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<document id="E79F1BF2E966CA3E05B44F9ED55680F3" ID-DOI="10.5281/zenodo.188001" ID-GBIF-Dataset="ae77a7c0-d0d3-4319-850a-d73dcc5cc218" ID-ISSN="1175-5326" ID-Zenodo-Dep="188001" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461079012509" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Jaschhof, Catrin" docDate="2009" docId="AF5F87C0FFB2BF20FF29FD318B253151" docLanguage="en" docName="zt02117p048.pdf" docOrigin="Zootaxa 2117" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cowanomyia hillaryi Jaschhof &amp; Jaschhof, new species" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="5366FFB8FFB0BF25FFBEFF848A353077" masterDocTitle="Cowanomyia hillaryi gen. et sp. n., a remarkable new gnoristine (Diptera: Mycetophilidae: Sciophilinae) from New Zealand" masterLastPageNumber="48" masterPageNumber="43" pageNumber="45" updateTime="1698587231274" updateUser="plazi">
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<mods:title id="D8C95ACD967BCEE7202C2732E61B2A18">Cowanomyia hillaryi gen. et sp. n., a remarkable new gnoristine (Diptera: Mycetophilidae: Sciophilinae) from New Zealand</mods:title>
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<mods:namePart id="42BA67E562B38CD4B7CD21227E31BA36">Jaschhof, Mathias</mods:namePart>
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<mods:namePart id="2D87C41D748BB18CED396986EE35147D">Jaschhof, Catrin</mods:namePart>
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<mods:date id="C9FEC71EE892E64A910E1E671E2BA11F">2009</mods:date>
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<mods:identifier id="0065F92F36885901D65D48829F83359B" type="DOI">10.5281/zenodo.188001</mods:identifier>
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<treatment id="AF5F87C0FFB2BF20FF29FD318B253151" ID-DOI="http://doi.org/10.5281/zenodo.6214378" ID-GBIF-Taxon="119618340" ID-Zenodo-Dep="6214378" LSID="urn:lsid:plazi:treatment:AF5F87C0FFB2BF20FF29FD318B253151" httpUri="http://treatment.plazi.org/id/AF5F87C0FFB2BF20FF29FD318B253151" lastPageId="5" lastPageNumber="47" pageId="2" pageNumber="45">
<subSubSection id="6FEC655DFFB2BF27FF29FD318B253285" pageId="2" pageNumber="45" type="nomenclature">
<paragraph id="274936D6FFB2BF27FF29FD31890232B8" blockId="2.[151,823,693,754]" box="[151,823,693,719]" pageId="2" pageNumber="45">
<heading id="7C0181BAFFB2BF27FF29FD31890232B8" bold="true" box="[151,823,693,719]" fontSize="11" level="1" pageId="2" pageNumber="45" reason="1">
<emphasis id="1582EAC4FFB2BF27FF29FD31890232B8" bold="true" box="[151,823,693,719]" pageId="2" pageNumber="45">
<taxonomicName id="E0F64D55FFB2BF27FF29FD31889532B8" ID-CoL="Z34C" authority="Jaschhof &amp; Jaschhof" authorityName="Jaschhof &amp; Jaschhof" box="[151,672,693,719]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="hillaryi" status="sp. nov.">
<emphasis id="1582EAC4FFB2BF27FF29FD318BA132B8" bold="true" box="[151,404,693,719]" italics="true" pageId="2" pageNumber="45">Cowanomyia hillaryi</emphasis>
Jaschhof &amp; Jaschhof
</taxonomicName>
<taxonomicNameLabel id="0EB157BFFFB2BF27FD19FD31890232B8" box="[679,823,693,719]" pageId="2" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="274936D6FFB2BF27FF29FD5C8B253285" blockId="2.[151,823,693,754]" box="[151,272,728,754]" pageId="2" pageNumber="45">
(
<figureCitation id="BFCD2A53FFB2BF27FF21FD5C8B323285" box="[159,263,728,754]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 7" captionStartId-0="3.[151,270,1641,1665]" captionStartId-1="4.[151,274,1860,1884]" captionTargetBox-0="[195,1405,192,1553]" captionTargetBox-1="[186,1389,295,1859]" captionTargetId-0="figure@3.[151,1436,141,1616]" captionTargetId-1="figure@4.[151,1436,283,1863]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 – 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." captionText-1="FIGURES 5 – 7. Cowanomyia hillaryi gen. et sp. n., male. 5: Thorax, lateral view. 6: Terminalia, ventral view. 7: Terminalia, dorsal view. Length of scale bar = 0.1 mm (for 5) and 0.05 mm (for 6 and 7)." httpUri-0="https://zenodo.org/record/188002/files/figure.png" httpUri-1="https://zenodo.org/record/188003/files/figure.png" pageId="2" pageNumber="45">Figs 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6FEC655DFFB2BF27FF29FCA18FAD3403" pageId="2" pageNumber="45" type="description">
<paragraph id="274936D6FFB2BF27FF29FCA18E263348" blockId="2.[151,1437,805,2030]" box="[151,1043,805,831]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF29FCA18B1A3348" bold="true" box="[151,303,805,831]" pageId="2" pageNumber="45">Description.</emphasis>
Male.
<emphasis id="1582EAC4FFB2BF27FE3FFCA18BFE3348" bold="true" box="[385,459,805,831]" pageId="2" pageNumber="45">Head.</emphasis>
<figureCitation id="BFCD2A53FFB2BF27FE6DFCA1880E3348" box="[467,571,805,831]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[195,1405,192,1553]" captionTargetId="figure@3.[151,1436,141,1616]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 – 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="2" pageNumber="45">Figs 1–2</figureCitation>
. Antennal flagellomeres 3–5, see
<figureCitation id="BFCD2A53FFB2BF27FC76FCA18E3A3348" box="[968,1039,805,831]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[195,1405,192,1553]" captionTargetId="figure@3.[151,1436,141,1616]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 – 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="2" pageNumber="45">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="274936D6FFB2BF27FF7BFCCF88D03311" blockId="2.[151,1437,805,2030]" box="[197,741,843,870]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF7BFCCF8B1C3312" bold="true" box="[197,297,843,869]" pageId="2" pageNumber="45">Thorax.</emphasis>
<figureCitation id="BFCD2A53FFB2BF27FE8EFCC88B423311" box="[304,375,844,870]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,274,1860,1884]" captionTargetBox="[186,1389,295,1859]" captionTargetId="figure@4.[151,1436,283,1863]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Cowanomyia hillaryi gen. et sp. n., male. 5: Thorax, lateral view. 6: Terminalia, ventral view. 7: Terminalia, dorsal view. Length of scale bar = 0.1 mm (for 5) and 0.05 mm (for 6 and 7)." httpUri="https://zenodo.org/record/188003/files/figure.png" pageId="2" pageNumber="45">Fig. 5</figureCitation>
.
<emphasis id="1582EAC4FFB2BF27FE3DFCCF8BFB3312" bold="true" box="[387,462,843,869]" pageId="2" pageNumber="45">Wing.</emphasis>
<figureCitation id="BFCD2A53FFB2BF27FE68FCC888283311" box="[470,541,844,870]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[195,1405,192,1553]" captionTargetId="figure@3.[151,1436,141,1616]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 – 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="2" pageNumber="45">Fig. 3</figureCitation>
. Length
<quantity id="E00E9B33FFB2BF27FD3DFCC888D43311" box="[643,737,844,870]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="45" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
<paragraph id="274936D6FFB2BF27FF7BFCF68FAD3403" blockId="2.[151,1437,805,2030]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF7BFCF68BA033FB" bold="true" box="[197,405,882,908]" pageId="2" pageNumber="45">Male terminalia.</emphasis>
<figureCitation id="BFCD2A53FFB2BF27FE22FCF6883333FB" box="[412,518,882,908]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,274,1860,1884]" captionTargetBox="[186,1389,295,1859]" captionTargetId="figure@4.[151,1436,283,1863]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Cowanomyia hillaryi gen. et sp. n., male. 5: Thorax, lateral view. 6: Terminalia, ventral view. 7: Terminalia, dorsal view. Length of scale bar = 0.1 mm (for 5) and 0.05 mm (for 6 and 7)." httpUri="https://zenodo.org/record/188003/files/figure.png" pageId="2" pageNumber="45">Figs 6–7</figureCitation>
. Not rotated. Sternite 9 not traceable. Tergite 9 with straight apical margin and strongly concave basal margin, setae concentrated apicolaterally. Gonocoxites broadly merged ventrobasally, ventrobasal portion without mesal and basal setae; apicolateral processes terminating in narrow, bare projections that cross each other. Gonostylus inserted at midlength of gonocoxite, elongate, narrow, with apical broadening and 2 fine subapical setulae, otherwise bare. Ejaculatory apodeme with rodlike basal portion and complex, rhomb-shaped apical extension. Aedeagus/paramere complex largely membranous. Sternite 10 unipartite, with 2 large apical setae. Cerci rounded, with numerous dorsal and 4 large apical setae.
</paragraph>
</subSubSection>
<subSubSection id="6FEC655DFFB2BF27FF7BFB0589F8352B" pageId="2" pageNumber="45" type="etymology">
<paragraph id="274936D6FFB2BF27FF7BFB058E47349F" blockId="2.[151,1437,805,2030]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF7BFB058B7434ED" box="[197,321,1153,1178]" italics="true" pageId="2" pageNumber="45">Etymology</emphasis>
. We name this new species in memory of Sir Edmund Hillary, the Patron of the
<collectingCountry id="5FE17646FFB2BF27FB43FB058FA934EC" box="[1277,1436,1153,1179]" name="New Zealand" pageId="2" pageNumber="45">New Zealand</collectingCountry>
Native Forests Restoration Trust until his death in
<date id="53481016FFB2BF27FD44FB2C89AB34B5" box="[762,926,1192,1218]" pageId="2" pageNumber="45" value="2008-01">January 2008</date>
, and to honour the Trust´s dedicated work from which forest-dwelling arthropods, such as fungus gnats, should benefit greatly.
</paragraph>
<paragraph id="274936D6FFB2BF27FF7BFB7189F8352B" blockId="2.[151,1437,805,2030]" pageId="2" pageNumber="45">
<typeStatus id="F84D8874FFB2BF27FF7BFB718B303579" box="[197,261,1269,1294]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF7BFB718B303579" box="[197,261,1269,1294]" italics="true" pageId="2" pageNumber="45">Types</emphasis>
</typeStatus>
.
<emphasis id="1582EAC4FFB2BF27FEA8FB718BB43579" box="[278,385,1269,1294]" italics="true" pageId="2" pageNumber="45">
<typeStatus id="F84D8874FFB2BF27FEA8FB718BB43579" box="[278,385,1269,1294]" pageId="2" pageNumber="45" type="holotype">Holotype</typeStatus>
</emphasis>
. Male,
<collectingCountry id="5FE17646FFB2BF27FE63FB7188483578" box="[477,637,1269,1295]" name="New Zealand" pageId="2" pageNumber="45">New Zealand</collectingCountry>
, South Island, Westland, Westland National Park, Canavan´s Knob, in grassy clearing in mixed podocarp forest,
<quantity id="E00E9B33FFB2BF27FD22FA9888DC3541" box="[668,745,1308,1334]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="45" unit="m" value="140.0">140 m</quantity>
,
<date id="53481016FFB2BF27FD4AFA9889853541" box="[756,944,1308,1334]" pageId="2" pageNumber="45" value="1982-10" valueMax="1982-11-31" valueMin="1982-10-00">Oct.–Nov. 1982</date>
, by Malaise trap, A. B. Miller (in NZAC).
<emphasis id="1582EAC4FFB2BF27FF29FAC78B20352B" box="[151,277,1347,1372]" italics="true" pageId="2" pageNumber="45">
<typeStatus id="F84D8874FFB2BF27FF29FAC78B27352B" box="[151,274,1347,1372]" pageId="2" pageNumber="45" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="31F0FD5FFFB2BF27FEA2FAC68B4C352B" box="[284,377,1346,1372]" pageId="2" pageNumber="45" type="male">2 males</specimenCount>
, same data as the
<typeStatus id="F84D8874FFB2BF27FDF2FAC68887352B" box="[588,690,1346,1372]" pageId="2" pageNumber="45" type="holotype">holotype</typeStatus>
(in NZAC and SMTD).
</paragraph>
</subSubSection>
<subSubSection id="6FEC655DFFB2BF20FF78FAEC8B253151" lastPageId="5" lastPageNumber="48" pageId="2" pageNumber="45" type="discussion">
<paragraph id="274936D6FFB2BF27FF78FAEC8E0D3799" blockId="2.[151,1437,805,2030]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF78FAEC8B6435F5" bold="true" box="[198,337,1384,1410]" pageId="2" pageNumber="45">Discussion.</emphasis>
<taxonomicName id="E0F64D55FFB2BF27FEE6FAED8BC635F5" box="[344,499,1385,1410]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB2BF27FEE6FAED8BC635F5" box="[344,499,1385,1410]" italics="true" pageId="2" pageNumber="45">Cowanomyia</emphasis>
</taxonomicName>
and
<taxonomicName id="E0F64D55FFB2BF27FD95FAED888B35F5" box="[555,702,1385,1410]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB2BF27FD95FAED888B35F5" box="[555,702,1385,1410]" italics="true" pageId="2" pageNumber="45">Adicroneura</emphasis>
</taxonomicName>
are the only
<taxonomicName id="E0F64D55FFB2BF27FCEAFAED8E3935F4" box="[852,1036,1385,1411]" class="Insecta" family="Mycetophilidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="family">Mycetophilidae</taxonomicName>
known to have only one complete median branch in combination with a furcate anterior cubitus. Apart from that, these two genera have little in common and are evidently not closely related. Even their vein patterns include a number of differences (character states in
<taxonomicName id="E0F64D55FFB2BF27FEC2FA5988223581" box="[380,535,1501,1526]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB2BF27FEC2FA5988223581" box="[380,535,1501,1526]" italics="true" pageId="2" pageNumber="45">Cowanomyia</emphasis>
</taxonomicName>
in parentheses): in
<taxonomicName id="E0F64D55FFB2BF27FCBFFA598E243581" box="[769,1041,1501,1526]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="biocellata">
<emphasis id="1582EAC4FFB2BF27FCBFFA598E243581" box="[769,1041,1501,1526]" italics="true" pageId="2" pageNumber="45">Adicroneura biocellata</emphasis>
</taxonomicName>
, the
<typeStatus id="F84D8874FFB2BF27FBF3FA598EB53580" box="[1101,1152,1501,1527]" pageId="2" pageNumber="45">type</typeStatus>
spcies, Sc is strong and reaches C (faint and ending free); R1 is very long and ends far in the distal half of the wing (much shorter and ending in the basal half of the wing); Rs and r-m form almost a right angle (oblique); the basal half of M is very faint (M is comparatively faint throughout); and all longitudinal veins except CuP are described to have dorsal and ventral setae (only dorsal setae present on R, R1 and a part of R5). As regards characters other than those of wing, in
<taxonomicName id="E0F64D55FFB2BF27FEDEF91B885A36CF" box="[352,623,1695,1720]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="biocellata">
<emphasis id="1582EAC4FFB2BF27FEDEF91B885A36CF" box="[352,623,1695,1720]" italics="true" pageId="2" pageNumber="45">Adicroneura biocellata</emphasis>
</taxonomicName>
the mesal ocellus is missing (present); mediotergite and laterotergites are setose (asetose); the hind coxae bear a vertical row of setae (such row absent); and the fore tibial organ is missing (well developed). There might be further differences, for instance in thorax structure, which however do not become apparent from Vockeroth´s description that does not fully illustrate the thorax of
<taxonomicName id="E0F64D55FFB2BF27FB40F8978FA1375B" box="[1278,1428,1811,1836]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="biocellata">
<emphasis id="1582EAC4FFB2BF27FB40F8978FA1375B" box="[1278,1428,1811,1836]" italics="true" pageId="2" pageNumber="45">A. biocellata</emphasis>
</taxonomicName>
.
<bibRefCitation id="43674B27FFB2BF27FF29F8BD8B0E3724" author="Matile" box="[151,315,1849,1875]" pageId="2" pageNumber="48" refString="Matile, L. (1995) Le genre Adicroneura Vockeroth decouvert en region neotropicale (Diptera, Mycetophilidae). Bulletin de la Societe entomologique de France, 100 (1), 7 - 10." type="journal article" year="1995">Matile (1995)</bibRefCitation>
described and discussed a number of morphological differences between
<taxonomicName id="E0F64D55FFB2BF27FB15F8BD8F773725" box="[1195,1346,1849,1874]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="biocellata">
<emphasis id="1582EAC4FFB2BF27FB15F8BD8F773725" box="[1195,1346,1849,1874]" italics="true" pageId="2" pageNumber="45">A. biocellata</emphasis>
</taxonomicName>
, on the one hand, and
<emphasis id="1582EAC4FFB2BF27FEF0F8E488F5370E" box="[334,704,1888,1913]" italics="true" pageId="2" pageNumber="45">
<taxonomicName id="E0F64D55FFB2BF27FEF0F8E4883A370E" box="[334,527,1888,1913]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="meridionalis">A. meridionalis</taxonomicName>
and
<taxonomicName id="E0F64D55FFB2BF27FDF0F8E488F5370E" box="[590,704,1888,1913]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="disjuncta">disjuncta</taxonomicName>
</emphasis>
, on the other hand. Contrary to his conclusion, we are not convinced that the species from
<collectingCountry id="5FE17646FFB2BF27FDAAF802886137D7" box="[532,596,1926,1952]" name="Chile" pageId="2" pageNumber="45">Chile</collectingCountry>
are congeneric with the
<typeStatus id="F84D8874FFB2BF27FCC9F802899F37D7" box="[887,938,1926,1952]" pageId="2" pageNumber="45">type</typeStatus>
species. Further, there is no indication for the affinity of the two Chilean species and
<taxonomicName id="E0F64D55FFB2BF27FD30F82989B337B1" box="[654,902,1965,1990]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="hillaryi">
<emphasis id="1582EAC4FFB2BF27FD30F82989B337B1" box="[654,902,1965,1990]" italics="true" pageId="2" pageNumber="45">Cowanomyia hillaryi</emphasis>
</taxonomicName>
. If our view is correct, then the
<taxonomicName id="E0F64D55FFB2BF27FB41F8298FA737B1" box="[1279,1426,1965,1990]" class="Insecta" family="Mycetophilidae" genus="Adicroneura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB2BF27FB41F8298FA737B1" box="[1279,1426,1965,1990]" italics="true" pageId="2" pageNumber="45">Adicroneura</emphasis>
</taxonomicName>
- like vein pattern must have evolved two or even three times in
<taxonomicName id="E0F64D55FFB2BF27FCC6F8508E013799" box="[888,1076,2004,2030]" class="Insecta" family="Mycetophilidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="family">Mycetophilidae</taxonomicName>
.
</paragraph>
<caption id="7389665EFFB3BF26FF29F9ED894B36B2" httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="3" pageNumber="46" targetBox="[195,1405,192,1553]" targetPageId="3">
<paragraph id="274936D6FFB3BF26FF29F9ED894B36B2" blockId="3.[151,1436,1641,1733]" pageId="3" pageNumber="46">
<emphasis id="1582EAC4FFB3BF26FF29F9ED8B7136F6" bold="true" box="[151,324,1641,1665]" pageId="3" pageNumber="46">FIGURES 1–4.</emphasis>
<taxonomicName id="E0F64D55FFB3BF26FEF5F9EE881A36F6" box="[331,559,1642,1665]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="hillaryi">
<emphasis id="1582EAC4FFB3BF26FEF5F9EE881A36F6" box="[331,559,1642,1665]" italics="true" pageId="3" pageNumber="46">Cowanomyia hillaryi</emphasis>
</taxonomicName>
<emphasis id="1582EAC4FFB3BF26FD88F9ED88F036F6" bold="true" box="[566,709,1641,1665]" pageId="3" pageNumber="46">gen. et sp. n.</emphasis>
, male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3–5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4).
</paragraph>
</caption>
<paragraph id="274936D6FFB3BF21FF7BF9708E343151" blockId="3.[151,1437,1780,2038]" lastBlockId="4.[151,1437,152,294]" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46">
The genus
<taxonomicName id="E0F64D55FFB3BF26FEF5F9708BDF377A" box="[331,490,1780,1805]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB3BF26FEF5F9708BDF377A" box="[331,490,1780,1805]" italics="true" pageId="3" pageNumber="46">Cowanomyia</emphasis>
</taxonomicName>
fits well the diagnosis for the Gnoristini as revised by
<bibRefCitation id="43674B27FFB3BF26FB33F9708FA23779" author="Vaisanen" box="[1165,1431,1780,1806]" pageId="3" pageNumber="48" refString="Vaisanen, R. (1986) The delimitation of the Gnoristinae: criteria for the classification of recent European genera (Diptera, Mycetophilidae). Annales Zoologici Fennici, 23, 197 - 206." type="journal article" year="1986">Väisänen (1986: 203)</bibRefCitation>
, irrespective of the fact that he did not distinguish between apomorphous and plesiomorphous character states. Characters supporting the affiliation of
<taxonomicName id="E0F64D55FFB3BF26FDDAF8C588CA372D" box="[612,767,1857,1882]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB3BF26FDDAF8C588CA372D" box="[612,767,1857,1882]" italics="true" pageId="3" pageNumber="46">Cowanomyia</emphasis>
</taxonomicName>
to the Gnoristini are: the presence of 3 ocelli, the laterals distant from the eye margins; the sharply delimited sensory depression on the third palpus segment; the anapleural suture declining posteriorly; the bare mediotergite; the absence of setae on the wing membrane; the short, oblique r-m; CuA with a long stem; and CuA2 evenly bent, not sinuous. Among Gnoristini sensu
<bibRefCitation id="43674B27FFB3BF26FF29F8588B693781" author="Vaisanen" box="[151,348,2012,2038]" pageId="3" pageNumber="48" refString="Vaisanen, R. (1986) The delimitation of the Gnoristinae: criteria for the classification of recent European genera (Diptera, Mycetophilidae). Annales Zoologici Fennici, 23, 197 - 206." type="journal article" year="1986">Väisänen (1986)</bibRefCitation>
,
<taxonomicName id="E0F64D55FFB3BF26FED6F85888363782" box="[360,515,2012,2037]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB3BF26FED6F85888363782" box="[360,515,2012,2037]" italics="true" pageId="3" pageNumber="46">Cowanomyia</emphasis>
</taxonomicName>
is peculiar for its enlarged proepimeron; the lack of distinct bare stripes on the scutum; the faint, free-ending Sc; the short R1; the one-branched media; and the sparsely setose hind coxae. Some of these peculiarities, such as those regarding wings and legs, might result from miniaturisation:
<taxonomicName id="E0F64D55FFB4BF21FF29FF618B0C3089" box="[151,313,229,254]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB4BF21FF29FF618B0C3089" box="[151,313,229,254]" italics="true" pageId="4" pageNumber="47">Cowanomyia</emphasis>
</taxonomicName>
males are among the smallest mycetophilids known to us and are actually as small as the micromyid gall midges with which they share the similar venation pattern.
</paragraph>
<caption id="7389665EFFB4BF21FF29F8C08E7F3709" httpUri="https://zenodo.org/record/188003/files/figure.png" pageId="4" pageNumber="47" targetBox="[186,1389,295,1859]" targetPageId="4">
<paragraph id="274936D6FFB4BF21FF29F8C08E7F3709" blockId="4.[151,1437,1860,1918]" pageId="4" pageNumber="47">
<emphasis id="1582EAC4FFB4BF21FF29F8C08B7E372B" bold="true" box="[151,331,1860,1884]" pageId="4" pageNumber="47">FIGURES 5–7.</emphasis>
<taxonomicName id="E0F64D55FFB4BF21FEEDF8C18874372B" box="[339,577,1861,1884]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="hillaryi">
<emphasis id="1582EAC4FFB4BF21FEEDF8C18874372B" box="[339,577,1861,1884]" italics="true" pageId="4" pageNumber="47">Cowanomyia hillaryi</emphasis>
</taxonomicName>
<emphasis id="1582EAC4FFB4BF21FDF7F8C088EC372B" bold="true" box="[585,729,1860,1884]" pageId="4" pageNumber="47">gen. et sp. n</emphasis>
., male. 5: Thorax, lateral view. 6: Terminalia, ventral view. 7: Terminalia, dorsal view. Length of scale bar = 0.1 mm (for 5) and 0.05 mm (for 6 and 7).
</paragraph>
</caption>
<paragraph id="274936D6FFB4BF20FF7BF8358B253151" blockId="4.[151,1436,1969,2033]" lastBlockId="5.[151,1436,152,294]" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47">
Due to its peculiar assemblage of characters, the relationship of
<taxonomicName id="E0F64D55FFB4BF21FC05F8358E6337BD" box="[955,1110,1969,1994]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB4BF21FC05F8358E6337BD" box="[955,1110,1969,1994]" italics="true" pageId="4" pageNumber="47">Cowanomyia</emphasis>
</taxonomicName>
to other Gnoristini remains obscure. To our knowledge there are no gnoristines in
<collectingCountry id="5FE17646FFB4BF21FCA1F853898B3786" box="[799,958,2007,2033]" name="New Zealand" pageId="4" pageNumber="47">New Zealand</collectingCountry>
and
<collectingCountry id="5FE17646FFB4BF21FC47F8538E5D3786" box="[1017,1128,2007,2033]" name="Australia" pageId="4" pageNumber="47">Australia</collectingCountry>
, particularly Tasmania, in which adult morphology indicates an affinity to
<taxonomicName id="E0F64D55FFB5BF20FD67FF1C894030C6" box="[729,885,152,177]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB5BF20FD67FF1C894030C6" box="[729,885,152,177]" italics="true" pageId="5" pageNumber="48">Cowanomyia</emphasis>
</taxonomicName>
. The same applies to the Gnoristini described from other southern hemisphere landmasses, which however are poorly known even on the generic level. For the time being this remarkable new genus holds an undefined position among world Gnoristini, both extant and fossil.
</paragraph>
</subSubSection>
</treatment>
</document>
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