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<document id="C3682862CC819779BA7B05FA9A374F92" ID-DOI="10.5281/zenodo.188001" ID-GBIF-Dataset="ae77a7c0-d0d3-4319-850a-d73dcc5cc218" ID-ISSN="1175-5326" ID-Zenodo-Dep="188001" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461079012509" checkinUser="plazi" docAuthor="Jaschhof, Mathias &amp; Jaschhof, Catrin" docDate="2009" docId="AF5F87C0FFB1BF27FF29FD508876322C" docLanguage="en" docName="zt02117p048.pdf" docOrigin="Zootaxa 2117" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cowanomyia Jaschhof &amp; Jaschhof" docType="treatment" docVersion="9" lastPageNumber="45" masterDocId="5366FFB8FFB0BF25FFBEFF848A353077" masterDocTitle="Cowanomyia hillaryi gen. et sp. n., a remarkable new gnoristine (Diptera: Mycetophilidae: Sciophilinae) from New Zealand" masterLastPageNumber="48" masterPageNumber="43" pageNumber="44" updateTime="1698587231274" updateUser="plazi">
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<mods:title id="4C966A9CAE471D3220B2190DC94A1426">Cowanomyia hillaryi gen. et sp. n., a remarkable new gnoristine (Diptera: Mycetophilidae: Sciophilinae) from New Zealand</mods:title>
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<mods:namePart id="9DE8B6AEDF3855DC2027A155C1C0E8E7">Jaschhof, Mathias</mods:namePart>
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<mods:namePart id="79E5F0688A368BF17C7A785592AA92D8">Jaschhof, Catrin</mods:namePart>
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<mods:date id="6EF70AEB90F2F92E42F4B60B757B3BCF">2009</mods:date>
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<mods:identifier id="062E30C630788C633FB2972E42686AAF" type="DOI">10.5281/zenodo.188001</mods:identifier>
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<treatment id="AF5F87C0FFB1BF27FF29FD508876322C" ID-DOI="http://doi.org/10.5281/zenodo.6214376" ID-GBIF-Taxon="119618339" ID-Zenodo-Dep="6214376" LSID="urn:lsid:plazi:treatment:AF5F87C0FFB1BF27FF29FD508876322C" httpUri="http://treatment.plazi.org/id/AF5F87C0FFB1BF27FF29FD508876322C" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
<subSubSection id="6FEC655DFFB1BF24FF29FD5088FF329A" box="[151,714,723,750]" pageId="1" pageNumber="44" type="nomenclature">
<paragraph id="274936D6FFB1BF24FF29FD5088FF329A" blockId="1.[151,714,723,750]" box="[151,714,723,750]" pageId="1" pageNumber="44">
<heading id="7C0181BAFFB1BF24FF29FD5088FF329A" bold="true" box="[151,714,723,750]" fontSize="11" level="1" pageId="1" pageNumber="44" reason="1">
<emphasis id="1582EAC4FFB1BF24FF29FD5088FF329A" bold="true" box="[151,714,723,750]" pageId="1" pageNumber="44">
<taxonomicName id="E0F64D55FFB1BF24FF29FD508875329A" ID-CoL="3V3Q" authority="Jaschhof &amp; Jaschhof" authorityName="Jaschhof &amp; Jaschhof" box="[151,576,723,750]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="1582EAC4FFB1BF24FF29FD508B003299" bold="true" box="[151,309,724,750]" italics="true" pageId="1" pageNumber="44">Cowanomyia</emphasis>
Jaschhof &amp; Jaschhof
</taxonomicName>
new genus
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6FEC655DFFB1BF24FF29FC998E233343" box="[151,1046,797,820]" pageId="1" pageNumber="44" type="materials_examined">
<paragraph id="274936D6FFB1BF24FF29FC998E233343" blockId="1.[151,1046,797,820]" box="[151,1046,797,820]" pageId="1" pageNumber="44">
<emphasis id="1582EAC4FFB1BF24FF29FC998B293343" box="[151,284,797,820]" italics="true" pageId="1" pageNumber="44">
<typeStatus id="F84D8874FFB1BF24FF29FC998AFC3343" box="[151,201,797,820]" pageId="1" pageNumber="44">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="E0F64D55FFB1BF24FE94FC9988DB3343" authority="Jaschhof &amp; Jaschhof" authorityName="Jaschhof &amp; Jaschhof" box="[298,750,797,820]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="hillaryi" status="sp. nov.">
<emphasis id="1582EAC4FFB1BF24FE94FC9988393343" box="[298,524,797,820]" italics="true" pageId="1" pageNumber="44">Cowanomyia hillaryi</emphasis>
Jaschhof &amp; Jaschhof
</taxonomicName>
<taxonomicNameLabel id="0EB157BFFFB1BF24FD4BFC99894D3343" box="[757,888,797,820]" pageId="1" pageNumber="44" rank="species">new species</taxonomicNameLabel>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="6FEC655DFFB1BF24FF29FCE78E273448" pageId="1" pageNumber="44" type="diagnosis">
<paragraph id="274936D6FFB1BF24FF29FCE78E273448" blockId="1.[151,1438,867,2015]" pageId="1" pageNumber="44">
<emphasis id="1582EAC4FFB1BF24FF29FCE78B22330A" bold="true" box="[151,279,867,893]" pageId="1" pageNumber="44">Diagnosis.</emphasis>
This new genus includes a single species of which males are only
<quantity id="E00E9B33FFB1BF24FB99FCE08EB43309" box="[1063,1153,868,894]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="44" unit="mm" value="1.5">1.5 mm</quantity>
long and of dark brown color, the legs being somewhat lighter than thorax and abdomen. Wing venation is characterized by presence of only 1 medial branch in combination with a furcate cubitus (
<figureCitation id="BFCD2A53FFB1BF24FC2BFC3589EB33BC" box="[917,990,945,971]" captionStart="FIGURES 1 4" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[195,1405,192,1553]" captionTargetId="figure@3.[151,1436,141,1616]" captionTargetPageId="3" captionText="FIGURES 1 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="1" pageNumber="44">Fig. 3</figureCitation>
), which is an unusual pattern among Gnoristini and other
<taxonomicName id="E0F64D55FFB1BF24FE34FC5C88733385" box="[394,582,984,1010]" class="Insecta" family="Mycetophilidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="family">Mycetophilidae</taxonomicName>
. Further characters in combination add to the generic distinctiveness: the presence of 3 ocelli; the enlarged proepimeron; the faint, free-ending Sc; the short R1; and the absence of setae on the wing membrane. Females and preimaginal stages are unknown.
</paragraph>
</subSubSection>
<subSubSection id="6FEC655DFFB1BF27FF7BFBCF88AD31ED" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44" type="description">
<paragraph id="274936D6FFB1BF24FF7BFBCF8BE0359F" blockId="1.[151,1438,867,2015]" pageId="1" pageNumber="44">
<emphasis id="1582EAC4FFB1BF24FF7BFBCF8B6F3412" bold="true" box="[197,346,1099,1125]" pageId="1" pageNumber="44">Description</emphasis>
(based on males).
<emphasis id="1582EAC4FFB1BF24FDFEFBCF88BE3412" bold="true" box="[576,651,1099,1125]" pageId="1" pageNumber="44">Head.</emphasis>
<figureCitation id="BFCD2A53FFB1BF24FD2AFBC889343411" box="[660,769,1100,1126]" captionStart="FIGURES 1 4" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[195,1405,192,1553]" captionTargetId="figure@3.[151,1436,141,1616]" captionTargetPageId="3" captionText="FIGURES 1 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="1" pageNumber="44">Figs 12</figureCitation>
. In lateral view head capsule higher than long, slightly flattened on top. Foramen situated above midheight of head. Median convexities of postgenae membranous. Frons asetose, in lateral profile strongly protruding. Frontal tubercle 2-pointed. Antennae inserted above midheight of head. Scape smaller than pedicel, both setose. Fourteen flagellomeres (
<figureCitation id="BFCD2A53FFB1BF24FB35FB448EE134AD" box="[1163,1236,1216,1242]" captionStart="FIGURES 1 4" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[195,1405,192,1553]" captionTargetId="figure@3.[151,1436,141,1616]" captionTargetPageId="3" captionText="FIGURES 1 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="1" pageNumber="44">Fig. 4</figureCitation>
), subcylindrical, with short stalks, nodes darker than stalks, slightly longer than wide, covered densely with trichia arising from large, circular basal pores, longest trichia little shorter than width of node, setae absent. Interommatidial setae numerous. Three ocelli in line, the laterals remote from eye margins. Postocular bristles about 8. Face setose, subrectangular, much larger than clypeus. Clypeus narrow, with 2 or so setae. Labrum subtriangular, asetose. Maxillary palpus short, 5-segmented, 2 segments present proximally to presumed third, third segment thickest of all, with sharply delimited mesal pit of sensilla. Premental apodemes fused mesally, with 2 posterior processes. Labellum small.
</paragraph>
<paragraph id="274936D6FFB1BF27FF7BFA718E833151" blockId="1.[151,1438,867,2015]" lastBlockId="2.[151,1436,152,603]" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
<emphasis id="1582EAC4FFB1BF24FF7BFA718B1F3678" bold="true" box="[197,298,1525,1551]" pageId="1" pageNumber="44">Thorax.</emphasis>
<figureCitation id="BFCD2A53FFB1BF24FE8DFA718B493678" box="[307,380,1525,1551]" captionStart="FIGURES 5 7" captionStartId="4.[151,274,1860,1884]" captionTargetBox="[186,1389,295,1859]" captionTargetId="figure@4.[151,1436,283,1863]" captionTargetPageId="4" captionText="FIGURES 5 7. Cowanomyia hillaryi gen. et sp. n., male. 5: Thorax, lateral view. 6: Terminalia, ventral view. 7: Terminalia, dorsal view. Length of scale bar = 0.1 mm (for 5) and 0.05 mm (for 6 and 7)." httpUri="https://zenodo.org/record/188003/files/figure.png" pageId="1" pageNumber="44">Fig. 5</figureCitation>
. Proepimeron comparatively large, extending posteriad of proepisternum. Basisternite 1 setose. Scutum with lateral and dorsocentral setae of various sizes. Scutellum with 2 large lateral setae and several smaller setae elsewhere. Mediotergite and laterotergite asetose. Anapleural suture declining posteriorly. Mesopreepisternum setose ventromesally. Anepimeron short, ending far before ventral margin of laterotergite. Mid-pleural pit well developed. Metepisternum finely setose posteroventrally.
<emphasis id="1582EAC4FFB1BF24FB49F90B8F7136DE" bold="true" box="[1271,1348,1679,1705]" pageId="1" pageNumber="44">Wing.</emphasis>
<figureCitation id="BFCD2A53FFB1BF24FAF3F9148FA236DD" box="[1357,1431,1680,1706]" captionStart="FIGURES 1 4" captionStartId="3.[151,270,1641,1665]" captionTargetBox="[195,1405,192,1553]" captionTargetId="figure@3.[151,1436,141,1616]" captionTargetPageId="3" captionText="FIGURES 1 4. Cowanomyia hillaryi gen. et sp. n., male. 1: Head, lateral view. 2: Head, frontal view. 3: Wing, dorsal view, alar microtrichia and marginal setae drawn only partially. 4: Antennal flagellomeres 3 5, lateral view. Length of scale bar = 0.1 mm (for 1 and 2), 0.5 mm (for 3) and 0.05 mm (for 4)." httpUri="https://zenodo.org/record/188002/files/figure.png" pageId="1" pageNumber="44">Fig. 3</figureCitation>
. Membrane transparent, with dense, irregular microtrichia on both sides, setae absent. C extending much beyond apex of R5 and up to apex of wing. Sc very weak, ending free. R1 short, ending in proximal half of wing. Rs oblique, about as long as r-m, situated in proximal half of wing. R-m oblique. Bm-cu comparatively weak throughout. R5 almost straight. M simple, wide, not sharply delimited from membrane which applies also to CuA. CuA furcate, stem and fork subequal in length. CuP weak, wide, extending to point of furcation of CuA. Both A1 and A2 weak, wide, comparatively long, A2 curved slightly anteriorly. Dorsal setae on R, R1 and R5 proximally.
<emphasis id="1582EAC4FFB1BF24FE18F81A8BDD37CF" bold="true" box="[422,488,1950,1976]" pageId="1" pageNumber="44">Legs.</emphasis>
Fore and mid coxae setose, hind coxa with 12 setae in upper third and 34 setae apically. Femora with setae and microtrichia. Tibiae covered with large microtrichia sparsely intermingled with setae including 2 sparse rows of strong, semi-erect setae, longest setae shorter than tibial diameter. Fore tibial organ with semicircular rim and regular comb of some 15 trichia. Tibial spurs 1:2:2, with serrated rims, on mid and hind tibiae unequal in length. Mid and hind tarsi with sole of semi-erect setae. Empodia slightly shorter than pretarsal claws. Claws each with 1 large tooth, thus seemingly split in 2 tines.
</paragraph>
<paragraph id="274936D6FFB2BF27FF7BFEB68EC23104" blockId="2.[151,1436,152,603]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF7BFEB68B72313B" bold="true" box="[197,327,306,332]" pageId="2" pageNumber="45">Abdomen.</emphasis>
Segments 16 evenly setose, segment 1 short, segments 26 longer, subequal in length, only sternites of segments 78 visible, setose. Presence of foldlines uncertain. Tergal plaques absent.
</paragraph>
<paragraph id="274936D6FFB2BF27FF7BFEFB88AD31ED" blockId="2.[151,1436,152,603]" box="[197,664,383,410]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF7BFEFB8BA031EE" bold="true" box="[197,405,383,409]" pageId="2" pageNumber="45">Male terminalia.</emphasis>
See under
<taxonomicName id="E0F64D55FFB2BF27FDA6FE0488A531EE" box="[536,656,384,409]" class="Insecta" family="Mycetophilidae" genus="Cowanomyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="hillaryi">
<emphasis id="1582EAC4FFB2BF27FDA6FE0488A531EE" box="[536,656,384,409]" italics="true" pageId="2" pageNumber="45">C. hillaryi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="6FEC655DFFB2BF27FF7BFE228876322C" pageId="2" pageNumber="45" type="etymology">
<paragraph id="274936D6FFB2BF27FF7BFE228876322C" blockId="2.[151,1436,152,603]" pageId="2" pageNumber="45">
<emphasis id="1582EAC4FFB2BF27FF7BFE228B6631B7" bold="true" box="[197,339,422,448]" pageId="2" pageNumber="45">Etymology.</emphasis>
We name this new genus to honour Pat and Arthur B. Cowan of Otorohanga, North Island,
<collectingCountry id="5FE17646FFB2BF27FF29FE498B093190" box="[151,316,461,487]" name="New Zealand" pageId="2" pageNumber="45">New Zealand</collectingCountry>
, for their contribution to forest conservation and restoration in the Waikato region and many other parts of
<collectingCountry id="5FE17646FFB2BF27FEFFFE708BD03279" box="[321,485,500,526]" name="New Zealand" pageId="2" pageNumber="45">New Zealand</collectingCountry>
. Among manifold other capacities, Arthur was a founding member of the
<collectingCountry id="5FE17646FFB2BF27FADAFE708AC03243" name="New Zealand" pageId="2" pageNumber="45">New Zealand</collectingCountry>
Native Forests Restoration Trust and, in spite of being in his nineties now, is still actively involved in pest control and native tree planting.
</paragraph>
</subSubSection>
</treatment>
</document>