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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.885.38655" ID-GBIF-Dataset="8e14d288-ddb5-485a-bd43-0cd3f46754ff" ID-PMC="PMC6848236" ID-Pensoft-Pub="1313-2970-885-51" ID-Pensoft-UUID="BCEA8CB1014D5D329A2E4B2CF99881FF" ID-PubMed="31736618" ID-ZooBank="BA152050AF7344CA8CED6D30F963CBC9" ModsDocID="1313-2970-885-51" checkinTime="1572912880629" checkinUser="pensoft" docAuthor="Zlatkov, Boyan &amp; Huemer, Peter" docDate="2019" docId="C929F10354A05685B877430BB5355771" docLanguage="en" docName="ZooKeys 885: 51-87" docOrigin="ZooKeys 885" docSource="http://dx.doi.org/10.3897/zookeys.885.38655" docTitle="Clepsis consimilana Huebner 1817" docType="treatment" docVersion="4" id="BCEA8CB1014D5D329A2E4B2CF99881FF" lastPageNumber="51" masterDocId="BCEA8CB1014D5D329A2E4B2CF99881FF" masterDocTitle="Remarkable confusion in some Western Palearctic Clepsis leads to a revised taxonomic concept (Lepidoptera, Tortricidae)" masterLastPageNumber="87" masterPageNumber="51" pageNumber="51" updateTime="1668168009670" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Remarkable confusion in some Western Palearctic Clepsis leads to a revised taxonomic concept (Lepidoptera, Tortricidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zlatkov, Boyan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huemer, Peter</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>885</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>87</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.885.38655</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.885.38655</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-885-51</mods:identifier>
<mods:identifier type="ZooBank">BA152050AF7344CA8CED6D30F963CBC9</mods:identifier>
<mods:identifier type="Pensoft-UUID">BCEA8CB1014D5D329A2E4B2CF99881FF</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="160133792" LSID="urn:lsid:plazi:treatment:C929F10354A05685B877430BB5355771" httpUri="http://treatment.plazi.org/id/C929F10354A05685B877430BB5355771" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
<paragraph pageId="0" pageNumber="51">
<taxonomicName LSID="C929F103-54A0-5685-B877-430BB5355771" authority="Huebner, 1817" authorityName="Huebner" authorityYear="1817" class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
Clepsis consimilana (
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
, 1817)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="reference_group">
<paragraph pageId="0" pageNumber="51">
[
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="genus">Tortrix</taxonomicName>
]
<taxonomicName family="Tortricidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="consimilana" order="Lepidoptera" pageId="0" pageNumber="51" rank="species" species="consimilana">consimilana</taxonomicName>
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
, 1817: pl. 38, fig. 239 (Europe).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix unifasciana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="unifasciana">Tortrix unifasciana</taxonomicName>
Duponchel, 1842: 135, pl.61, fig. 6 (France).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" form="cinnamomeana" genus="Tortrix" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Animalia" lsidName="Tortrix unifasciana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="form" species="unifasciana">Tortrix unifasciana f. cinnamomeana</taxonomicName>
Dufrane, 1957: 6 (Belgium).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix fallaciana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="fallaciana">Tortrix fallaciana</taxonomicName>
Fuchs, 1903: 4 (Sicily).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" form="fuscana" genus="Tortrix" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Animalia" lsidName="Tortrix unifasciana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="form" species="unifasciana">Tortrix unifasciana f. fuscana</taxonomicName>
Dufrane, 1957: 6 (Belgium).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" form="obliterana" genus="Tortrix" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Animalia" lsidName="Tortrix unifasciana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="form" species="unifasciana">Tortrix unifasciana f. obliterana</taxonomicName>
Dufrane, 1957: 6 (Belgium).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" genus="Paramesia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramesia peregrinana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="peregrinana">Paramesia peregrinana</taxonomicName>
Stephens, 1852: 90 (Britain), unavailable, published in synonymy with
<taxonomicName class="Insecta" family="Tortricidae" genus="Lozotaenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lozotaenia obliquana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="obliquana">Lozotaenia obliquana</taxonomicName>
Humphreys and Westwood 1845.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix productana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="productana">Tortrix productana</taxonomicName>
Zeller, 1847: 660 (Sicily).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName family="Tortricidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="obliterana" order="Lepidoptera" pageId="0" pageNumber="51" rank="species" species="obliterana">obliterana</taxonomicName>
<normalizedToken originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
, 1847 (uninominal): pl. 9, fig. 60.
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix (Lozotaenia) obliterana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="obliterana" subGenus="Lozotaenia">Tortrix (Lozotaenia) obliterana</taxonomicName>
<normalizedToken originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
, 1851: 164 (Italy).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<taxonomicName class="Insecta" family="Tortricidae" genus="Siclobola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siclobola placida" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="placida">Siclobola placida</taxonomicName>
Diakonoff, 1948: 25 (Madagascar).
</paragraph>
<paragraph pageId="0" pageNumber="51">
non
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix eatoniana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="eatoniana">Tortrix eatoniana</taxonomicName>
Ragonot, 1881
</paragraph>
<paragraph pageId="0" pageNumber="51">
non
<taxonomicName class="Insecta" family="Tortricidae" genus="Cacoecia" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Cacoecia unifasciana var. semiana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="variety" species="unifasciana" variety="semiana">Cacoecia unifasciana var. semiana</taxonomicName>
<normalizedToken originalValue="Chrétien">Chretien</normalizedToken>
, 1915
</paragraph>
<paragraph pageId="0" pageNumber="51">
non
<taxonomicName class="Insecta" family="Tortricidae" genus="Cacoecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cacoecia acclivana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="acclivana">Cacoecia acclivana</taxonomicName>
Zerny, 1933
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="materials_examined">
<paragraph pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="51">
Neotype ♂ (here designated): pinned, with 5 labels: &quot;Gehegs / Dresden / 18.vi.1922 / E.
<normalizedToken originalValue="Möbius”">Moebius&quot;</normalizedToken>
&quot;Coll. E.
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
/ Ankauf 1946&quot; &quot;Neotype /
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix consimilana</emphasis>
/
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
, 1817 / Zlatkov &amp; Huemer, 2019 des.&quot; [red printed] &quot;Museum
<normalizedToken originalValue="für">fuer</normalizedToken>
Tierkunde Dresden / Gen. prep. / ♂ / No. 2/30.11.2018&quot; &quot;
<emphasis italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
/ (
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
, 1817) / Zlatkov &amp; Huemer, 2019 det.&quot;; genitalia on a slide with two labels: &quot;Neotype /
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix consimilana</emphasis>
/
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
, 1817 / [Germany] Dresden, Gehegs / 18.vi.1922, E.
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
/ Zlatkov &amp; Huemer, 2019 des.&quot; [red printed] &quot;Museum
<normalizedToken originalValue="für">fuer</normalizedToken>
Tierkunde Dresden / Gen. Prep. / ♂ / No. 2/30.11.2018 / B. Zlatkov 2018 Euparal&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="51">
GERMANY • 1 ♂; Dresden, Gehegs; 18 Jun. 1922; E.
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
leg.; GS 2/30.11.2018; MTD.
</paragraph>
<paragraph pageId="0" pageNumber="51">
Lectotype ♂ of
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix unifasciana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="unifasciana">
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix unifasciana</emphasis>
</taxonomicName>
(here designated): pinned, with 5 labels:
<normalizedToken originalValue="“unifasciana”">&quot;unifasciana&quot;</normalizedToken>
[handwritten]
<normalizedToken originalValue="“Duponch[el]”">&quot;Duponch[el]&quot;</normalizedToken>
[round, handwritten]
<normalizedToken originalValue="“713”">&quot;713&quot;</normalizedToken>
[round handwritten]
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
[red printed] &quot;Chr. Gibeaux
<normalizedToken originalValue="dét">det</normalizedToken>
. /
<normalizedToken originalValue="prép">prep</normalizedToken>
.
<normalizedToken originalValue="génit">genit</normalizedToken>
. n° 4231 ♂ / unifasciana / Dup.&quot; [printed and handwritten]; male genitalia on a slide with two labels: &quot;Tortrix / unifasciana / Dup., 1842 / Type&quot; [handwritten with red border] &quot;Chr. Gibeaux
<normalizedToken originalValue="dét">det</normalizedToken>
. /
<normalizedToken originalValue="prép">prep</normalizedToken>
.
<normalizedToken originalValue="génit">genit</normalizedToken>
. n° 4231 ♂ / Clepsis / consimilana Hb. / 30.x.91 Euparal M. P.&quot; [printed and handwritten].
</paragraph>
<paragraph pageId="0" pageNumber="51">FRANCE • 1 ♂; GS 4231; MNHN.</paragraph>
<paragraph pageId="0" pageNumber="51">
Holotype ♂ of
<taxonomicName class="Insecta" family="Tortricidae" genus="Siclobola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siclobola placida" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="placida">
<emphasis italics="true" pageId="0" pageNumber="51">Siclobola placida</emphasis>
</taxonomicName>
: pinned, with four labels:
<normalizedToken originalValue="“Tananarive”">&quot;Tananarive&quot;</normalizedToken>
&quot;Type: ♂ / Siclobola / placida / A. Diakonoff 1946&quot; [printed and handwritten]
<normalizedToken originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
[printed red] &quot;Siclobola / unifasciana / placida / 1959 Diak. / det. A. Diakonoff&quot; [printed and handwritten]; genitalia on glass slide pinned to the holotype, with one label:
<normalizedToken originalValue="“536.D.”">&quot;536.D.&quot;</normalizedToken>
</paragraph>
<paragraph pageId="0" pageNumber="51">MADAGASCAR • 1 ♂; Antananarivo; MNHN 536.D.</paragraph>
<paragraph pageId="0" pageNumber="51">
Other material: GERMANY • 1 ♂; Dresden, Gehegs; 24 Jun. 1921; E.
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
leg.; GS 1/30.11.2018; MTD • 1♂; Dresden,
<normalizedToken originalValue="Lössaltz">Loessaltz</normalizedToken>
; 28 Jun. 1935; E.
<normalizedToken originalValue="Möbius">Moebius</normalizedToken>
leg.; GS 2/3.12.2018; MTD • 1 ♂; same collection data; 3 Aug. 1935; GS 1/3.12.2018; MTD • 1 ♀; Fritzsche leg.; GS 1/4.12.2018; MTD • 1 ♂; Sylt; GS 2/5.11.2018; ZSM • 2 ♂♂, 2 ♀♀; Baden
<normalizedToken originalValue="Württemberg">Wuerttemberg</normalizedToken>
, Marbach am Neckar; 29 Jun. 1957; L.
<normalizedToken originalValue="Süssner">Suessner</normalizedToken>
leg.; TLMF • 1 ♂; Bayern,
<normalizedToken originalValue="München">Muenchen</normalizedToken>
, Hochmutting; 19 Aug. 1988; H. Kolbeck leg.; TLMF • 1 ♀; Kehlheim; 12 Jun. 1997; H. Kolbeck leg.; TLMF • ITALY • 1 ♂; prov. Verona, Monte; alt. 300 m; 20 Aug. 1994; Franz leg.; GS 1/16.10.2017; TLMF • 1 ♂, same collection data; 7 Jun. 1991; K. Burmann et al. leg.; GS 2/16.10.2017; TLMF • 2 ♀♀; same collection data; 1 Oct. 1994; Burmann et Erlebach leg.; GS 3/16.10.2017, 4/16.10.2017; TLMF • 1 ♂; prov. Verona, Albisano; alt. 450 m; 27
<normalizedToken originalValue="May">May-</normalizedToken>
3 Jun. 1962; K. Burmann leg.; TLMF • 1 ♂; prov. Trentino, Pietramurata; alt. 250 m; mid Jun. 1978; F.
<normalizedToken originalValue="Zürnbauer">Zuernbauer</normalizedToken>
leg.; TLMF • 1 ♀; Verona, Mte. Maderno; alt. 250 m; mid Sep. 1963; K. Burmann leg.; TLMF • 1 ♀;
<normalizedToken originalValue="Südtirol">Suedtirol</normalizedToken>
, Auer; 21 Jun. 1957; Hernegger leg.; TLMF • 3 ♂♂, 1 ♀;
<normalizedToken originalValue="Südtirol">Suedtirol</normalizedToken>
, Eing. Schnalstal; alt. 700 m; mid Jun.1968; K. Burmann leg.; TLMF • 2 ♀♀;
<normalizedToken originalValue="Südtirol">Suedtirol</normalizedToken>
, S Kalterer See; alt. 230 m; 26 Jun. 1990; P. Huemer leg.; TLMF • 1 ♂; same collection data; 14 Jun. 1991; TLMF • 2 ♂♂;
<normalizedToken originalValue="Südtirol">Suedtirol</normalizedToken>
, Umg. Bozen, Kampenn; 26 Jun. 1991; P. Huemer leg.; TLMF • 1 ♂; Carrara; alt. 200 m; early Jun. 1979; F.
<normalizedToken originalValue="Zürnbauer">Zuernbauer</normalizedToken>
leg.; TLMF • 1 ♂, 1 ♀; Elba, Porto Ferroio; alt. 40 m; late May 1967; F.
<normalizedToken originalValue="Zürnbauer">Zuernbauer</normalizedToken>
leg.; TLMF • 1 ♂; Sardinia, Umg. Tempio, Mte. Limbara; alt. 1200 m; 27 May1973; Laubmeier, Sommerer &amp; Witt leg.; TLMF • 1 ♂; Sardinia, Umg. Aritzo, Gennargentu; alt. 750 m; 4-5 Jun. 1973; Laubmeier, Sommerer &amp; Witt leg.; TLMF • POLAND • 1 ♂;
<normalizedToken originalValue="Poznań">Poznan</normalizedToken>
, Ogrody; 13 Jun. 2012; W. Kubasik leg; GS 1/15.3.2019; • CWK 1♂;
<normalizedToken originalValue="Poznań">Poznan</normalizedToken>
, Ogrody; 19 Jun. 2012; W. Kubasik leg.; GS 2/15.3.2019; CWK • 1 ♀; Komorniki; 19 Jun. 2002; GS 1/15.3.2002; W. Kubasuk leg.; CWK • AUSTRIA • 2 ♂♂; Wien,
<normalizedToken originalValue="Döbling">Doebling</normalizedToken>
, Cobenzl; alt. 385 m; 15 Jun. 2012; F. Lichtenberger leg.; TLMF • 3 ♂♂;
<normalizedToken originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, 10 km SE Schwechat, Seiherwiese; 1 Jul. 1998; F. Lichtenberger leg.; TLMF • 3 ♂♂;
<normalizedToken originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, Ebreichsdorf, Welscherhalten; 2 Jul. 1999, F. Lichtenberger leg.; TLMF • 1 ♂;
<normalizedToken originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, Rubring; 20 Jun. 1998; F. Hofmann leg.; TLMF • 1 ♂;
<normalizedToken originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, Amstetten, Sonnleiten; 16 Jun. 1997; H. Brandstetter leg.; TLMF • 1 ♂;
<normalizedToken originalValue="Niederösterreich">Niederoesterreich</normalizedToken>
, Purgstall/Erlauf; 7 Jul. 1995; F. Ressl leg.; TLMF • 1 ♂;
<normalizedToken originalValue="Oberösterreich">Oberoesterreich</normalizedToken>
, Linz, Biologiezentrum; 11 Jun. 2002; J. Wimmer leg.; GS 3771; TLMF • 2 ♂♂, 2 ♀♀; Vorarlberg, Hard,
<normalizedToken originalValue="Fußach">Fussach</normalizedToken>
, NSG Rheindelta-Rohrspitz; alt. 397 m; 29 Jun. 1992; P. Huemer leg.; TLMF • FRANCE • 1 ♂; Alpes Maritimes, Nice, Bd. Tzarewitsch; 26 May 1971; F. Dujardin leg.; TLMF • 1 ♂; Alpes Martitimes, mt. Alban; alt. 200 m; 4 Jun. 1962; F. Dujardin leg.; TLMF • 1 ♂; Bouches du Rhone, La Ciotat; 4 May 1994; J. Nel leg.; GS 9763 TLMF • 1 ♂; Var, Massif du Pradet; J. Nel leg.; TLMF • 1 ♂; Vaucluse, Mormoiron; alt. 20 m; 6 May 2000; J. Nel leg.; TLMF • 2 ♂♂; Corse, Pinarello; alt. 10 m; early Jun. 1972; F.
<normalizedToken originalValue="Zürnbauer">Zuernbauer</normalizedToken>
leg.; TLMF • CROATIA • 2 ♂♂, 1 ♀; Rovinij; alt. 50 m; 10 Sep. 2002; H. Deutsch leg.; TLMF • 1 ♂; Cres isl., Stivian; 14-17 May 1996; F. Lichtenberger leg.; TLMF • GREECE • 1 ♂; Crete, Heraklion prefecture, Fodele; alt. 100 m; 25 May 2000; W. Ruckdeschel leg.; GS 5/16.10.2017; TLMF • BULGARIA • 1 ♂; Petrich district, Rupite area; 41.4620°N, 23.2583°E; alt. 200 m; 15 May 2007; B. Zlatkov leg.; GS 6/16.10.2017; NMNHS • ALBANIA • 1 ♂;
<normalizedToken originalValue="Korça">Korca</normalizedToken>
Region, Boboshtica Village; 40.5405°N, 20.7918°E; alt. 1220 m; 4 Jun. 2018; S. Beshkov leg.; GS 1/17.10.2018; NMNHS • MACEDONIA (Republic of Northern) • 1 ♂; Vardar River Valley, Demir Kapiya; 41.3826°N, 22.1958°E; alt. 240 m; 10 Jun. 2018; S. Beshkov &amp; A. Nahirnic leg.; GS 2/17.10.2018; NMNHS • SPAIN • 4 ♂♂, 3 ♀♀; Catalonia, Vidreras; 6-15 Jun. 1993; J. Wimmer leg.; TLMF.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="diagnosis">
<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="51">
The wing pattern of males resembles that of
<taxonomicName lsidName="C. neglectana" pageId="0" pageNumber="51" rank="species" species="neglectana">
<emphasis italics="true" pageId="0" pageNumber="51">C. neglectana</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
. The male genitalia are most similar to
<taxonomicName lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
but the phallic process is smaller, the phallus lacks a keel, the cornuti are smaller, the valvae are more slender, and the labis is different. The females of
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
externally are very similar but the genitalia (colliculum) show considerable difference, it is much larger in
<taxonomicName lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
, with a long process at the left side.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">
Adult. Sexual dimorphism prominent. Male (
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Adults of Clepsis consimilana species group A-H C. consimilana: A illustration from the original description of Tortrix consimilana B neotype of T. consimilana C male, Bulgaria D lectotype of T. unifasciana E Holotype of Siclobola placida F male, Poland G female, Italy H female, Poland I-L C. eatoniana: I Lectotype of T. eatoniana J male, Spain K Lectotype of Tortrix xylotoma (courtesy by D. Lees and Trustees of the Natural History Museum, London) L female, Spain. Scale bar: 5 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure11" httpUri="https://binary.pensoft.net/fig/352389" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure11">
Fig. 11
<normalizedToken originalValue="AF">A-F</normalizedToken>
</figureCitation>
). Head. Vertex, frons and labial palps fulvous. Antennae with scapus and pedicellus ochreous, flagellum with fulvous brown-tipped scales and numerous sensilla trichodea as long as width of flagellomeres. Thorax. Dorsally fulvous, ventrally creamy, legs brownish. Tegula fulvous. Forewing relatively wide, with length 6.3-7.8 mm (mean 7.0,
<emphasis italics="true" pageId="0" pageNumber="51">N</emphasis>
= 13). Costa basally convex, apically straight, costal fold extending from base to ca. 0.4-0.6 of costa (
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Costal folds of males of Clepsis spp. A C. neglectana, Germany B C. striolana, Austria C C. trivia, Crete D C. acclivana, lectotype of Cacoecia acclivana E C. consimilana, neotype of Tortrix consimilana F C. eatoniana, Spain. Scale bar: 1 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure3" httpUri="https://binary.pensoft.net/fig/352381" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure3">Fig. 3E</figureCitation>
). Upperside background dark yellow with rusty reticulate pattern. Markings fulvous to brown: basal blotch atrophied, more prominent at dorsum forming dark dot; median fascia often with darker borders; subapical blotch triangular, often ill-defined. Cilia concolourous with background. Underside pale brown with creamy longitudinal blotch in the distal half of costal area. Hindwings upperside monochrome grey with paler cilia, underside with creamy costal half pale grey with scattered pale brown scales and monochrome pale grey dorsal half. Forewings with variable colouration, darker or paler, sometimes with reduced markings similarly to female (
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Adults of Clepsis consimilana species group A-H C. consimilana: A illustration from the original description of Tortrix consimilana B neotype of T. consimilana C male, Bulgaria D lectotype of T. unifasciana E Holotype of Siclobola placida F male, Poland G female, Italy H female, Poland I-L C. eatoniana: I Lectotype of T. eatoniana J male, Spain K Lectotype of Tortrix xylotoma (courtesy by D. Lees and Trustees of the Natural History Museum, London) L female, Spain. Scale bar: 5 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure11" httpUri="https://binary.pensoft.net/fig/352389" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure11">Fig. 11F</figureCitation>
). Abdomen. Grey. Male genitalia (
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Male genitalia of the Clepsis consimilana species group (some without phallus) A-E C. consimilana: A Tortrix consimilana neotype B C. consimilana, Italy C C. consimilana, Albania D Siclobola placida holotype E T. unifasciana lectotype F T. eatoniana lectotype. Scale bar: 500 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure12" httpUri="https://binary.pensoft.net/fig/352390" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure12">
Fig. 12
<normalizedToken originalValue="AE">A-E</normalizedToken>
</figureCitation>
). Uncus with variable shape and median incision depending on preparation, more or less ovoid, widened distally, rounded. Gnathos plough-shaped. Socius small, membranous. Valvae pointed dorsolaterally when mounted on slide. Costal sclerite protruded medially into large triangular labis with elongated tip (
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Labides of Clepsis spp. A, B C. neglectana: A Southern Germany B Finland C C. striolana, Austria, North Tyrol D C. trivia, Crete E C. acclivana, paralectotype of Cacoecia acclivana F, G C. consimilana: F Germany, neotype of Tortrix consimilana G Italy H C. eatoniana, Spain. Scale bar: 100 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure5" httpUri="https://binary.pensoft.net/fig/352383" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure5">Fig. 5F, G</figureCitation>
). Apical part of sacculus ca. 1.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than basal part, both forming angle of 160-165°, saccular process small, curved, thorn-shaped. Distal part of valva membranous, widened apically and protruding into small brachiola, costal edge slightly convex, with longitudinal fold on the median surface bearing row of 5-8 large modified setae. Phallus smooth, coecum medially concave, basal part curved ventrad at ca. 140°, distal part smoothly curved dorsad, with small sharp tipped lateral process as long as 0.26
<normalizedToken originalValue="×">x</normalizedToken>
distance between anterior opening and tip of phallus, slightly curved laterally and not overpassing tip of phallus. Caulis small, adjoining coecum. Vesica cylindrical, curved at angle of 100-120° to phallus, with expansion basally and finger-like sinistrodorsal diverticulum (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Phalli with vesicae everted of the Clepsis consimilana species group A-B C. consimilana: A neotype of Tortrix consimilana B specimen from Italy C C. eatoniana, Spain. The phallus of each species is shown in three aspects from top to bottom: sclerotised phallic tube (without vesica) in dorsal view, tip of vesica in dorsal view and whole phallus in left view. Scale bar: 250 μm." figureDoi="10.3897/zookeys.885.38655.figure13" httpUri="https://binary.pensoft.net/fig/352391" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure13">Fig. 13A, B</figureCitation>
). Three to four deciduous aciculate, robust, weakly sinuate, slightly flattened cornuti attached ventroapically (
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Specialised male setae in Clepsis spp. A-E cornuti: A C. neglectana, Finland B C. striolana, Austria C C. trivia, Crete D C. consimilana, neotype of Tortrix consimilana E C. eatoniana, Spain F modified large seta from the medial surface of valva of C. consimilana, Germany. Scale bar: 100 μm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure7" httpUri="https://binary.pensoft.net/fig/352385" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure7">Fig. 7D</figureCitation>
). Gonopore located at left to cornuti sockets and diverticulum. Female darker than male, with uniform forewings (
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Adults of Clepsis consimilana species group A-H C. consimilana: A illustration from the original description of Tortrix consimilana B neotype of T. consimilana C male, Bulgaria D lectotype of T. unifasciana E Holotype of Siclobola placida F male, Poland G female, Italy H female, Poland I-L C. eatoniana: I Lectotype of T. eatoniana J male, Spain K Lectotype of Tortrix xylotoma (courtesy by D. Lees and Trustees of the Natural History Museum, London) L female, Spain. Scale bar: 5 mm, all to scale." figureDoi="10.3897/zookeys.885.38655.figure11" httpUri="https://binary.pensoft.net/fig/352389" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure11">Fig. 11G, H</figureCitation>
). Head. Frons, vertex and labial palps fulvous to ochreous, antennae with fulvous brown-tipped scales and sparse sensilla trichodea shorter than width of flagellomeres. Thorax dorsally ochreous, ventrally creamy, legs brownish. Forewing length 6.3-7.6 mm (mean 6.8,
<emphasis italics="true" pageId="0" pageNumber="51">N</emphasis>
= 3). Upperside ground colour ochreous to rusty with darker reticulated pattern and more or less atrophied ill-defined markings. Basal blotch reduced to dark spot at dorsum, median fascia more or less prominent in costal and dorsal area or completely reduced. Cilia paler than background. Underside pale brown, costal and terminal areas paler or creamy with reticulate pattern. Hindwing upperside pale grey, underside with whitish or pale grey reticulate patterned costal half and monochrome pale grey dorsal half. Abdomen. Grey. Female genitalia with papillae anales not modified (
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Female genitalia of the Clepsis consimilana species group A C. consimilana, Italy B C. eatoniana, Spain. The sterigma and colliculum are shown enlarged to the right of the whole genitalia. Scale bars: 1 mm (whole genitalia), 250 μm (sterigma and colliculum)." figureDoi="10.3897/zookeys.885.38655.figure14" httpUri="https://binary.pensoft.net/fig/352392" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure14">Fig. 14A</figureCitation>
). Apophyses anteriores 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than apophyses posteriores. Sterigma widened caudad, with lateral sclerotised pockets cephalad, large excavation on the dorsal wall and wide v-shaped lamella antevaginalis. Colliculum large, with length 0.17
<normalizedToken originalValue="×">x</normalizedToken>
length of ductus bursae, slightly bent to left, funnel-shaped, with plicate longitudinal sclerotisation forming small process at left, and spherical protrusion at cranial end consisting of colourless thick cuticle. Ductus bursae long and narrow, emerging at left side of cuticular protrusion, with cestum extending along cranial 0.8
<normalizedToken originalValue="×">x</normalizedToken>
of its length. Ductus seminalis inserted dorsally at caudal end of ductus bursae. Corpus bursae ovoid, with large falcate signum with capitulum and flat signum located near end of cestum consisting of sclerotised papillae (
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Signa of bursa copulatrix in Clepsis spp. A C. neglectana B C. striolana C C. trivia D C. consimilana E C. eatoniana F A detail of plate shaped signum of C. consimilana consisting of sclerotised papillae. 1 falcate signum, 2 plate shaped signum, 3 cestum. Scale bars: 250 μm (A-E, all to scale), 25 μm (F)." figureDoi="10.3897/zookeys.885.38655.figure9" httpUri="https://binary.pensoft.net/fig/352387" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure9">Fig. 9D, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.885.38655.figure11" httpUri="https://binary.pensoft.net/fig/352389" pageId="0" pageNumber="51" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 11.</emphasis>
Adults of
<taxonomicName class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
</taxonomicName>
species group
<emphasis bold="true" pageId="0" pageNumber="51">
<normalizedToken originalValue="AH">A-H</normalizedToken>
</emphasis>
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
illustration from the original description of
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix consimilana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
neotype of
<taxonomicName lsidName="T. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">T. consimilana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
male, Bulgaria
<emphasis bold="true" pageId="0" pageNumber="51">D</emphasis>
lectotype of
<taxonomicName lsidName="T. unifasciana" pageId="0" pageNumber="51" rank="species" species="unifasciana">
<emphasis italics="true" pageId="0" pageNumber="51">T. unifasciana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="51">E</emphasis>
Holotype of
<taxonomicName class="Insecta" family="Tortricidae" genus="Siclobola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siclobola placida" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="placida">
<emphasis italics="true" pageId="0" pageNumber="51">Siclobola placida</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="51">F</emphasis>
male, Poland
<emphasis bold="true" pageId="0" pageNumber="51">G</emphasis>
female, Italy
<emphasis bold="true" pageId="0" pageNumber="51">H</emphasis>
female, Poland
<emphasis bold="true" pageId="0" pageNumber="51">
<normalizedToken originalValue="IL">I-L</normalizedToken>
</emphasis>
<taxonomicName lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="51">I</emphasis>
Lectotype of
<taxonomicName lsidName="T. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">T. eatoniana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="51">J</emphasis>
male, Spain
<emphasis bold="true" pageId="0" pageNumber="51">K</emphasis>
Lectotype of
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix xylotoma" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="xylotoma">
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix xylotoma</emphasis>
</taxonomicName>
(courtesy by D. Lees and Trustees of the Natural History Museum, London)
<emphasis bold="true" pageId="0" pageNumber="51">L</emphasis>
female, Spain. Scale bar: 5 mm, all to scale.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.885.38655.figure12" httpUri="https://binary.pensoft.net/fig/352390" pageId="0" pageNumber="51" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 12.</emphasis>
Male genitalia of the
<taxonomicName class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
</taxonomicName>
species group (some without phallus)
<emphasis bold="true" pageId="0" pageNumber="51">
<normalizedToken originalValue="AE">A-E</normalizedToken>
</emphasis>
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix consimilana</emphasis>
</taxonomicName>
neotype
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
, Italy
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
, Albania
<emphasis bold="true" pageId="0" pageNumber="51">D</emphasis>
<taxonomicName class="Insecta" family="Tortricidae" genus="Siclobola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siclobola placida" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="placida">
<emphasis italics="true" pageId="0" pageNumber="51">Siclobola placida</emphasis>
</taxonomicName>
holotype
<emphasis bold="true" pageId="0" pageNumber="51">E</emphasis>
<taxonomicName lsidName="T. unifasciana" pageId="0" pageNumber="51" rank="species" species="unifasciana">
<emphasis italics="true" pageId="0" pageNumber="51">T. unifasciana</emphasis>
</taxonomicName>
lectotype
<emphasis bold="true" pageId="0" pageNumber="51">F</emphasis>
<taxonomicName lsidName="T. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">T. eatoniana</emphasis>
</taxonomicName>
lectotype. Scale bar: 500
<normalizedToken originalValue="μm">μm</normalizedToken>
, all to scale.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.885.38655.figure13" httpUri="https://binary.pensoft.net/fig/352391" pageId="0" pageNumber="51" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 13.</emphasis>
Phalli with vesicae everted of the
<taxonomicName class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
</taxonomicName>
species group
<emphasis bold="true" pageId="0" pageNumber="51">
<normalizedToken originalValue="AB">A-B</normalizedToken>
</emphasis>
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
:
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
neotype of
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix consimilana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
specimen from Italy
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
<taxonomicName lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
, Spain. The phallus of each species is shown in three aspects from top to bottom: sclerotised phallic tube (without vesica) in dorsal view, tip of vesica in dorsal view and whole phallus in left view. Scale bar: 250
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.885.38655.figure14" httpUri="https://binary.pensoft.net/fig/352392" pageId="0" pageNumber="51" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 14.</emphasis>
Female genitalia of the
<taxonomicName class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
</taxonomicName>
species group
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
, Italy
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
<taxonomicName lsidName="C. eatoniana" pageId="0" pageNumber="51" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">C. eatoniana</emphasis>
</taxonomicName>
, Spain. The sterigma and colliculum are shown enlarged to the right of the whole genitalia. Scale bars: 1 mm (whole genitalia), 250
<normalizedToken originalValue="μm">μm</normalizedToken>
(sterigma and colliculum).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="preimaginal stages">
<paragraph pageId="0" pageNumber="51">Preimaginal stages.</paragraph>
<paragraph pageId="0" pageNumber="51">
Detailed descriptions of the ovum, larvae of all instars, and pupa are provided by
<bibRefCitation author="Sheldon, WG" journalOrPublisher="Entomologist" pageId="0" pageNumber="51" pagination="49 - 52" refId="B38" refString="Sheldon, WG, 1920. The life-cycle of Cacoeciaunifasciana Duponchel. Entomologist 53: 49 - 52" title="The life-cycle of Cacoeciaunifasciana Duponchel." volume="53" year="1920">Sheldon (1920)</bibRefCitation>
and
<bibRefCitation author="Bradley, JD" journalOrPublisher="The Entomologist's Gazette" pageId="0" pageNumber="51" refId="B4" refString="Bradley, JD, Tremewan, WG, Smith, A, 1973. British tortricoid moths. Cochylidae and Tortricidae: Tortricinae. The Ray Society, London, 251 pp [47 pls.]" title="British tortricoid moths. Cochylidae and Tortricidae: Tortricinae. The Ray Society, London, 251 pp [47 pls.]" year="1973">Bradley et al. (1973)</bibRefCitation>
. The chaetotaxy of the larva is described by
<bibRefCitation author="Swatschek, B" journalOrPublisher="Entomologist" pageId="0" pageNumber="51" refId="B40" refString="Swatschek, B, 1958. Die Larvalsystematik der Wickler (Tortricidae und Carposinidae). Abhandlungen zur Larvalsystematik der Insekten 3. Berlin, 269 pp." title="Die Larvalsystematik der Wickler (Tortricidae und Carposinidae). Abhandlungen zur Larvalsystematik der Insekten 3. Berlin, 269 pp." year="1958">Swatschek (1958)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="molecular data">
<paragraph pageId="0" pageNumber="51">Molecular data</paragraph>
<paragraph pageId="0" pageNumber="51">
(
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Neighbour-joining tree of Clepsis neglectana and C. consimilana species groups (Kimura 2 parameter, built with MEGA 6; cf. Tamura et al. 2013). Note: the scale bar only applies to internal branches between species. The width of the triangles represents the sample size, and the depth the relative genetic variation within the cluster (2 x scale bar). Source: DNA Barcode data from BOLD (Barcode of Life Database, cf. Ratnasingham and Hebert 2007)." figureDoi="10.3897/zookeys.885.38655.figure16" httpUri="https://binary.pensoft.net/fig/352394" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure16">Fig. 16</figureCitation>
). BIN: BOLD:AAC4212. The intraspecific average of the barcode region is 0.34%, the maximum distance 1.08% (p-dist) (
<emphasis italics="true" pageId="0" pageNumber="51">N</emphasis>
= 29). The minimum distance to the nearest neighbour,
<taxonomicName class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis eatoniana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">Clepsis eatoniana</emphasis>
</taxonomicName>
, is 2.25%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="distribution">
<paragraph pageId="0" pageNumber="51">Distribution</paragraph>
<paragraph pageId="0" pageNumber="51">
(
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution map of examined material of Clepsis neglectana and C. consimilana species groups. Map created with SimpleMappr (http: // www. simplemappr. net)." figureDoi="10.3897/zookeys.885.38655.figure17" httpUri="https://binary.pensoft.net/fig/352395" pageId="0" pageNumber="51" tableDoi="10.3897/zookeys.885.38655.figure17">Fig. 17</figureCitation>
). Europe, Asia Minor, Syria, European Russia, W Africa to Lebanon, Madagascar (introduced; the data come from the type specimen only), North America (introduced) (
<bibRefCitation author="Bradley, JD" journalOrPublisher="The Entomologist's Gazette" pageId="0" pageNumber="51" refId="B4" refString="Bradley, JD, Tremewan, WG, Smith, A, 1973. British tortricoid moths. Cochylidae and Tortricidae: Tortricinae. The Ray Society, London, 251 pp [47 pls.]" title="British tortricoid moths. Cochylidae and Tortricidae: Tortricinae. The Ray Society, London, 251 pp [47 pls.]" year="1973">Bradley et al. 1973</bibRefCitation>
,
<bibRefCitation author="Razowski, J" journalOrPublisher="Slamka, Bratislava" pageId="0" pageNumber="51" refId="B35" refString="Razowski, J, 2002. Tortricidae of Europe. Vol. 1. Tortricinae and Chlidanothinae. Slamka, Bratislava" title="Tortricidae of Europe. Vol. 1. Tortricinae and Chlidanothinae." year="2002">Razowski 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="ecology">
<paragraph pageId="0" pageNumber="51">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="51">
Moths are on the wing from May to October in shrubby habitats. The larvae feed on dead or withered leaves of
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Crataegus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Crataegus" order="Rosales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Crataegus</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Malus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Malus" order="Rosales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Malus</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carpinus" order="Fagales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Carpinus</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Polygonaceae" genus="Polygonum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Polygonum" order="Caryophyllales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Polygonum</emphasis>
</taxonomicName>
,
<taxonomicName class="Anthozoa" family="Alcyoniidae" genus="Hedera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hedera" order="Alcyonacea" pageId="0" pageNumber="51" phylum="Cnidaria" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Hedera</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Caprifoliaceae" genus="Lonicera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lonicera" order="Dipsacales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Lonicera</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Oleaceae" genus="Ligustrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ligustrum" order="Lamiales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Ligustrum</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Oleaceae" genus="Syringa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Syringa" order="Lamiales" pageId="0" pageNumber="51" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Syringa</emphasis>
</taxonomicName>
, and overwinter in the third instar (
<bibRefCitation author="Bradley, JD" journalOrPublisher="The Entomologist's Gazette" pageId="0" pageNumber="51" refId="B4" refString="Bradley, JD, Tremewan, WG, Smith, A, 1973. British tortricoid moths. Cochylidae and Tortricidae: Tortricinae. The Ray Society, London, 251 pp [47 pls.]" title="British tortricoid moths. Cochylidae and Tortricidae: Tortricinae. The Ray Society, London, 251 pp [47 pls.]" year="1973">Bradley et al. 1973</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="remarks">
<paragraph pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="51">
The collection of Jakob
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
was acquired by Vincenz Abbate Edler von Mazzola in the early part of the 19th Century, and the European material was deposited in the
<normalizedToken originalValue="“Hof-Naturalien-Kabinett”">&quot;Hof-Naturalien-Kabinett&quot;</normalizedToken>
in 1823 where most of the material was destroyed by fire during the Vienna Rebellion of 1848 (
<bibRefCitation author="Calhoun, JV" journalOrPublisher="Journal of the Lepidopterists' Society" pageId="0" pageNumber="51" pagination="204 - 219" refId="B5" refString="Calhoun, JV, 2003. The history and true identity of Melitaeaismeria (Nymphalidae): A remarkable tale of duplication, misinterpretation, and presumption. Journal of the Lepidopterists' Society 57: 204 - 219" title="The history and true identity of Melitaeaismeria (Nymphalidae): A remarkable tale of duplication, misinterpretation, and presumption." volume="57" year="2003">Calhoun 2003</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3630.3.5" author="Gilligan, TM" journalOrPublisher="Zootaxa" pageId="0" pageNumber="51" pagination="489 - 504" refId="B17" refString="Gilligan, TM, Wright, DJ, 2013. The type species of Eucosma Huebner (Lepidoptera: Tortricidae: Eucosmini). Zootaxa 3630: 489 - 504" title="The type species of Eucosma Huebner (Lepidoptera: Tortricidae: Eucosmini)." url="https://doi.org/10.11646/zootaxa.3630.3.5" volume="3630" year="2013">Gilligan and Wright 2013</bibRefCitation>
). Despite a personal search by PH we found no potential type material and conclude that the syntypes are lost or destroyed. As the boundaries of this taxon are revised, designation of a neotype is necessary to preserve the stability of nomenclature. The type locality is presumably Europe, as indicated by the title of the original description (
<bibRefCitation author="Huebner, J" journalOrPublisher="Regensburg, Manz" pageId="0" pageNumber="51" refId="B20" refString="Huebner, J, 1817. Der Sammlung europaeischer Schmetterlinge siebente Horde. Tortrices-Wickler. Augsburg, 340 pls." title="Der Sammlung europaeischer Schmetterlinge siebente Horde. Tortrices-Wickler. Augsburg, 340 pls." year="1817">
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
1817
</bibRefCitation>
) and material may have originated from Germany as many other species described by
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
. Here we designate as neotype a male specimen with an everted vesica and preserved cornuti, and wing pattern resembling the illustration in the work by
<bibRefCitation author="Huebner, J" journalOrPublisher="Regensburg, Manz" pageId="0" pageNumber="51" refId="B20" refString="Huebner, J, 1817. Der Sammlung europaeischer Schmetterlinge siebente Horde. Tortrices-Wickler. Augsburg, 340 pls." title="Der Sammlung europaeischer Schmetterlinge siebente Horde. Tortrices-Wickler. Augsburg, 340 pls." year="1817">
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
(1817
</bibRefCitation>
: pl. 38, fig. 239). Though not very detailed, this painting demonstrates some important differences with
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
sensu auctt., namely the shape of basal fascia and costa, and questions the interpretation of the taxon. The painting either refers to another representative of the tribe
<taxonomicName lsidName="" pageId="0" pageNumber="51" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
or may be a result of an artistic decision. Without studying the type material any interpretation of the figure may be incorrect, therefore we prefer to preserve the present-day concept for
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
.
<bibRefCitation author="Obraztsov, NS" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="51" pagination="147 - 228" refId="B29" refString="Obraztsov, NS, 1955. Die Gattungen der Palaearktischen Tortricidae I. Allgemenie Aufteilung der Familie und due Unterfamilien Tortricinae und Sparganothinae. Tijdschrift voor Entomologie 98: 147 - 228" title="Die Gattungen der Palaearktischen Tortricidae I. Allgemenie Aufteilung der Familie und due Unterfamilien Tortricinae und Sparganothinae." volume="98" year="1955">Obraztsov (1955)</bibRefCitation>
considered
<taxonomicName lsidName="consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">consimilana</emphasis>
</taxonomicName>
as a probable synonym of
<taxonomicName lsidName="unifasciana" pageId="0" pageNumber="51" rank="species" species="unifasciana">
<emphasis italics="true" pageId="0" pageNumber="51">unifasciana</emphasis>
</taxonomicName>
.
<bibRefCitation author="Bradley, JD" journalOrPublisher="The Entomologist's Gazette" pageId="0" pageNumber="51" pagination="151 - 156" refId="B3" refString="Bradley, JD, Martin, EL, 1956. An illustrated list of British Tortricidae. The Entomologist's Gazette 7: 151 - 156" title="An illustrated list of British Tortricidae." volume="7" year="1956">Bradley and Martin (1956)</bibRefCitation>
established the name
<taxonomicName lsidName="consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">consimilana</emphasis>
</taxonomicName>
as having priority over
<taxonomicName lsidName="unifasciana" pageId="0" pageNumber="51" rank="species" species="unifasciana">
<emphasis italics="true" pageId="0" pageNumber="51">unifasciana</emphasis>
</taxonomicName>
, though they did not state explicitly this nomenclatorial act. We found support for this synonymy as the lectotype of
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix unifasciana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="unifasciana">
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix unifasciana</emphasis>
</taxonomicName>
appears conspecific with the neotype of
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
.
<bibRefCitation author="Razowski, J" journalOrPublisher="Acta zoologica cracoviensia" pageId="0" pageNumber="51" pagination="101 - 198" refId="B34" refString="Razowski, J, 1979. Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I. Acta zoologica cracoviensia 23: 101 - 198" title="Revision of the Genus Clepsis Guenee (Lepidoptera, Tortricidae). Part I." volume="23" year="1979">Razowski (1979)</bibRefCitation>
synonymised
<taxonomicName class="Insecta" family="Tortricidae" genus="Tortrix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tortrix eatoniana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="eatoniana">
<emphasis italics="true" pageId="0" pageNumber="51">Tortrix eatoniana</emphasis>
</taxonomicName>
with
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
(see below); this is not justified in our opinion. Some taxa from the synonymic list were not examined by us but, minding the minimal genetic diversity throughout Europe, we consider all of them correctly synonymised with
<taxonomicName lsidName="C. consimilana" pageId="0" pageNumber="51" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">C. consimilana</emphasis>
</taxonomicName>
.
<taxonomicName class="Insecta" family="Tortricidae" genus="Cacoecia" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Cacoecia unifasciana var. semiana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="variety" species="unifasciana" variety="semiana">Cacoecia unifasciana var. semiana</taxonomicName>
was erroneously regarded as a synonym of
<taxonomicName class="Insecta" family="Tortricidae" genus="Clepsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clepsis consimilana" order="Lepidoptera" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="consimilana">
<emphasis italics="true" pageId="0" pageNumber="51">Clepsis consimilana</emphasis>
</taxonomicName>
(see above).
</paragraph>
</subSubSection>
</treatment>
</document>