versus sericeous); and its stamens with glabrous or sparsely setulose filaments (versus moderately villose).
</paragraph>
</subSubSection>
</treatment>
</document>