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<document id="E486073831450D683F8C04BB77D9EF23" ID="10.11646/zootaxa.4121.3.3" ID-CLB-Dataset="38910" ID-DOI="10.11646/zootaxa.4121.3.3" ID-GBIF-Dataset="364fbc6a-ead2-4e76-90b2-5b2c746b594c" ID-ISSN="1175-5326" ID-Zenodo-Dep="255115" ID-ZooBank="43691D85-5544-42B2-8F57-F87AC3EAD869" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465453111221" checkinUser="plazi" docAuthor="Ho, Hsuan-Ching, Kawai, Toshio &amp; Amaoka, Kunio" docDate="2016" docId="EF45FB7CC26AFFA43DC0446BFDC9F918" docLanguage="en" docName="zootaxa.4121.3.3.pdf" docOrigin="Zootaxa 4121 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bufoceratias shaoi Pietsch, Ho &amp; Chen 2004" docType="treatment" docVersion="11" lastPageNumber="276" masterDocId="137C8304C262FFAD3D57422DFFC6FF8B" masterDocTitle="Records of deep-sea anglerfishes (Lophiiformes: Ceratioidei) from Indonesia, with descriptions of three new species" masterLastPageNumber="294" masterPageNumber="267" pageNumber="275" updateTime="1698662783365" updateUser="ExternalLinkService">
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<mods:title id="25B7DB0207A9411DA0F6698D4874C08F">Records of deep-sea anglerfishes (Lophiiformes: Ceratioidei) from Indonesia, with descriptions of three new species</mods:title>
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<mods:namePart id="18E2C84480A3603D62C712BDC93A08A3">Ho, Hsuan-Ching</mods:namePart>
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<mods:namePart id="675CC2F0875B1E90E09451B5DF8FD371">Kawai, Toshio</mods:namePart>
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<mods:namePart id="68F3E035B2F3500F09064EA353F4CA8D">Amaoka, Kunio</mods:namePart>
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<mods:title id="600B603C068F99E28FC451884B078665">Zootaxa</mods:title>
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<mods:date id="19842EE798E84BAF28372E4871D65FF8">2016</mods:date>
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<treatment id="EF45FB7CC26AFFA43DC0446BFDC9F918" ID-DOI="http://doi.org/10.5281/zenodo.5617538" ID-GBIF-Taxon="120826658" ID-Zenodo-Dep="5617538" LSID="urn:lsid:plazi:treatment:EF45FB7CC26AFFA43DC0446BFDC9F918" httpUri="http://treatment.plazi.org/id/EF45FB7CC26AFFA43DC0446BFDC9F918" lastPageId="9" lastPageNumber="276" pageId="8" pageNumber="275">
<subSubSection id="2FF619E1C26AFFA53DC0446BFED4F909" pageId="8" pageNumber="275" type="nomenclature">
<paragraph id="67534A6AC26AFFA53DC0446BFD04F9EA" blockId="8.[151,706,1606,1666]" box="[151,706,1606,1633]" pageId="8" pageNumber="275">
<heading id="3C1BFD06C26AFFA53DC0446BFD04F9EA" bold="true" box="[151,706,1606,1633]" fontSize="11" level="1" pageId="8" pageNumber="275" reason="1">
<taxonomicName id="A0EC31E9C26AFFA53DC0446BFD04F9EA" ID-CoL="NPHK" authority="Pietsch, Ho &amp; Chen, 2004" authorityName="Pietsch, Ho &amp; Chen" authorityYear="2004" box="[151,706,1606,1633]" class="Actinopterygii" family="Diceratiidae" genus="Bufoceratias" kingdom="Animalia" order="Lophiiformes" pageId="8" pageNumber="275" phylum="Chordata" rank="species" species="shaoi">
<emphasis id="55989678C26AFFA53DC0446BFD04F9EA" bold="true" box="[151,706,1606,1633]" pageId="8" pageNumber="275">
<emphasis id="55989678C26AFFA53DC0446BFEBCF9EB" bold="true" box="[151,378,1606,1632]" italics="true" pageId="8" pageNumber="275">Bufoceratias shaoi</emphasis>
<bibRefCitation id="037D379BC26AFFA53CD6446AFD04F9EA" author="Pietsch" box="[385,706,1607,1633]" pageId="8" pageNumber="275" refString="Pietsch, T. W., Ho, H. - C. &amp; Chen, H. - M. (2004) Revision of the deep-sea anglerfish genus Bufoceratias Whitley (Lophiiformes: Ceratioidei: Diceratiidae), with description of a new species from the Indo-West Pacific Ocean. Copeia, 2004 (1), 98 - 107. http: // dx. doi. org / 10.1643 / CI- 03 - 166 R" type="journal article" year="2004">Pietsch, Ho &amp; Chen, 2004</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="67534A6AC26AFFA53DC04444FED4F909" blockId="8.[151,706,1606,1666]" box="[151,274,1641,1666]" pageId="8" pageNumber="275">
<figureCitation id="FFD756EFC26AFFA53DC04444FF22F909" box="[151,228,1641,1666]" captionStart="FIGURE 6. A" captionStartId="7.[151,250,1899,1921]" captionTargetBox="[378,1175,768,1859]" captionTargetId="figure@7.[378,1195,767,1884]" captionTargetPageId="7" captionText="FIGURE 6. A. Bufoceratias microcephalus, HUMZ 193747, 151.0 mm SL, holotype. B. Bufoceratias thele, HUMZ 191147, 127.4 mm SL. C. Bufoceratias shaoi, HUMZ 193746, 68.2 mm SL. D. Bufoceratias wedli, HUMZ 31934, 124.0 mm SL." httpUri="https://zenodo.org/record/255120/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
C, 8
</paragraph>
</subSubSection>
<subSubSection id="2FF619E1C26AFFA53DC0449CFA85F94C" box="[151,1347,1713,1735]" pageId="8" pageNumber="275" type="reference_group">
<paragraph id="67534A6AC26AFFA53DC0449CFA85F94C" blockId="8.[151,1347,1713,1735]" box="[151,1347,1713,1735]" pageId="8" pageNumber="275">
<treatmentCitationGroup id="47FC6D44C26AFFA53DC0449CFA85F94C" box="[151,1347,1713,1735]" pageId="8" pageNumber="275">
<taxonomicName id="A0EC31E9C26AFFA53DC0449CFD52F94C" ID-CoL="NPHK" authority="Pietsch, Ho &amp; Chen, 2004:100" authorityName="Pietsch, Ho &amp; Chen" authorityPageNumber="100" authorityYear="2004" box="[151,660,1713,1735]" class="Actinopterygii" family="Diceratiidae" genus="Bufoceratias" kingdom="Animalia" order="Lophiiformes" pageId="8" pageNumber="275" phylum="Chordata" rank="species" species="shaoi">
<emphasis id="55989678C26AFFA53DC0449CFE93F94C" box="[151,341,1713,1735]" italics="true" pageId="8" pageNumber="275">Bufoceratias shaoi</emphasis>
<treatmentCitation id="E64D6C7BC26AFFA53C0C449CFD52F94C" author="Pietsch" box="[347,660,1713,1735]" page="100" pageId="8" pageNumber="275" year="2004">
<bibRefCitation id="037D379BC26AFFA53C0C449CFDACF94C" author="Pietsch" box="[347,618,1713,1735]" pageId="8" pageNumber="275" refString="Pietsch, T. W., Ho, H. - C. &amp; Chen, H. - M. (2004) Revision of the deep-sea anglerfish genus Bufoceratias Whitley (Lophiiformes: Ceratioidei: Diceratiidae), with description of a new species from the Indo-West Pacific Ocean. Copeia, 2004 (1), 98 - 107. http: // dx. doi. org / 10.1643 / CI- 03 - 166 R" type="journal article" year="2004">Pietsch, Ho &amp; Chen, 2004</bibRefCitation>
:100
</treatmentCitation>
</taxonomicName>
, figs. 1, 3A (
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locality: northeastern
<collectingCountry id="1FFB0AFAC26AFFA53970449CFBB0F94C" box="[1063,1142,1713,1735]" name="Taiwan" pageId="8" pageNumber="275">Taiwan</collectingCountry>
).
<treatmentCitation id="E64D6C7BC26AFFA539D1449CFAF8F94C" author="Pietsch" box="[1158,1342,1713,1735]" page="358" pageId="8" pageNumber="275" year="2009">
<bibRefCitation id="037D379BC26AFFA539D1449CFAD5F94C" author="Pietsch" box="[1158,1299,1713,1735]" pageId="8" pageNumber="275" refString="Pietsch, T. W. (2009) Oceanic anglerfishes. Extraordinary diversity in the Deep Sea. University of California Press, Oakland, 557 pp." type="book" year="2009">Pietsch, 2009</bibRefCitation>
:358
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2FF619E1C26AFFA53DC044D9FBA2F8DE" pageId="8" pageNumber="275" type="materials_examined">
<paragraph id="67534A6AC26AFFA53DC044D9FBA2F8DE" blockId="8.[151,1437,1780,2021]" pageId="8" pageNumber="275">
<emphasis id="55989678C26AFFA53DC044D9FED7F886" bold="true" box="[151,273,1780,1805]" pageId="8" pageNumber="275">Materials.</emphasis>
<collectionCode id="01FDD2AFC26AFFA53C4E44D8FEAAF887" box="[281,364,1781,1804]" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="8" pageNumber="275">HUMZ</collectionCode>
193746 (1, 68.2), st. 29.,
<geoCoordinate id="02D82CADC26AFFA53FC144D8FD30F886" box="[662,758,1781,1805]" direction="south" orientation="latitude" pageId="8" pageNumber="275" precision="92" value="-8.521667">8°31.3'S</geoCoordinate>
,
<geoCoordinate id="02D82CADC26AFFA53E5344D9FC42F886" box="[772,900,1780,1805]" direction="east" orientation="longitude" pageId="8" pageNumber="275" precision="92" value="110.416664">110°25.0'E</geoCoordinate>
–
<geoCoordinate id="02D82CADC26AFFA53EC344D9FC28F886" box="[916,1006,1780,1805]" direction="south" orientation="latitude" pageId="8" pageNumber="275" precision="92" value="-8.513333">8°30.8'S</geoCoordinate>
,
<geoCoordinate id="02D82CADC26AFFA53EAC44D9FBBFF886" box="[1019,1145,1780,1805]" direction="east" orientation="longitude" pageId="8" pageNumber="275" precision="92" value="110.39333">110°23.6'E</geoCoordinate>
, off Java,
<quantity id="A014E78FC26AFFA539A744D9FAB4F887" box="[1264,1394,1780,1805]" metricMagnitude="2" metricUnit="m" metricValue="7.625" metricValueMax="7.7" metricValueMin="7.55" pageId="8" pageNumber="275" unit="m" value="762.5" valueMax="770.0" valueMin="755.0">755–770 m</quantity>
,
<date id="13526CAAC26AFFA538D744D8FECBF8BA" pageId="8" pageNumber="275" value="2005-05-13">13 May 2005</date>
.
<collectionCode id="01FDD2AFC26AFFA53C404534FEACF8BB" box="[279,362,1817,1840]" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="8" pageNumber="275">HUMZ</collectionCode>
193895 (1, 44.8),
<collectionCode id="01FDD2AFC26AFFA53F6C4534FD4BF8BB" box="[571,653,1817,1840]" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="8" pageNumber="275">HUMZ</collectionCode>
193896 (1, 52.5),
<collectionCode id="01FDD2AFC26AFFA53E094534FC76F8BB" box="[862,944,1817,1840]" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="8" pageNumber="275">HUMZ</collectionCode>
193897 (1, 46.1),
<collectionCode id="01FDD2AFC26AFFA539D74534FB14F8BB" box="[1152,1234,1817,1840]" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="8" pageNumber="275">HUMZ</collectionCode>
193898 (1, 33.5), st. 22,
<geoCoordinate id="02D82CADC26AFFA53DB54510FE85F8DE" box="[226,323,1853,1877]" direction="south" orientation="latitude" pageId="8" pageNumber="275" precision="92" value="-8.3133335">8°18.8'S</geoCoordinate>
,
<geoCoordinate id="02D82CADC26AFFA53C194510FE0CF8DE" box="[334,458,1853,1877]" direction="east" orientation="longitude" pageId="8" pageNumber="275" precision="92" value="110.21833">110°13.1'E</geoCoordinate>
–
<geoCoordinate id="02D82CADC26AFFA53CB64510FD86F8DE" box="[481,576,1853,1877]" direction="south" orientation="latitude" pageId="8" pageNumber="275" precision="92" value="-8.303333">8°18.2'S</geoCoordinate>
,
<geoCoordinate id="02D82CADC26AFFA53F1C4510FD0FF8DE" box="[587,713,1853,1877]" direction="east" orientation="longitude" pageId="8" pageNumber="275" precision="92" value="109.18833">109°11.3'E</geoCoordinate>
, off Java,
<quantity id="A014E78FC26AFFA53E6B4511FC7BF8DF" box="[828,957,1852,1877]" metricMagnitude="2" metricUnit="m" metricValue="5.56" metricValueMax="5.82" metricValueMin="5.3" pageId="8" pageNumber="275" unit="m" value="556.0" valueMax="582.0" valueMin="530.0">530–582 m</quantity>
,
<date id="13526CAAC26AFFA53E9F4510FBA7F8DE" box="[968,1121,1852,1877]" pageId="8" pageNumber="275" value="2005-05-11">11 May 2005</date>
.
</paragraph>
</subSubSection>
<subSubSection id="2FF619E1C26AFFA43D90454DFCD6FA31" lastPageId="9" lastPageNumber="276" pageId="8" pageNumber="275" type="description">
<paragraph id="67534A6AC26AFFA53D90454DFABAF86E" blockId="8.[151,1437,1780,2021]" pageId="8" pageNumber="275">
<emphasis id="55989678C26AFFA53D90454DFE9FF8F2" bold="true" box="[199,345,1888,1913]" pageId="8" pageNumber="275">Description.</emphasis>
Dorsal-fin rays 6–7; pectoral-fin rays 13–14 (mode 14); anal-fin rays 4; caudal-fin rays 9. Teeth on vomer 6–10 (3–5 on each side); upper jaw teeth 25–45; lower jaw teeth 23–35. Head length 41.8–53.4% SL; head depth 48.7–60.3; head width 30.1–41.7; frontal width 14.0–17.0; illicial length 24.8–36.7; distance between upper jaw symphysis and origin of illicium 33.1–50.6; upper jaw length 38.0–49.9; lower jaw length 41.5–50.5.
</paragraph>
<caption id="33931AE2C26BFFA43DC041CDFB5DFC7D" box="[151,1179,992,1014]" httpUri="https://zenodo.org/record/255122/files/figure.png" pageId="9" pageNumber="276" targetBox="[264,1309,199,962]" targetPageId="9">
<paragraph id="67534A6AC26BFFA43DC041CDFB5DFC7D" blockId="9.[151,1179,992,1014]" box="[151,1179,992,1014]" pageId="9" pageNumber="276">
<emphasis id="55989678C26BFFA43DC041CDFED4FC7E" bold="true" box="[151,274,992,1014]" pageId="9" pageNumber="276">FIGURE 8.</emphasis>
<taxonomicName id="A0EC31E9C26BFFA43C4E41CDFD35FC7D" authority="Pietsch, Ho &amp; Chen, 2004" authorityName="Pietsch, Ho &amp; Chen" authorityYear="2004" box="[281,755,992,1014]" class="Actinopterygii" family="Diceratiidae" genus="Buforceratias" kingdom="Animalia" order="Lophiiformes" pageId="9" pageNumber="276" phylum="Chordata" rank="species" species="shaoi">
<emphasis id="55989678C26BFFA43C4E41CDFE27FC7D" box="[281,481,992,1014]" italics="true" pageId="9" pageNumber="276">Buforceratias shaoi</emphasis>
Pietsch, Ho &amp; Chen, 2004
</taxonomicName>
, HUMZ 193746, 68.2 mm SL, preserved.
</paragraph>
</caption>
<paragraph id="67534A6AC26BFFA43D904617FD49FAF8" blockId="9.[151,1437,1082,1683]" pageId="9" pageNumber="276">Body short, globular, rounded to oval profile in lateral view. Mouth large, oblique. Illicium emerging from dorsal surface of head behind sphenotic spines. illicium relatively short. Esca small, its total length 33.9–54.8% SL and its width 4.0–5.3% SL; extremely elongate, unpigmented terminal papilla, its length 29.2–39.7% SL and width 2.0–4.5% SL, cylindrical and gradually tapering to a terminal cirrus; escal pore situated at postero-basal margin of terminal papilla; anterior escal appendage, 18.8–28.4% SL, single main branch or divided into two branches, each subdivided into several secondary branches and bearing numerous slender filaments; a pair of lateral escal appendages, 27.2–36.7% SL, each divided into four or five branches, each branch bearing numerous long, slender filaments; posterior appendage absent. Slender fang-like teeth on both jaws, longest teeth on upper jaw 5.3–7.8% SL; longest teeth on lower jaw 6.2–9.2% SL.</paragraph>
<paragraph id="67534A6AC26BFFA43D904753FCD6FA31" blockId="9.[151,1437,1082,1683]" pageId="9" pageNumber="276">Body brownish to blackish; fin rays pale grayish to black; escal appendages variously pigmented, from colorless to slightly pigmented or uniformly transparent.</paragraph>
</subSubSection>
<subSubSection id="2FF619E1C26BFFA43D9047EBFDC9F918" pageId="9" pageNumber="276" type="discussion">
<paragraph id="67534A6AC26BFFA43D9047EBFB78F9C0" blockId="9.[151,1437,1082,1683]" pageId="9" pageNumber="276">
<emphasis id="55989678C26BFFA43D9047EBFEFDFA54" bold="true" box="[199,315,1478,1503]" pageId="9" pageNumber="276">Remarks.</emphasis>
All five Indonesian specimens have nearly identical morphological characters, except for the terminal papillae that are much longer than those of
<typeStatus id="B857F4C8C26BFFA43FB447C6FCD5F988" box="[739,787,1515,1539]" pageId="9" pageNumber="276">type</typeStatus>
series and more dorsal-fin rays (6 or 7 vs. all with 5 rays, except for one
<typeStatus id="B857F4C8C26BFFA43C14443DFE65F9A3" box="[323,419,1552,1576]" pageId="9" pageNumber="276" type="paratype">paratype</typeStatus>
with 6). The escal morphology varies slightly: the
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specimen has a main branch and one additional small branch on each side; the
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specimen has a single main branch.
</paragraph>
<paragraph id="67534A6AC26BFFA43D90447BFDC9F918" blockId="9.[151,1437,1082,1683]" pageId="9" pageNumber="276">
The lack of an escal posterior appendage in all specimens also distinguishes it from all congeners but
<taxonomicName id="A0EC31E9C26BFFA438684475FA52F9E4" box="[1343,1428,1623,1647]" class="Actinopterygii" family="Diceratiidae" genus="Buforceratias" kingdom="Animalia" order="Lophiiformes" pageId="9" pageNumber="276" phylum="Chordata" rank="species" species="thele">
<emphasis id="55989678C26BFFA438684475FA52F9E4" box="[1343,1428,1623,1647]" italics="true" pageId="9" pageNumber="276">B. thele</emphasis>
</taxonomicName>
, which lacks all escal appendages.
</paragraph>
</subSubSection>
</treatment>
</document>
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