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<mods:title id="A6EA78080BE7F1ED1806F227BCF2688D">A catalogue of the ants of Paraguay (Hymenoptera: Formicidae).</mods:title>
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<treatment id="DAE1ADFACF4EF21316C2468C5272C5A7" ID-DOI="http://doi.org/10.5281/zenodo.6239959" ID-GBIF-Taxon="100131750" ID-Zenodo-Dep="6239959" LSID="urn:lsid:plazi:treatment:DAE1ADFACF4EF21316C2468C5272C5A7" httpUri="http://treatment.plazi.org/id/DAE1ADFACF4EF21316C2468C5272C5A7" lastPageNumber="52" pageNumber="52">
<subSubSection id="F7EF8572BADF9FCB76DD7D439E32593F" type="nomenclature">
<paragraph id="6E31DB7A09B327518CE66F0468BEDDFD" pageNumber="52">
<taxonomicName id="0EC51BD7AD5F386B6FFB19C32B068600" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="rochai">Crematogaster rochai Forel</taxonomicName>
1903.
</paragraph>
</subSubSection>
<subSubSection id="58E23EC8DF99E08DCF53FFA97B495E5A" type="reference_group">
<paragraph id="530AC39253DD890AEAF521F09468C0AA" pageNumber="52">
<taxonomicName id="517FA3F0669C185A5E2AAD26872EA85F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="rochai">Crematogaster rochai Forel</taxonomicName>
1903: 255.
<materialsCitation id="82FE4165B2F14798B4B977CD9EAB1C9F" ID-GBIF-Occurrence="923905315" collectionCode="MHNG" collectorName="Rocha" country="Brazil" stateProvince="Ceara" typeStatus="syntypes">
[w
<typeStatus id="5CB58D76B7D07D1403E8B46BC0871FD2">syntypes</typeStatus>
examined,
<collectionCode id="3E8BEDC075C6BD14003A77F3D25ED5CC" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
;
<normalizedToken id="E89F9CFA2E3581AC48F8F5966D20D1A2" originalValue="Ceará">
<collectingRegion id="9B1389D9AD108ACCE27D90F96776BC22">Ceara</collectingRegion>
</normalizedToken>
,
<collectingCountry id="F918F4D5C629F761184227CE0027C3E9">Brazil</collectingCountry>
, (
<collectorName id="08414DD5EEF538A33FB233D1B1868823">Rocha</collectorName>
)]
</materialsCitation>
</paragraph>
<paragraph id="E51312BE1DDCB08814BFA629F6DF6EEC" pageNumber="52">
<taxonomicName id="05E1AFE758090C51A93F9D2D8CED9859" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:149405" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster brevispinosa rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="brevispinosa" subSpecies="rochai">Crematogaster brevispinosa r. rochai Forel</taxonomicName>
. Forel 1912c: 213.
</paragraph>
<paragraph id="24E6A36D96C0AC4A2009558EF651DF1E" pageNumber="52">
<taxonomicName id="2AAA6472ED38A9A2ACD8F7E14D0C97E1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136861" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster (Orthocrema) brevispinosa st. malevolens Santschi" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="brevispinosa" subGenus="Orthocrema" subSpecies="malevolens">Crematogaster (Orthocrema) brevispinosa st. malevolens Santschi</taxonomicName>
1919: 41.
<taxonomicNameLabel id="B1A0CF3A17F84B6B91F12395AA000026">NEW SYNONYMY</taxonomicNameLabel>
.
<materialsCitation id="FC16172EDEBCFBBE148E5843AB748DE9" ID-GBIF-Occurrence="923905313" collectionCode="NHMB" collectorName="Joergensen" country="Argentina" location="Nueva Pompeya" stateProvince="Chaco" typeStatus="syntypes">
[w
<typeStatus id="755B7FC2F1D30E05A10D9E6BE39990D8">syntypes</typeStatus>
examined,
<collectionCode id="17325D442C0FE76697E2108E381AD630" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
;
<location id="4C8A1E8762335A527C4C25B27BA74041" LSID="urn:lsid:plazi:treatment:DAE1ADFACF4EF21316C2468C5272C5A7:4C8A1E8762335A527C4C25B27BA74041" country="Argentina" name="Nueva Pompeya" stateProvince="Chaco">Nueva Pompeya</location>
,
<collectingRegion id="2A71607728DD46397DD4DE45CA8A828E">Chaco</collectingRegion>
,
<collectingCountry id="18534C923A06CC6695883466734173A1">Argentina</collectingCountry>
(
<collectorName id="407AC34162F3AAF5AFFC70937416A570">Joergensen</collectorName>
)]
</materialsCitation>
.
</paragraph>
<paragraph id="61D6131723B88EF89D1543BBD6C13574" pageNumber="52">
<taxonomicName id="DC003DF769366EB8BE93C7B0730A055D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239773" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster (Orthocrema) brevispinosa subsp. rochai" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="brevispinosa" subGenus="Orthocrema" subSpecies="rochai">Crematogaster (Orthocrema) brevispinosa subsp. rochai Forel</taxonomicName>
. Emery 1924a: 134.
</paragraph>
<paragraph id="0E8A0D150AA13D1EAE5AC703E173CD6D" pageNumber="52">
<taxonomicName id="649074E9F38E8742558BED40A6E0B976" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:239773" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster (Orthocrema) brevispinosa subsp. rochai" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="brevispinosa" subGenus="Orthocrema" subSpecies="rochai">Crematogaster (Orthocrema) brevispinosa rochai Forel</taxonomicName>
. Kempf 1972: 86.
</paragraph>
<paragraph id="A58809F9237BCD152D5AC08EE249817D" pageNumber="52">
<taxonomicName id="6462878AEBE2A63BA2A285B5056E2EFC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228299" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster (Crematogaster) rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="rochai" subGenus="Crematogaster">Crematogaster (Crematogaster) rochai Forel</taxonomicName>
. Longino 2003: 102.
</paragraph>
<paragraph id="A5DFA93E15D003FED54C6F89618BFFD3" pageNumber="52">
<taxonomicName id="1732D889D7FF216F2C6EBAB4911817CE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228098" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster (Crematogaster) malevolens Santschi" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="malevolens" subGenus="Crematogaster">Crematogaster (Crematogaster) malevolens Santschi</taxonomicName>
. Longino 2003: 131. Raised to species.
</paragraph>
</subSubSection>
<subSubSection id="6DE66F12C671F2A2782E96E80788F1DF" type="diagnosis">
<paragraph id="CA078EF9249324159A74FE755008C1EC" pageNumber="52">
<taxonomicName id="338DD1F842E4FB35EF51DD40C44C3852" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="rochai">Crematogaster rochai</taxonomicName>
can be diagnosed by the following combination of characters: antennal scapes short, less than.8x head length, in repose failing to reach posterior margin of head in full-face view; dorsal surface of head largely devoid of erect setae; promesonotum slightly arched in profile; propodeal suture present but not deep; postero-dorsal face of propodeum forming a single concave slope, not broken into distinct dorsal and posterior faces; propodeal spines short and upturned; first gastric tergite (= abd. tergite 4) with &lt;6 standing setae exclusive of posterior row.
</paragraph>
</subSubSection>
<subSubSection id="8C0F5074D11AA90CEE392824C45857FB" type="discussion">
<paragraph id="8DB3C1674DD5E014BCF1C5ADB84F5023" pageNumber="52">
Mature colonies in the
<taxonomicName id="B06CB437FCCFBE24AD52694465498AC2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28145" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster crinosa Mayr" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="crinosa">Crematogaster crinosa</taxonomicName>
complex, including
<taxonomicName id="0E73256C0C8F5B8DDEC3C1B35E28608F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="rochai">C. rochai</taxonomicName>
, are polymorphic in the worker caste and occasionally produce rather large workers. Santschis
<taxonomicName id="A108EF4AD4A590587C7A9C13997523B4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136861" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster (Orthocrema) brevispinosa st. malevolens Santschi" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="brevispinosa" subGenus="Orthocrema" subSpecies="malevolens">C. malevolens</taxonomicName>
seems to be little other than a large
<taxonomicName id="6E77D7515566C09E5189DC20753F721B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="rochai">C. rochai</taxonomicName>
. I can find no consistent characters to separate
<taxonomicName id="8263399F6230C115ABA06AE9B1FE006A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136861" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster (Orthocrema) brevispinosa st. malevolens Santschi" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="brevispinosa" subGenus="Orthocrema" subSpecies="malevolens">C. malevolens</taxonomicName>
from the
<taxonomicName id="C94D933A1F2C24B4C5A37CEE2D7E4E96" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="52" phylum="Arthropoda" rank="species" species="rochai">C. rochai</taxonomicName>
type or from Paraguayan material that Jack Longino has identified as
<taxonomicName id="286B6C35BB9D847D32F9D09A66C4F729" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="rochai">C. rochai</taxonomicName>
. Longino (2003) elevated
<taxonomicName id="6AB662C891A6927204A2C62614038A1C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:183077" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster malevolens Santschi" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="malevolens">C. malevolens</taxonomicName>
to species with the intent to clear trinomials from the
<taxonomicName id="DC4A583914A208A5A11671DC53912464" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28145" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster crinosa Mayr" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="crinosa">C. crinosa</taxonomicName>
complex but did not provide additional reasoning. If
<taxonomicName id="ACFA00BD36F7C57187DB09C140D63517" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:183077" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster malevolens Santschi" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="malevolans">C. malevolans</taxonomicName>
is indeed conspecific with what I call
<taxonomicName id="733957293071A5D879F24FA3F652213E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="rochai">C. rochai</taxonomicName>
in Paraguay, an alternate resolution would be to split these southern populations off from
<taxonomicName id="6837B59BCE1CEF01BFF5D8884595B743" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:136868" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster rochai Forel" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="rochai">C. rochai</taxonomicName>
under the name
<taxonomicName id="365193F6B4A857DD87075288AE45A30A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:183077" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster malevolens Santschi" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="malevolens">C. malevolens</taxonomicName>
. Specimens from southern South America are more pilose and bear a stronger propodeal suture than more northerly material Longino (2003), character states that apply equally to
<taxonomicName id="72B82D4B3E5203ED8D7F29FC5D2853AB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:183077" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Crematogaster malevolens Santschi" order="Hymenoptera" pageNumber="53" phylum="Arthropoda" rank="species" species="malevolens">C. malevolens</taxonomicName>
. Considering the allopatric nature of the variation, however, I prefer to retain a single species.
</paragraph>
</subSubSection>
</treatment>
</document>