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<document id="8BE4D58BCEFD19C49D2F106FB39937ED" ID-CLB-Dataset="25459" ID-DOI="10.11646/zootaxa.4671.1.11" ID-GBIF-Dataset="46f14143-5170-4760-b3f2-9452ed80bdaf" ID-ISSN="1175-5326" ID-Zenodo-Dep="3450106" ID-ZooBank="3FC57739-88C9-44B7-9632-E452755089A6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568898309288" checkinUser="plazi" docAuthor="Deschodt, Christian M. &amp; Sole, Catherine L." docDate="2019" docId="A4598792C741420546A89600D3B0FEC1" docLanguage="en" docName="zootaxa.4671.1.11.pdf" docOrigin="Zootaxa 4671 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Versicorpus daures Deschodt &amp; Sole 2019, new species" docType="treatment" docUuid="3BAB8003-6334-4F3F-94C0-680BBCF86FF2" docUuidSource="ZooBank" docVersion="9" lastPageNumber="143" masterDocId="5860FFEAC7414200463F900CD757FFE1" masterDocTitle="A new species of Versicorpus Deschodt, Davis &amp; Scholtz, 2011 (Scarabaeidae Scarabaeinae: Byrrhidiini) from the Brandberg Mountain, Namibia" masterLastPageNumber="144" masterPageNumber="139" pageNumber="139" updateTime="1698755846860" updateUser="ExternalLinkService">
<mods:mods id="7640B56C0D86531651447D62086529CA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="17FC3015CC972B8450C19BB2A5EB7805">A new species of Versicorpus Deschodt, Davis &amp; Scholtz, 2011 (Scarabaeidae Scarabaeinae: Byrrhidiini) from the Brandberg Mountain, Namibia</mods:title>
</mods:titleInfo>
<mods:name id="1A3E0089C28F0457B22C048B8FA0405D" type="personal">
<mods:role id="55261B6CB10D44A39233BBEF4BBD6A2D">
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</mods:role>
<mods:namePart id="607A9665102B5BCA118DC55BCC95A595">Deschodt, Christian M.</mods:namePart>
</mods:name>
<mods:name id="7F0F95DEDACA04FF3265B31FE95B0EF5" type="personal">
<mods:role id="0A0FD431C19AF0CA5A85FBC4880DE2D3">
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<mods:namePart id="7A10018116F4FCAEC04F6A8687F2B75D">Sole, Catherine L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C8AF478F6E8C0EB3E72CBF4C0788686E">text</mods:typeOfResource>
<mods:relatedItem id="545144111A4F165C37694A07DFD14752" type="host">
<mods:titleInfo id="CA7D024A21D5344158B467079C6CEE51">
<mods:title id="21B2ED73B8B3461C7E7D08BF43B4305C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="27595C8C38DCFB68AB85E30AF4B8299C">
<mods:date id="1C593A20BD8909058357E0C30B163EC2">2019</mods:date>
<mods:detail id="EE3209D2112CAEE9FB2C30ADF7C50E59" type="pubDate">
<mods:number id="BBBA35CBC3FA454C30F022D11506BD95">2019-09-16</mods:number>
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<mods:number id="9D809DFD83453073138B9FE042837303">4671</mods:number>
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<mods:number id="A83BDBED1B0E68657FE6DA387C0E99D2">1</mods:number>
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<mods:classification id="CC5E5E668DD883235BF42997B27DFA63">journal article</mods:classification>
<mods:identifier id="B010A2BCC9003572156EAF365C1F8965" type="CLB-Dataset">25459</mods:identifier>
<mods:identifier id="855E924B03E92A0F2F6750B958B0A1D1" type="DOI">10.11646/zootaxa.4671.1.11</mods:identifier>
<mods:identifier id="317BE1DBBE45E4874424014E24587A8A" type="GBIF-Dataset">46f14143-5170-4760-b3f2-9452ed80bdaf</mods:identifier>
<mods:identifier id="D0BB213DF0CFCC7188084024071604E3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C947CCFC2E75F558F92FA69F5958FC07" type="Zenodo-Dep">3450106</mods:identifier>
<mods:identifier id="E01CF72092B4C5C0AAA5CE02A5188544" type="ZooBank">3FC57739-88C9-44B7-9632-E452755089A6</mods:identifier>
</mods:mods>
<treatment id="A4598792C741420546A89600D3B0FEC1" ID-GBIF-Taxon="159565868" LSID="urn:lsid:zoobank.org:act:3BAB8003-6334-4F3F-94C0-680BBCF86FF2" httpUri="http://treatment.plazi.org/id/A4598792C741420546A89600D3B0FEC1" lastPageId="5" lastPageNumber="143" pageId="0" pageNumber="139">
<subSubSection id="64EA650FC741420046A89600D5C0F9C7" box="[151,663,1548,1575]" pageId="0" pageNumber="139" type="nomenclature">
<paragraph id="2C4F3684C741420046A89600D5C0F9C7" blockId="0.[151,663,1548,1610]" box="[151,663,1548,1575]" pageId="0" pageNumber="139">
<heading id="770781E8C741420046A89600D5C0F9C7" bold="true" box="[151,663,1548,1575]" fontSize="11" level="1" pageId="0" pageNumber="139" reason="1">
<emphasis id="1E84EA96C741420046A89600D5C0F9C7" bold="true" box="[151,663,1548,1575]" pageId="0" pageNumber="139">
<taxonomicName id="EBF04D07C741420046A89600D6ACF9C6" ID-CoL="5B8QM" authority="Deschodt" authorityName="Deschodt &amp; Sole" authorityYear="2019" box="[151,507,1548,1575]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="daures" status="sp. nov.">
<emphasis id="1E84EA96C741420046A89600D62AF9C6" bold="true" box="[151,381,1548,1575]" italics="true" pageId="0" pageNumber="139">Versicorpus daures</emphasis>
Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="05B757EDC741420044379601D5C0F9C7" box="[520,663,1548,1575]" pageId="0" pageNumber="139" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="64EA650FC741420046A8963CD628F9AB" box="[151,383,1584,1610]" pageId="0" pageNumber="139" type="description">
<paragraph id="2C4F3684C741420046A8963CD628F9AB" blockId="0.[151,663,1548,1610]" box="[151,383,1584,1610]" pageId="0" pageNumber="139">
(
<figureCitation id="B4CB2A01C741420046A0963CD7BAF9AB" box="[159,237,1584,1610]" captionStart="FIGURE 1" captionStartId="1.[151,250,1132,1157]" captionTargetBox="[151,1436,179,1104]" captionTargetId="figure@1.[151,1436,179,1104]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of the live holotype of Versicorpus daures Deschodt, new species walking on granite rock. Scale bar = 2.0 mm. Photo: Hennie De Klerk at Brandberg, 16 April 2019." figureDoi="http://doi.org/10.5281/zenodo.3450108" httpUri="https://zenodo.org/record/3450108/files/figure.png" pageId="0" pageNumber="139">Figs. 1</figureCitation>
,
<figureCitation id="B4CB2A01C741420046C4963CD65EF9AB" box="[251,265,1584,1610]" captionStart="FIGURE 2" captionStartId="2.[151,250,1895,1920]" captionTargetBox="[208,1380,199,1838]" captionTargetId="figure@2.[198,1388,179,1868]" captionTargetPageId="2" captionText="FIGURE 2. Habitus of the holotype of Versicorpus daures Deschodt, new species (A) in comparison with the habitus of the holotype of Versicorpus streyi (Frolov, 2005) (D) with the aedeagus of the holotype of V. daures Deschodt, new species (B, C) and the aedeagus of the holotype of V. streyi (Frolov, 2005) (E, F) in lateral (left) and dorsal (right) view for comparison. Scale bar for habitus = 2.0 mm and scale bar for aedeagi = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3450110" httpUri="https://zenodo.org/record/3450110/files/figure.png" pageId="0" pageNumber="139">2</figureCitation>
A–C, 3, 4)
</paragraph>
</subSubSection>
<subSubSection id="64EA650FC741420046A89674D4C4F853" pageId="0" pageNumber="139" type="materials_examined">
<paragraph id="2C4F3684C741420046A89674D685F93B" blockId="0.[151,1437,1655,2043]" pageId="0" pageNumber="139">
<emphasis id="1E84EA96C741420046A89674D611F973" bold="true" box="[151,326,1655,1682]" pageId="0" pageNumber="139">Type material.</emphasis>
<materialsCitation id="9C983CD9C741420047719674D685F93B" ID-GBIF-Occurrence="2416712606" collectingDate="2019-04-16" collectionCode="SMWN" collectorName="C. M. Deschodt" elevation="839" latitude="-21.105" location="Brandberg Mountain" longLatPrecision="75" longitude="14.691" pageId="0" pageNumber="139" specimenCount="1" typeStatus="holotype">
<typeStatus id="F34B8826C741420047719674D6E1F973" box="[334,438,1656,1682]" pageId="0" pageNumber="139" type="holotype">Holotype</typeStatus>
♂: “
<location id="292F605FC741420047D29674D58CF973" LSID="urn:lsid:plazi:treatment:A4598792C741420546A89600D3B0FEC1:292F605FC741420047D29674D58CF973" box="[493,731,1656,1682]" latitude="-21.105" longLatPrecision="75" longitude="14.691" name="Brandberg Mountain" pageId="0" pageNumber="139">Brandberg Mountain</location>
|
<geoCoordinate id="49C45043C741420044CF967BD40EF973" box="[752,857,1655,1682]" degrees="21.105" direction="south" orientation="latitude" pageId="0" pageNumber="139" precision="55" value="-21.105">S21.105°</geoCoordinate>
<geoCoordinate id="49C45043C7414200455F9674D49DF973" box="[864,970,1656,1682]" degrees="14.691" direction="east" orientation="longitude" pageId="0" pageNumber="139" precision="55" value="14.691">E14.691°</geoCoordinate>
<quantity id="EB089B61C741420045ED9674D346F973" box="[978,1041,1656,1682]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="0" pageNumber="139" unit="m" value="839.0">
<elevation id="A7DDD1B7C741420045ED9674D346F973" box="[978,1041,1656,1682]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="0" pageNumber="139" unit="m" value="839.0">839m</elevation>
</quantity>
|
<date id="584E1044C741420042179674D3F7F973" box="[1064,1184,1656,1682]" pageId="0" pageNumber="139" value="2019-04-16">
<collectingDate id="480AE9ACC741420042179674D3F7F973" box="[1064,1184,1656,1682]" pageId="0" pageNumber="139" value="2019-04-16">16.iv.2019</collectingDate>
</date>
,
<collectorName id="81055352C741420042929674D20FF973" box="[1197,1368,1656,1682]" pageId="0" pageNumber="139">C.M. Deschodt</collectorName>
|| Ex- cavated live from | accumulated silt under a | layer of hyrax dung” (
<collectionCode id="4AE1AE41C741420045AF9690D4B9F957" box="[912,1006,1692,1718]" country="Namibia" httpUri="http://grbio.org/cool/s32g-pczn" name="State Museum" pageId="0" pageNumber="139">SMWN</collectionCode>
). The specimen was preserved in 99% ethanol for molecular study.
</materialsCitation>
</paragraph>
<paragraph id="2C4F3684C741420046F896EFD4D8F853" blockId="0.[151,1437,1655,2043]" pageId="0" pageNumber="139">
<materialsCitation id="9C983CD9C741420046F896EFD781F8A7" ID-GBIF-Occurrence="2422755396" collectingDate="2019-04-16" collectionCode="UPSA" collectorName="C. M. Deschodt" elevation="839" latitude="-21.105" location="Disarticulated" longLatPrecision="75" longitude="14.691" municipality="One" pageId="0" pageNumber="139" specimenCount="1">
<emphasis id="1E84EA96C741420046F896EFD560F91F" bold="true" box="[199,567,1763,1790]" pageId="0" pageNumber="139">Additional material examined.</emphasis>
<location id="292F605FC7414200440096E8D58BF91F" LSID="urn:lsid:plazi:treatment:A4598792C741420546A89600D3B0FEC1:292F605FC7414200440096E8D58BF91F" box="[575,732,1764,1790]" latitude="-21.105" longLatPrecision="75" longitude="14.691" municipality="One" name="Disarticulated" pageId="0" pageNumber="139">Disarticulated</location>
parts of different specimens that include: elytra, pronota with and without front legs, a single pygidium and one complete set of abdominal sternites, all mounted together on cards
</materialsCitation>
.
<materialsCitation id="9C983CD9C741420046DF9720D384F88B" ID-GBIF-Occurrence="2422755397" collectingDate="2019-04-16" collectionCode="UPSA" collectorName="C. M. Deschodt" elevation="839" latitude="-21.105" location="One" longLatPrecision="75" longitude="14.691" municipality="One" pageId="0" pageNumber="139" specimenCount="1">
<collectingMunicipality id="CC2BACFEC741420046DF9720D658F8A7" box="[224,271,1836,1862]" pageId="0" pageNumber="139">One</collectingMunicipality>
lot from the same locality as the holotype with labels: “Brandberg Mountain |
<geoCoordinate id="49C45043C741420042419727D3B1F8A7" box="[1150,1254,1835,1862]" degrees="21.105" direction="south" orientation="latitude" pageId="0" pageNumber="139" precision="55" value="-21.105">S21.105°</geoCoordinate>
<geoCoordinate id="49C45043C741420042D39720D201F8A7" box="[1260,1366,1836,1862]" degrees="14.691" direction="east" orientation="longitude" pageId="0" pageNumber="139" precision="55" value="14.691">E14.691°</geoCoordinate>
<quantity id="EB089B61C741420043639720D2CBF8A7" box="[1372,1436,1836,1862]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="0" pageNumber="139" unit="m" value="839.0">
<elevation id="A7DDD1B7C741420043639720D2CBF8A7" box="[1372,1436,1836,1862]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="0" pageNumber="139" unit="m" value="839.0">839m</elevation>
</quantity>
|
<date id="584E1044C7414200469A975CD64AF88B" box="[165,285,1872,1898]" pageId="0" pageNumber="139" value="2019-04-16">
<collectingDate id="480AE9ACC7414200469A975CD64AF88B" box="[165,285,1872,1898]" pageId="0" pageNumber="139" value="2019-04-16">16.iv.2019</collectingDate>
</date>
,
<collectorName id="81055352C74142004715975CD681F88B" box="[298,470,1872,1898]" pageId="0" pageNumber="139">C.M. Deschodt</collectorName>
|| Excavated from | accumulated silt under a | layer of hyrax dung”
</materialsCitation>
<materialsCitation id="9C983CD9C741420042E4975CD4D8F853" ID-GBIF-Occurrence="2422755398" collectingDate="2019-04-16" collectionCode="UPSA" collectorName="C. M. Deschodt" elevation="839" latitude="-21.105" location="One" longLatPrecision="75" longitude="14.691" municipality="One" pageId="0" pageNumber="139" specimenCount="1">
and a second lot: “Brandberg Mountain |
<geoCoordinate id="49C45043C7414200479D977FD55DF86F" box="[418,522,1907,1934]" degrees="21.075" direction="south" orientation="latitude" pageId="0" pageNumber="139" precision="55" value="-21.075">S21.075°</geoCoordinate>
<geoCoordinate id="49C45043C7414200442E9778D52DF86F" box="[529,634,1908,1934]" degrees="14.677" direction="east" orientation="longitude" pageId="0" pageNumber="139" precision="55" value="14.677">E14.677°</geoCoordinate>
<quantity id="EB089B61C741420044BD977FD596F86F" box="[642,705,1907,1934]" metricMagnitude="2" metricUnit="m" metricValue="5.58" pageId="0" pageNumber="139" unit="m" value="558.0">
<elevation id="A7DDD1B7C741420044BD977FD596F86F" box="[642,705,1907,1934]" metricMagnitude="2" metricUnit="m" metricValue="5.58" pageId="0" pageNumber="139" unit="m" value="558.0">558m</elevation>
</quantity>
|
<date id="584E1044C741420044E99778D419F86F" box="[726,846,1908,1934]" pageId="0" pageNumber="139" value="2019-04-16">
<collectingDate id="480AE9ACC741420044E99778D419F86F" box="[726,846,1908,1934]" pageId="0" pageNumber="139" value="2019-04-16">16.iv.2019</collectingDate>
</date>
,
<collectorName id="81055352C741420045659778D352F86F" box="[858,1029,1908,1934]" pageId="0" pageNumber="139">C.M. Deschodt</collectorName>
|| Excavated from | accumulated silt under a | layer of hyrax dung” about three kilometres away (
<collectionCode id="4AE1AE41C741420045069794D4D0F853" box="[825,903,1944,1970]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="0" pageNumber="139">UPSA</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph id="2C4F3684C741420045B09794D4C4F853" blockId="0.[151,1437,1655,2043]" box="[911,915,1944,1970]" pageId="0" pageNumber="139">.</paragraph>
</subSubSection>
<subSubSection id="64EA650FC741420446F897B7D499F963" lastPageId="4" lastPageNumber="143" pageId="0" pageNumber="139" type="description">
<paragraph id="2C4F3684C741420046F897B7D630F81A" blockId="0.[151,1437,1655,2043]" pageId="0" pageNumber="139">
<emphasis id="1E84EA96C741420046F897B7D5E6F837" bold="true" box="[199,689,1979,2006]" pageId="0" pageNumber="139">
Description of the
<typeStatus id="F34B8826C7414200479697B0D546F837" box="[425,529,1980,2006]" pageId="0" pageNumber="139" type="holotype">holotype</typeStatus>
, male. Body:
</emphasis>
9.1 (maximum length with head extended) x
<quantity id="EB089B61C741420042FE97B7D24EF836" box="[1217,1305,1979,2007]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="0" pageNumber="139" unit="mm" value="5.5">5.5 mm</quantity>
(maximum width over elytra).
</paragraph>
<caption id="788F660CC740420146A89460D5BCFB49" ID-DOI="http://doi.org/10.5281/zenodo.3450108" ID-Zenodo-Dep="3450108" httpUri="https://zenodo.org/record/3450108/files/figure.png" pageId="1" pageNumber="140" startId="1.[151,250,1132,1157]" targetBox="[151,1436,179,1104]" targetPageId="1">
<paragraph id="2C4F3684C740420146A89460D5BCFB49" blockId="1.[151,1437,1132,1193]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146A89460D644FB64" bold="true" box="[151,275,1132,1157]" pageId="1" pageNumber="140">FIGURE 1.</emphasis>
Habitus of the live holotype of
<taxonomicName id="EBF04D07C7404201446B9461D429FB64" authority="Deschodt" authorityName="Deschodt &amp; Sole" authorityYear="2019" box="[596,894,1132,1157]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="daures" status="sp. nov.">
<emphasis id="1E84EA96C7404201446B9461D441FB65" box="[596,790,1132,1156]" italics="true" pageId="1" pageNumber="140">Versicorpus daures</emphasis>
Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="05B757EDC740420145B79461D356FB64" box="[904,1025,1132,1157]" pageId="1" pageNumber="140" rank="species">new species</taxonomicNameLabel>
walking on granite rock. Scale bar = 2.0 mm. Photo: Hennie De Klerk at Brandberg, 16 April 2019.
</paragraph>
</caption>
<paragraph id="2C4F3684C740420146F894DBD630FABC" blockId="1.[151,1437,1239,2021]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146F894DBD659FB10" bold="true" box="[199,270,1239,1265]" pageId="1" pageNumber="140">Head.</emphasis>
Clypeus bidentate, with two small teeth that are well separated; clypeal margin between teeth very slightly sinuate. Surface clearly punctate with small punctures that are separated by less than one puncture diameter. One very small tooth on lower clypeal margin between dorsal teeth. Clypeogenal suture indistinct and forming a fairly straight line.
</paragraph>
<paragraph id="2C4F3684C740420146F8956BD676FA28" blockId="1.[151,1437,1239,2021]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146F8956BD610FA60" bold="true" box="[199,327,1383,1409]" pageId="1" pageNumber="140">Pronotum.</emphasis>
Convex, surface with small oval to round punctures covering the pronotal surface, punctures separated by less than one puncture diameter; most punctures with very short associated setae; clear puncture-free midline present.
</paragraph>
<paragraph id="2C4F3684C740420146F895DFD79EF9B8" blockId="1.[151,1437,1239,2021]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146F895DFD64DFA0C" bold="true" box="[199,282,1491,1517]" pageId="1" pageNumber="140">Elytra.</emphasis>
Convex, slightly elongated, surface covered with microscopically fine granules giving velutinous appearance under
<date id="584E1044C7404201477895FBD626F9F0" box="[327,369,1527,1553]" pageId="1" pageNumber="140" value="1940-10">40x</date>
or higher magnification. Striae shallow but visible, not punctate. Interstriae with punctures with associated short yellow setae arranged in fairly straight double lines except for the first which only has a single line.
</paragraph>
<paragraph id="2C4F3684C740420146F8966ED26CF924" blockId="1.[151,1437,1239,2021]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146F8966ED649F99C" bold="true" box="[199,286,1634,1661]" pageId="1" pageNumber="140">Sterna.</emphasis>
Prosternum with small and well separated punctures; with short yellow irregularly spaced setae. Meso- and metasternum fused, mesometasternal suture indistinct and straight. Surface of mesosternum shagreened, faint, medium-sized shallow punctures present. Surface of metasternum shagreened with a few small punctures.
</paragraph>
<paragraph id="2C4F3684C740420146F896C2D435F8B4" blockId="1.[151,1437,1239,2021]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146F896C2D667F908" bold="true" box="[199,304,1742,1769]" pageId="1" pageNumber="140">Protibia.</emphasis>
Internal margin approximately straight, with short yellow setae and lacking denticles. Tibial apex blunt with a small conical slightly outward curving spur. Tarsi attached under spur close to inner corner of tibia. Setae on internal margin of tarsi short. Apical external margin of protibiae with three teeth with the medial tooth being the smallest. Basal two thirds of external margin serrated.
</paragraph>
<paragraph id="2C4F3684C740420146F89752D4DFF87C" blockId="1.[151,1437,1239,2021]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146F89752D612F898" bold="true" box="[199,325,1886,1913]" pageId="1" pageNumber="140">Mesotibia.</emphasis>
Curving slightly inward, with smooth margins and five rows of short yellow setae. Apically with two short spurs. Tarsi with short yellow setae, ending in two claws.
</paragraph>
<paragraph id="2C4F3684C740420146F897AAD321F804" blockId="1.[151,1437,1239,2021]" pageId="1" pageNumber="140">
<emphasis id="1E84EA96C740420146F897AAD66BF820" bold="true" box="[199,316,1958,1985]" pageId="1" pageNumber="140">Metatibia</emphasis>
. Fairly straight, with internal margin saw-toothed along entire length, external margin smooth; five rows of short yellow setae present. Tarsi with short yellow setae, ending with two claws.
</paragraph>
<caption id="788F660CC743420246A8976BD452F80A" ID-DOI="http://doi.org/10.5281/zenodo.3450110" ID-Zenodo-Dep="3450110" httpUri="https://zenodo.org/record/3450110/files/figure.png" pageId="2" pageNumber="141" startId="2.[151,250,1895,1920]" targetBox="[208,1380,199,1838]" targetPageId="2">
<paragraph id="2C4F3684C743420246A8976BD452F80A" blockId="2.[151,1437,1895,2028]" pageId="2" pageNumber="141">
<emphasis id="1E84EA96C743420246A8976BD643F861" bold="true" box="[151,276,1895,1920]" pageId="2" pageNumber="141">FIGURE 2.</emphasis>
Habitus of the holotype of
<taxonomicName id="EBF04D07C7434202440B9765D436F861" authority="Deschodt" authorityName="Deschodt &amp; Sole" authorityYear="2019" box="[564,865,1895,1920]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="daures" status="sp. nov.">
<emphasis id="1E84EA96C7434202440B9765D5A0F861" box="[564,759,1896,1920]" italics="true" pageId="2" pageNumber="141">Versicorpus daures</emphasis>
Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="05B757EDC743420245529764D4BFF861" box="[877,1000,1895,1920]" pageId="2" pageNumber="141" rank="species">new species</taxonomicNameLabel>
(A) in comparison with the habitus of the holotype of
<taxonomicName id="EBF04D07C7434202472D9781D536F845" authority="(Frolov, 2005)" authorityName="Frolov" authorityYear="2005" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[274,609,1931,1956]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="streyi">
<emphasis id="1E84EA96C7434202472D9781D69FF845" box="[274,456,1933,1956]" italics="true" pageId="2" pageNumber="141">Versicorpus streyi</emphasis>
(Frolov, 2005)
</taxonomicName>
(D) with the aedeagus of the holotype of
<taxonomicName id="EBF04D07C743420242349781D386F845" authority="Deschodt" authorityName="Deschodt &amp; Sole" authorityYear="2019" box="[1035,1233,1931,1956]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="daures" status="sp. nov.">
<emphasis id="1E84EA96C743420242349781D348F845" box="[1035,1055,1933,1956]" italics="true" pageId="2" pageNumber="141">V.</emphasis>
<emphasis id="1E84EA96C7434202421B9780D33FF845" box="[1060,1128,1932,1956]" italics="true" pageId="2" pageNumber="141">daures</emphasis>
Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="05B757EDC743420242E39780D201F845" box="[1244,1366,1931,1956]" pageId="2" pageNumber="141" rank="species">new species</taxonomicNameLabel>
(B, C) and the aedeagus of the holotype of
<taxonomicName id="EBF04D07C7434202443997BDD5B8F829" authority="(Frolov, 2005)" authorityName="Frolov" authorityYear="2005" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[518,751,1967,1992]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="streyi">
<emphasis id="1E84EA96C7434202443997BDD501F829" box="[518,598,1969,1992]" italics="true" pageId="2" pageNumber="141">V. streyi</emphasis>
(Frolov, 2005)
</taxonomicName>
(E, F) in lateral (left) and dorsal (right) view for comparison. Scale bar for habitus = 2.0 mm and scale bar for aedeagi = 1.0 mm.
</paragraph>
</caption>
<caption id="788F660CC742420346A89743D48AF84E" ID-DOI="http://doi.org/10.5281/zenodo.3450118" ID-Zenodo-Dep="3450118" httpUri="https://zenodo.org/record/3450118/files/figure.png" pageId="3" pageNumber="142" startId="3.[151,250,1871,1896]" targetBox="[249,1339,201,1846]" targetPageId="3">
<paragraph id="2C4F3684C742420346A89743D48AF84E" blockId="3.[151,1436,1869,1968]" pageId="3" pageNumber="142">
<emphasis id="1E84EA96C742420346A89743D642F889" bold="true" box="[151,277,1871,1896]" pageId="3" pageNumber="142">FIGURE 3.</emphasis>
Map indicating the Brandberg and the type locality of
<taxonomicName id="EBF04D07C74242034568975CD3D1F889" authority="Deschodt" authorityName="Deschodt &amp; Sole" authorityYear="2019" box="[855,1158,1871,1896]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="daures" status="sp. nov.">
<emphasis id="1E84EA96C74242034568975CD34CF886" box="[855,1051,1871,1895]" italics="true" pageId="3" pageNumber="142">Versicorpus daures</emphasis>
Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="05B757EDC742420342AD975CD259F889" box="[1170,1294,1871,1896]" pageId="3" pageNumber="142" rank="species">new species</taxonomicNameLabel>
(
<emphasis id="1E84EA96C742420343209741D260F887" bold="true" box="[1311,1335,1869,1894]" pageId="3" pageNumber="142">*</emphasis>
) with the distribution of all other Byrrhidiini (
<emphasis id="1E84EA96C74242034438977DD54CF86B" bold="true" box="[519,539,1905,1930]" pageId="3" pageNumber="142">•</emphasis>
), clearly following the Namibian section of the great escarpment of southern Africa. For detailed distribution of the other Byrrhidiini species see
<bibRefCitation id="48614B75C742420344EE979BD48EF84E" author="Deschodt, C. M. &amp; Davis, A. L. V." box="[721,985,1943,1968]" pageId="3" pageNumber="142" pagination="1" refId="ref2520" refString="Deschodt, C. M. &amp; Davis, A. L. V. (2018) New deltochiline (Coleoptera: Scarabaeidae: Scarabaeinae) taxa associated with hyrax dung in arid south western Namibia. Zootaxa, 4450 (1), 066 - 076. https: // doi. org / 10.11646 / zootaxa. 4450.1.4" type="journal article" year="2018">Deschodt &amp; Davis (2018)</bibRefCitation>
.
</paragraph>
</caption>
<caption id="788F660CC745420446A89424D539FB84" ID-DOI="http://doi.org/10.5281/zenodo.3450122" ID-Zenodo-Dep="3450122" httpUri="https://zenodo.org/record/3450122/files/figure.png" pageId="4" pageNumber="143" startId="4.[151,250,1064,1089]" targetBox="[151,1436,179,1037]" targetPageId="4">
<paragraph id="2C4F3684C745420446A89424D539FB84" blockId="4.[151,1437,1064,1125]" pageId="4" pageNumber="143">
<emphasis id="1E84EA96C745420446A89424D644FBA0" bold="true" box="[151,275,1064,1089]" pageId="4" pageNumber="143">FIGURE 4.</emphasis>
Habitat and collection site (with the arrow indicating the exact spot) in the Brandberg Mountain, Namibia. Photo: Hennie De Klerk at Brandberg, 16 April 2019.
</paragraph>
</caption>
<paragraph id="2C4F3684C745420446F8949ED423FB4F" blockId="4.[151,1437,1170,2026]" box="[199,884,1170,1198]" pageId="4" pageNumber="143">
<emphasis id="1E84EA96C745420446F8949ED66DFB4C" bold="true" box="[199,314,1170,1197]" pageId="4" pageNumber="143">Pygidium</emphasis>
. Surface shagreened with large shallow punctures.
</paragraph>
<paragraph id="2C4F3684C745420446F894BAD40AFAFB" blockId="4.[151,1437,1170,2026]" pageId="4" pageNumber="143">
<emphasis id="1E84EA96C745420446F894BAD616FB30" bold="true" box="[199,321,1206,1233]" pageId="4" pageNumber="143">Aedeagus.</emphasis>
Same general plan as for the other known
<taxonomicName id="EBF04D07C7454204452694B4D4CAFB30" authorityName="Deschodt, Davis &amp; Scholtz" authorityYear="2011" box="[793,925,1208,1233]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="1E84EA96C7454204452694B4D4CAFB30" box="[793,925,1208,1233]" italics="true" pageId="4" pageNumber="143">Versicorpus</emphasis>
</taxonomicName>
species. Parameres asymmetrical at base with the left paramere forming a lobe slightly covering the base of the right one, nearly symmetrical towards apex; rows of setae dorsally on distal half of parameres (
<figureCitation id="B4CB2A01C745420444AF94F3D407FAFB" box="[656,848,1279,1306]" captionStart="FIGURE 2" captionStartId="2.[151,250,1895,1920]" captionTargetBox="[208,1380,199,1838]" captionTargetId="figure@2.[198,1388,179,1868]" captionTargetPageId="2" captionText="FIGURE 2. Habitus of the holotype of Versicorpus daures Deschodt, new species (A) in comparison with the habitus of the holotype of Versicorpus streyi (Frolov, 2005) (D) with the aedeagus of the holotype of V. daures Deschodt, new species (B, C) and the aedeagus of the holotype of V. streyi (Frolov, 2005) (E, F) in lateral (left) and dorsal (right) view for comparison. Scale bar for habitus = 2.0 mm and scale bar for aedeagi = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3450110" httpUri="https://zenodo.org/record/3450110/files/figure.png" pageId="4" pageNumber="143">Figs. 2B, C, E, F</figureCitation>
).
</paragraph>
<paragraph id="2C4F3684C745420446F8952ED34DFA67" blockId="4.[151,1437,1170,2026]" pageId="4" pageNumber="143">
<emphasis id="1E84EA96C745420446F8952ED61AFADC" bold="true" box="[199,333,1314,1341]" pageId="4" pageNumber="143">Variability.</emphasis>
Except for a very small size difference and that most specimens were quite worn out, there was very little other variation observed between the
<typeStatus id="F34B8826C745420444FF954BD475FA80" box="[704,802,1351,1377]" pageId="4" pageNumber="143" type="holotype">holotype</typeStatus>
and the disarticulated specimens. All pronota had very worn out front legs but still had the short setae present at almost every puncture.
</paragraph>
<paragraph id="2C4F3684C745420446F89582D499F963" blockId="4.[151,1437,1170,2026]" pageId="4" pageNumber="143">
<emphasis id="1E84EA96C745420446F89582D698FA48" bold="true" box="[199,463,1422,1449]" pageId="4" pageNumber="143">Differential diagnosis.</emphasis>
This new species clearly belongs to the genus
<taxonomicName id="EBF04D07C745420445D8959CD33CFA48" authorityName="Deschodt, Davis &amp; Scholtz" authorityYear="2011" box="[999,1131,1424,1449]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="1E84EA96C745420445D8959CD33CFA48" box="[999,1131,1424,1449]" italics="true" pageId="4" pageNumber="143">Versicorpus</emphasis>
</taxonomicName>
by virtue of the somewhat elongated body and the basal part of front tibiae being serrated and the saw-toothed internal margin of the hind tibiae and very short clypeal horns. It is closest in appearance to
<taxonomicName id="EBF04D07C745420445BF95D4D3A1FA10" authority="(Frolov, 2005)" authorityName="Frolov" authorityYear="2005" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[896,1270,1494,1521]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="streyi">
<emphasis id="1E84EA96C745420445BF95D4D31DFA10" box="[896,1098,1496,1521]" italics="true" pageId="4" pageNumber="143">Versicorpus streyi</emphasis>
(Frolov, 2005)
</taxonomicName>
but can easily be distinguished from it by the presence of short setae on the pronotum (visible only under a microscope with
<date id="584E1044C7454204434D95F7D2CBF9F4" box="[1394,1436,1531,1557]" pageId="4" pageNumber="143" value="1920-10">20x</date>
magnification and higher), whereas
<emphasis id="1E84EA96C74542044415962CD56CF9DB" box="[554,571,1568,1594]" italics="true" pageId="4" pageNumber="143">Ľ</emphasis>
.
<taxonomicName id="EBF04D07C74542044476962CD5D1F9D8" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[585,646,1568,1593]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="streyi">
<emphasis id="1E84EA96C74542044476962CD5D1F9D8" box="[585,646,1568,1593]" italics="true" pageId="4" pageNumber="143">streyi</emphasis>
</taxonomicName>
is glabrous. It is also smaller than
<emphasis id="1E84EA96C74542044235962CD34CF9DB" box="[1034,1051,1568,1594]" italics="true" pageId="4" pageNumber="143">Ľ</emphasis>
.
<taxonomicName id="EBF04D07C74542044216962CD331F9D8" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[1065,1126,1568,1593]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="streyi">
<emphasis id="1E84EA96C74542044216962CD331F9D8" box="[1065,1126,1568,1593]" italics="true" pageId="4" pageNumber="143">streyi</emphasis>
</taxonomicName>
(
<figureCitation id="B4CB2A01C7454204424A9613D3ABF9DB" box="[1141,1276,1567,1594]" captionStart="FIGURE 2" captionStartId="2.[151,250,1895,1920]" captionTargetBox="[208,1380,199,1838]" captionTargetId="figure@2.[198,1388,179,1868]" captionTargetPageId="2" captionText="FIGURE 2. Habitus of the holotype of Versicorpus daures Deschodt, new species (A) in comparison with the habitus of the holotype of Versicorpus streyi (Frolov, 2005) (D) with the aedeagus of the holotype of V. daures Deschodt, new species (B, C) and the aedeagus of the holotype of V. streyi (Frolov, 2005) (E, F) in lateral (left) and dorsal (right) view for comparison. Scale bar for habitus = 2.0 mm and scale bar for aedeagi = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3450110" httpUri="https://zenodo.org/record/3450110/files/figure.png" pageId="4" pageNumber="143">Figs. 2A, D</figureCitation>
). The setae on the aedeagus of
<taxonomicName id="EBF04D07C745420447709648D6E1F9BC" authority="Deschodt &amp; Sole, 2019" authorityName="Deschodt &amp; Sole" authorityYear="2019" box="[335,438,1603,1629]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="daures" status="sp. nov.">
<emphasis id="1E84EA96C745420447709648D6E1F9BC" box="[335,438,1603,1629]" italics="true" pageId="4" pageNumber="143">V. daures</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05B757EDC745420447819648D512F9BC" box="[446,581,1603,1630]" pageId="4" pageNumber="143" rank="species">new species</taxonomicNameLabel>
are shorter and less dense than on the aedeagus of
<emphasis id="1E84EA96C745420442B69648D3CDF9BF" box="[1161,1178,1604,1630]" italics="true" pageId="4" pageNumber="143">Ľ</emphasis>
.
<taxonomicName id="EBF04D07C745420442969648D3B1F9BC" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[1193,1254,1604,1629]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="streyi">
<emphasis id="1E84EA96C745420442969648D3B1F9BC" box="[1193,1254,1604,1629]" italics="true" pageId="4" pageNumber="143">streyi</emphasis>
</taxonomicName>
and the apex of the parameres of
<emphasis id="1E84EA96C745420447669664D63DF963" box="[345,362,1640,1666]" italics="true" pageId="4" pageNumber="143">Ľ</emphasis>
.
<taxonomicName id="EBF04D07C745420447479664D6E2F960" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[376,437,1640,1665]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="streyi">
<emphasis id="1E84EA96C745420447479664D6E2F960" box="[376,437,1640,1665]" italics="true" pageId="4" pageNumber="143">streyi</emphasis>
</taxonomicName>
are also more elongated (see
<figureCitation id="B4CB2A01C7454204453D966BD495F963" box="[770,962,1639,1666]" captionStart="FIGURE 2" captionStartId="2.[151,250,1895,1920]" captionTargetBox="[208,1380,199,1838]" captionTargetId="figure@2.[198,1388,179,1868]" captionTargetPageId="2" captionText="FIGURE 2. Habitus of the holotype of Versicorpus daures Deschodt, new species (A) in comparison with the habitus of the holotype of Versicorpus streyi (Frolov, 2005) (D) with the aedeagus of the holotype of V. daures Deschodt, new species (B, C) and the aedeagus of the holotype of V. streyi (Frolov, 2005) (E, F) in lateral (left) and dorsal (right) view for comparison. Scale bar for habitus = 2.0 mm and scale bar for aedeagi = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3450110" httpUri="https://zenodo.org/record/3450110/files/figure.png" pageId="4" pageNumber="143">Figs. 2B, C, E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="64EA650FC745420446F89687D591F928" pageId="4" pageNumber="143" type="etymology">
<paragraph id="2C4F3684C745420446F89687D591F928" blockId="4.[151,1437,1170,2026]" pageId="4" pageNumber="143">
<emphasis id="1E84EA96C745420446F89687D619F944" bold="true" box="[199,334,1675,1701]" pageId="4" pageNumber="143">Etymology.</emphasis>
The name Dâures is a noun and in the local Damara people’s language it is the name for the Brandberg Mountain which means “burning mountain”.
</paragraph>
</subSubSection>
<subSubSection id="64EA650FC745420546F896DED3B0FEC1" lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143" type="distribution">
<paragraph id="2C4F3684C745420546F896DED3B0FEC1" blockId="4.[151,1437,1170,2026]" lastBlockId="5.[151,1436,153,288]" lastPageId="5" lastPageNumber="144" pageId="4" pageNumber="143">
<emphasis id="1E84EA96C745420446F896DED484F90C" bold="true" box="[199,979,1746,1773]" pageId="4" pageNumber="143">Distribution, ecology, conservation status, and potential threats.</emphasis>
This new species is currently the most northerly recorded of all the known Byrrhidiini (
<figureCitation id="B4CB2A01C7454204448F96FBD5A3F8F0" box="[688,756,1783,1809]" captionStart="FIGURE 3" captionStartId="3.[151,250,1871,1896]" captionTargetBox="[249,1339,201,1846]" captionTargetId="figure@3.[249,1339,201,1846]" captionTargetPageId="3" captionText="FIGURE 3. Map indicating the Brandberg and the type locality of Versicorpus daures Deschodt, new species (*) with the distribution of all other Byrrhidiini (•), clearly following the Namibian section of the great escarpment of southern Africa. For detailed distribution of the other Byrrhidiini species see Deschodt &amp; Davis (2018)." figureDoi="http://doi.org/10.5281/zenodo.3450118" httpUri="https://zenodo.org/record/3450118/files/figure.png" pageId="4" pageNumber="143">Fig. 3</figureCitation>
). However it is anticipated that there might be further species to the north in suitable habitats. The
<typeStatus id="F34B8826C745420444159717D5DBF8D4" box="[554,652,1819,1845]" pageId="4" pageNumber="143" type="holotype">holotype</typeStatus>
and disarticulated body parts were collected between boulders (
<figureCitation id="B4CB2A01C7454204436E9717D2C3F8D4" box="[1361,1428,1819,1845]" captionStart="FIGURE 4" captionStartId="4.[151,250,1064,1089]" captionTargetBox="[151,1436,179,1037]" captionTargetId="figure@4.[151,1436,179,1037]" captionTargetPageId="4" captionText="FIGURE 4. Habitat and collection site (with the arrow indicating the exact spot) in the Brandberg Mountain, Namibia. Photo: Hennie De Klerk at Brandberg, 16 April 2019." figureDoi="http://doi.org/10.5281/zenodo.3450122" httpUri="https://zenodo.org/record/3450122/files/figure.png" pageId="4" pageNumber="143">Fig. 4</figureCitation>
) in a relatively small area against the lower slopes of the Brandberg, but the species might also occur higher up and probably all around the circumference of the mountain. As in all the other species of Byrrhidiini the live specimen was found amongst hyrax dung pellets which are probably used as a sustainable food source. It appears that the preferred dwelling area of specimens is deep in between boulders where the accumulated silt stays moist for much longer after rain. This species may be restricted to the isolated Brandberg massif in
<collectingCountry id="54E77614C7454204427597C3D3F9F808" box="[1098,1198,1999,2025]" name="Namibia" pageId="4" pageNumber="143">Namibia</collectingCountry>
. As this is a declared National Monument of
<collectingCountry id="54E77614C744420547989095D550FF52" box="[423,519,153,179]" name="Namibia" pageId="5" pageNumber="144">Namibia</collectingCountry>
(
<bibRefCitation id="48614B75C744420544279095D5CAFF52" author="Voigt, A." box="[536,669,153,179]" pageId="5" pageNumber="144" refId="ref2657" refString="Voigt, A. (2004) National Monuments in Namibia: An Inventory of Proclaimed National Monuments in the Republic of Namibia. Gamsberg Macmillan Publishers, Windhoek, 252 pp." type="book" year="2004">Voigt 2004</bibRefCitation>
), the hyrax colonies should remain protected, while anthropogenic activities should be limited and thus the new species is probably not facing any current threats. However, it should be assessed as Data Deficient (DD) as the true extent of occurrence remains unknown (this species could encompass a geographic area as small as or smaller than 450 square kilometres (main area of the Brandberg)).
</paragraph>
</subSubSection>
</treatment>
</document>
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