437 lines
28 KiB
XML
437 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1027.54888" ID-GBIF-Dataset="a108311a-65d1-431d-b108-40e585c2646b" ID-PMC="PMC8035127" ID-Pensoft-Pub="1313-2970-1027-1" ID-Pensoft-UUID="C5103AABD06058C595280B912CBE9B8B" ID-PubMed="33867800" ID-ZooBank="7D179ED7D7A540A2A972E07BC648F3B9" ModsDocID="1313-2970-1027-1" checkinTime="1617435236653" checkinUser="pensoft" docAuthor="Godwin, Rebecca L. & Bond, Jason E." docDate="2021" docId="CB612683080E53029075D30877C27500" docLanguage="en" docName="ZooKeys 1027: 1-165" docOrigin="ZooKeys 1027" docPubDate="2021-04-02" docSource="http://dx.doi.org/10.3897/zookeys.1027.54888" docTitle="Ummidia quepoa Godwin & Bond 2021, sp. nov." docType="treatment" docUuid="94DD790E-40BB-481B-868C-7670674A83EF" docUuidSource="ZooBank" docVersion="4" id="C5103AABD06058C595280B912CBE9B8B" lastPageNumber="1" masterDocId="C5103AABD06058C595280B912CBE9B8B" masterDocTitle="Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae)" masterLastPageNumber="165" masterPageNumber="1" pageNumber="1" updateTime="1668151966284" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Godwin, Rebecca L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2439-2868</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology and Nematology, University of California, Davis 1 Shields Ave, Davis, CA, 95616</mods:affiliation>
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<mods:nameIdentifier type="email">rlc0004@tigermail.auburn.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bond, Jason E.</mods:namePart>
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<mods:affiliation>Department of Entomology and Nematology, University of California, Davis 1 Shields Ave, Davis, CA, 95616</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1027</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>165</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1027.54888</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1027.54888</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1027-1</mods:identifier>
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<mods:identifier type="ZooBank">7D179ED7D7A540A2A972E07BC648F3B9</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C5103AABD06058C595280B912CBE9B8B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="180850615" LSID="urn:lsid:zoobank.org:act:94DD790E-40BB-481B-868C-7670674A83EF" httpUri="http://treatment.plazi.org/id/CB612683080E53029075D30877C27500" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/94DD790E-40BB-481B-868C-7670674A83EF" authority="Godwin & Bond, 2021" authorityName="Godwin & Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia quepoa" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quepoa" status="sp. nov.">Ummidia quepoa</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 61" captionStartId="F66" captionText="Figure 61. Ummidia quepoa sp. nov. from Quepos, Puntarenas, Costa Rica. A male habitus illustration UMM 757 B female habitus illustration UMM 210. Scale bars: 4.0 mm." figureDoi="10.3897/zookeys.1027.54888.figure61" httpUri="https://binary.pensoft.net/fig/527525" pageId="0" pageNumber="1">Figs 61</figureCitation>
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<figureCitation captionStart="Figure 62" captionStartId="F67" captionText="Figure 62. Ummidia quepoa sp. nov. from Quepos, Puntarenas, Costa Rica A-D male holotype (UMM 757) A prolateral aspect, leg I B retrolateral aspect, leg I C line drawings, leg I prolateral and retrolateral aspects D retrolateral aspect, pedipalp E cleared spermathecae female paratype (UMM 210). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1027.54888.figure62" httpUri="https://binary.pensoft.net/fig/527526" pageId="0" pageNumber="1">, 62</figureCitation>
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<figureCitation captionStart="Map 5" captionStartId="F46" captionText="Map 5. Locality records for southern Mexico and Central American species." figureDoi="10.3897/zookeys.1027.54888.map5" httpUri="https://binary.pensoft.net/fig/527503" pageId="0" pageNumber="1">, Map 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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HOLOTYPE: 1 ♂ (UMM757) from
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<normalizedToken originalValue="Lucía">Lucia</normalizedToken>
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Lutz farm, Quepos, Puntarenas, Costa Rica,
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<geoCoordinate degrees="9.4828" direction="north" orientation="latitude" precision="5" value="9.4828">9.4828</geoCoordinate>
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<geoCoordinate degrees="84.0948" direction="west" orientation="longitude" precision="5" value="-84.0948">-84.0948</geoCoordinate>
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1, 102 m a.s.l., 26-28.v.2010, coll. A Solis BME.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The specific epithet is a noun taken in apposition and refers to the Quepoa, who inhabited the region of the type locality during the colonial era.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Godwin & Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia quepoa" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quepoa">
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<emphasis italics="true" pageId="0" pageNumber="1">Ummidia quepoa</emphasis>
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</taxonomicName>
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are relatively large spiders (CL 8-13). Males can be distinguished from
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. hondurena" order="Araneae" pageId="0" pageNumber="1" rank="species" species="hondurena">
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<emphasis italics="true" pageId="0" pageNumber="1">U. hondurena</emphasis>
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</taxonomicName>
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,
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. yojoa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="yojoa">
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<emphasis italics="true" pageId="0" pageNumber="1">U. yojoa</emphasis>
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</taxonomicName>
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,
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. rugosa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="rugosa">
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<emphasis italics="true" pageId="0" pageNumber="1">U. rugosa</emphasis>
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</taxonomicName>
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, and
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. erema" order="Araneae" pageId="0" pageNumber="1" rank="species" species="erema">
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<emphasis italics="true" pageId="0" pageNumber="1">U. erema</emphasis>
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</taxonomicName>
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by having a relatively short embolus with a single even curve and from
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. yojoa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="yojoa">
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<emphasis italics="true" pageId="0" pageNumber="1">U. yojoa</emphasis>
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</taxonomicName>
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and
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. varablanca" order="Araneae" pageId="0" pageNumber="1" rank="species" species="varablanca">
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<emphasis italics="true" pageId="0" pageNumber="1">U. varablanca</emphasis>
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</taxonomicName>
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by the presence of many prolateral (23 vs 0) and retrolateral (44 vs 2-3) spines on tibia I. Males can be further differentiated from
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. rugosa" order="Araneae" pageId="0" pageNumber="1" rank="species" species="rugosa">
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<emphasis italics="true" pageId="0" pageNumber="1">U. rugosa</emphasis>
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</taxonomicName>
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and
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<taxonomicName family="Halonoproctidae" kingdom="Animalia" lsidName="U. riverai" order="Araneae" pageId="0" pageNumber="1" rank="species" species="riverai">
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<emphasis italics="true" pageId="0" pageNumber="1">U. riverai</emphasis>
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</taxonomicName>
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by the lack of a pale dorsal heart patch on the abdomen, though occasionally with a faint opalescence. Females can be differentiated from all geographically proximate species by having spermathecae which curve laterally and then bend posteriorly and then anteriorly at the distal end.
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</paragraph>
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<caption doi="10.3897/zookeys.1027.54888.figure61" httpUri="https://binary.pensoft.net/fig/527525" pageId="0" pageNumber="1" start="Figure 61" startId="F66">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 61.</emphasis>
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<taxonomicName authorityName="Godwin & Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia quepoa" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quepoa">
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<emphasis italics="true" pageId="0" pageNumber="1">Ummidia quepoa</emphasis>
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</taxonomicName>
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sp. nov. from Quepos, Puntarenas, Costa Rica.
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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male habitus illustration UMM757
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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female habitus illustration UMM210. Scale bars: 4.0 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description of male holotype.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Specimen preparation and condition</emphasis>
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. Specimen preserved in 80% EtOH.
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<emphasis italics="true" pageId="0" pageNumber="1">General coloration</emphasis>
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. Carapace and chelicerae reddish black 2.5YR 2.5/1, legs dark reddish brown 5YR 3/4. Abdomen very dark brown 10YR 2/2.
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<emphasis italics="true" pageId="0" pageNumber="1">Cephalothorax</emphasis>
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. Carapace 8.81 long, 8.35 wide. Pars cephalica 6.14 long. Foveal groove procurved, 0.84 long, 1.77 wide. All eyes on moderate tubercle, highest under AME. AER procurved. PER slightly recurved. Eye group 0.99 long, 2.07 wide, AME 0.48, PME 0.26, ALE 0.48, PLE 0.39. Sternum sparsely setose around outer 1/3, STRl 5.06, STRw 4.78. Chelicerae with anterior tooth row comprising five teeth, posterior margin with seven teeth. Palpal endites with 22 small cuspules across proximal half of endite face, lacking distal endite cuspules, ENDw 1.93, ENDl 3.56. Labium with three cuspules, LBw 1.67, LBl 1.25. Rastellum with many spines on process. Abdomen setose.
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<emphasis italics="true" pageId="0" pageNumber="1">Legs</emphasis>
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. F1 7.47; F1w 1.91; P1 3.8; Ti1 5.03; Mt1 3.09; Tr1 1.48; F3 5.99; F3w 2.36; P3 3.11; Ti3 3.76; Sd3 2.31; Mt3 3.5; Tr3 2.33; F4 8.12; F4w 2.27; P4 3.51; Ti4 4.91; Mt4 5.87; Tr4 2.71. Retrolateral face of tarsus IV with loose comb surrounded in setae. Leg I spination pattern: TSp 11, TSpv 13, TSrd 0, TSr 2, TSrv 42, MtSp 11, MtSr 16, TrSp 11, TrSr 16.
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<emphasis italics="true" pageId="0" pageNumber="1">Pedipalps</emphasis>
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. PTl 4, PTw 1.37, Bl 2.88. Embolus evenly curved.
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</paragraph>
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<caption doi="10.3897/zookeys.1027.54888.figure62" httpUri="https://binary.pensoft.net/fig/527526" pageId="0" pageNumber="1" start="Figure 62" startId="F67">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 62.</emphasis>
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<taxonomicName authorityName="Godwin & Bond" authorityYear="2021" class="Arachnida" family="Halonoproctidae" genus="Ummidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ummidia quepoa" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quepoa">
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<emphasis italics="true" pageId="0" pageNumber="1">Ummidia quepoa</emphasis>
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</taxonomicName>
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sp. nov. from Quepos, Puntarenas, Costa Rica
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<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
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male holotype (UMM757)
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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prolateral aspect, leg I
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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retrolateral aspect, leg I
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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line drawings, leg I prolateral and retrolateral aspects
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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retrolateral aspect, pedipalp
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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cleared spermathecae female paratype (UMM210). Scale bars: 1.0 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="variation">
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<paragraph pageId="0" pageNumber="1">Variation, males</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">(n = 4).</emphasis>
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CL 8.15-10.57, 9.18
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.51; CW 7.75-10.11, 8.64
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.51; Cap 5.73-7.2, 6.21
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.34; ENDl 0.91-1.25, 1.07
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.08; ENDw 1.74-2.31, 2.06
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.12; STRl 4.71-6.19, 5.27
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.32; STRw 4.32-5.56, 4.77
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.28; LBl 1.22-1.6, 1.33
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.09; LBw 1.49-1.98, 1.7
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.1; F1 6.8-10.43, 8.38
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.8; F1w 1.88-2.39, 2.02
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.12; P1 3.65-5.18, 4.22
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.34; Ti1 4.58-7.31, 5.81
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.62; Mt1 2.83-4.84, 3.7
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.46; Tr1 1.26-1.81, 1.56
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.12; F3 5.53-7.22, 6.14
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.37; F3w 2.28-2.87, 2.45
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.14; P3 2.98-3.87, 3.26
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2; Ti3 3.3-4.44, 3.78
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.24; Mt3 3.04-4.77, 3.71
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.37; Tr3 2.15-3.13, 2.5
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.22; F4 7.49-9.73, 8.41
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.47; F4w 2.04-2.6, 2.29
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.12; P4 3.21-4.23, 3.61
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.22; Ti4 4.76-6.57, 5.49
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.42; Mt4 5.2-7.45, 6.18
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.47; Tr4 2.41-3.19, 2.71
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.17; TSp 0-11, 7.5
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<normalizedToken originalValue="±">+/-</normalizedToken>
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2.6; TSpv 12-30, 20
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<normalizedToken originalValue="±">+/-</normalizedToken>
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4.45; TSr 1-2, 1.75
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.25; TSrv 25-57, 42.5
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<normalizedToken originalValue="±">+/-</normalizedToken>
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6.64; PTl 3.67-5.1, 4.4
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.34; PTw 1.27-1.61, 1.41
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.07; BL 2.39-3.38, 2.89
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description of female paratype.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Specimen preparation and condition</emphasis>
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. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen.
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<emphasis italics="true" pageId="0" pageNumber="1">General coloration</emphasis>
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. Carapace, chelicerae, and legs very dusky red 10R 2.5/2. Abdomen very dark brown 10YR 2/2, spinnerets yellowish brown 10YR 5/6.
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<emphasis italics="true" pageId="0" pageNumber="1">Cephalothorax</emphasis>
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. Carapace 13.04 long, 11.21 wide. Pars cephalica 8.59 long. Foveal groove procurved, 1.6 long, 3.48 wide. Eye tubercle moderate under ME. AER procurved. PER straight. Eye group 1.41 long, 2.1 wide, AME 0.47, PME 0.32, ALE 0.73, PLE 0.38. Sternum sparsely setose, STRl 7.56, STRw 7.08. Chelicerae with anterior row comprising six teeth, posterior margin with five teeth. Palpal endites with 19 cuspules spread across proximal half of endite and 29 smaller cuspules distally, ENDw 3.14, ENDl 4.86. Labium with seven cuspules, LBw 2.66, LBl 1.85. Rastellum with many strong spines on process and up cheliceral face for ~ 2
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<normalizedToken originalValue="×">x</normalizedToken>
|
||
length of process.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
||
. Evenly setose with pale speckles concentrated at apodemes and with a dorsal light patch.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Legs</emphasis>
|
||
. F1 7.77; F1w 2.73; P1 4.97; Ti1 5.18; Mt1 3.6; Tr1 1.58; F3 6.05; F3w 3.32; P3 4.69; Ti3 3.77, Sd3 2.3; Mt3 3.31; Tr3 2.27; F4 7.93; F4w 3.11; P4 4.86; Ti4 4.66; Mt4 5.33; Tr4 2.36. Retrolateral face tarsus IV with brush 2-3 spinules deep and setose ventrally.
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pedipalps</emphasis>
|
||
. PF 7.28, PP 4.38, PTi 5.06, PTr 3.95. Spermathecae with lateral and then anterior bend, bulbs facing anterolaterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="variation">
|
||
<paragraph pageId="0" pageNumber="1">Variation, females</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">(n = 2).</emphasis>
|
||
CL 10.75-12.1, 11.43
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.68; CW 9.38-10.69, 10.03
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.66; Cap 7.36-8.23, 7.8
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.44; ENDl 1.27-1.41, 1.34
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.07; ENDw 1.97-2.1, 2.04
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.07; STRl 6.66-8.01, 7.33
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.68; STRw 5.71-6.6, 6.15
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.45; LBl 1.78-1.84, 1.81
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.03; LBw 2.37-2.52, 2.45
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.07; F1 6.76-7.92, 7.34
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.58; F1w 2.36-2.82, 2.59
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.23; P1 4.1-4.83, 4.46
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.36; Ti1 4.34-5.26, 4.8
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.46; Mt1 2.99-3.49, 3.24
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.25; Tr1 1.52-1.63, 1.58
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.05; F3 5.77-6.62, 6.2
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.42; F3w 2.83-3.48, 3.16
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.33; P3 3.5-4.4, 3.95
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.45; Ti3 3.21-3.75, 3.48
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.27; Mt3 2.56-2.92, 2.74
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.18; Tr3 2.17-2.39, 2.28
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.11; F4 7.32-8.4, 7.86
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.54; F4w 2.77-3.31, 3.04
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.27; P4 3.87-4.57, 4.22
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.35; Ti4 3.96-4.63, 4.3
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.33; Mt4 4.34-5.42, 4.88
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.54; Tr4 1.93-2.53, 2.23
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.3; PF 6.11-7.34, 6.72
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.62; PP 3.63-4.36, 4
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.36; PTi 4.22-5.33, 4.78
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.56; PTr 3.19-3.93, 3.56
|
||
<normalizedToken originalValue="±">+/-</normalizedToken>
|
||
0.37.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<materialsCitation ID-GBIF-Occurrence="3067212301" collectingDate="2010-06-14" collectingDateMax="2015-12-08" collectingDateMin="2010-06-14" collectorName="JA Azofeifa, C Chacoa" country="Costa Rica" county="Guatuso" elevation="1478" latitude="10.898" location="Finca de Jose Martinez" longLatPrecision="55" longitude="-85.0195" municipality="Cabanga" specimenCount="2" specimenCount-male="2" stateProvince="Alajuela">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<collectingCountry name="Costa Rica">Costa Rica</collectingCountry>
|
||
:
|
||
<collectingRegion country="Costa Rica" name="Alajuela">Alajuela</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty>Guatuso</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Cabanga</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:CB612683080E53029075D30877C27500:642C02D737FF7861F02B70CC66FBDEBD" country="Costa Rica" county="Guatuso" latitude="10.898" longLatPrecision="55" longitude="-85.0195" municipality="Cabanga" name="Finca de Jose Martinez" stateProvince="Alajuela">
|
||
Finca de
|
||
<normalizedToken originalValue="José">Jose</normalizedToken>
|
||
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
|
||
</location>
|
||
, 10.59 -84.8528
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.478" unit="m" value="1478.0">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.478" unit="m" value="1478.0">1, 478 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(UMM0764,
|
||
<collectingDate value="2010-06-14" valueMax="2010-07-14" valueMin="2010-06-14">14.vi-14.vii.2010</collectingDate>
|
||
,
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
,
|
||
<collectorName>JA Azofeifa</collectorName>
|
||
, BME);
|
||
<location LSID="urn:lsid:plazi:treatment:CB612683080E53029075D30877C27500:F41953E77F9E6561474EC0DD8816A54E" country="Costa Rica" county="Guatuso" latitude="10.898" longLatPrecision="55" longitude="-85.0195" municipality="Cabanga" name="Upala" stateProvince="Alajuela">Upala</location>
|
||
,
|
||
<geoCoordinate degrees="10.898" direction="north" orientation="latitude" precision="55" value="10.898">10.898</geoCoordinate>
|
||
<geoCoordinate degrees="85.0195" direction="west" orientation="longitude" precision="5" value="-85.0195">-85.0195</geoCoordinate>
|
||
1, 300-
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(UMM0683,
|
||
<collectingDate value="2015-10-05" valueMax="2015-12-08" valueMin="2015-10-05">5.x-8.xii.2015</collectingDate>
|
||
,
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
,
|
||
<collectorName>C Chacoa</collectorName>
|
||
, BME)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3067212346" collectingDate="2010-05-26" collectingDateMax="2010-05-28" collectingDateMin="2010-05-26" collectorName="A Solis, Agujas" country="Costa Rica" elevation="1102" latitude="8.5794" location="Finca de Lucia Lutz" longLatPrecision="54" longitude="-83.387" municipality="Aguirre" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<collectingRegion country="Costa Rica" name="Puntarenas">Puntarenas</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality>Aguirre</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:CB612683080E53029075D30877C27500:CA72AAB4637657ED671E2BF2C4C6538F" country="Costa Rica" latitude="8.5794" longLatPrecision="54" longitude="-83.387" municipality="Aguirre" name="Finca de Lucia Lutz" stateProvince="Puntarenas">
|
||
Finca de
|
||
<normalizedToken originalValue="Lucía">Lucia</normalizedToken>
|
||
Lutz
|
||
</location>
|
||
,
|
||
<geoCoordinate degrees="9.4828" direction="north" orientation="latitude" precision="5" value="9.4828">9.4828</geoCoordinate>
|
||
<geoCoordinate degrees="84.0948" direction="west" orientation="longitude" precision="5" value="-84.0948">-84.0948</geoCoordinate>
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.1019999999999999" unit="m" value="1102.0">
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1019999999999999" unit="m" value="1102.0">1, 102 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(UMM0757,
|
||
<collectingDate value="2010-05-26" valueMax="2010-05-28" valueMin="2010-05-26">26-28.v.2010</collectingDate>
|
||
,
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
,
|
||
<collectorName>A Solis</collectorName>
|
||
, BME);
|
||
<collectorName>Agujas</collectorName>
|
||
,
|
||
<geoCoordinate degrees="8.5794" direction="north" orientation="latitude" precision="5" value="8.5794">8.5794</geoCoordinate>
|
||
<geoCoordinate degrees="83.387" direction="west" orientation="longitude" precision="55" value="-83.387">-83.387</geoCoordinate>
|
||
1, 29 m a.s.l. (UMM0676,
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
, BME)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="3067212374" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="N Banks" country="Costa Rica" latitude="9.933" location="San Jose" longLatPrecision="55" longitude="-84.0831" specimenCount="1" specimenCount-female="1" stateProvince="San Jose">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<collectingRegion country="Costa Rica" name="San Jose">
|
||
San
|
||
<normalizedToken originalValue="José">Jose</normalizedToken>
|
||
</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:CB612683080E53029075D30877C27500:99E05BF37E88E499A39DB2CF68FF5F05" country="Costa Rica" latitude="9.933" longLatPrecision="55" longitude="-84.0831" name="San Jose" stateProvince="San Jose">
|
||
San
|
||
<normalizedToken originalValue="José">Jose</normalizedToken>
|
||
</location>
|
||
,
|
||
<geoCoordinate degrees="9.933" direction="north" orientation="latitude" precision="55" value="9.933">9.933</geoCoordinate>
|
||
<geoCoordinate degrees="84.0831" direction="west" orientation="longitude" precision="5" value="-84.0831">-84.0831</geoCoordinate>
|
||
6, 1156 m a.s.l. (UMM0210,
|
||
<specimenCount type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName>N Banks</collectorName>
|
||
, AMNH)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |