treatments-xml/data/A4/42/F6/A442F619FFC5CA68FF2CFB58739DFC7C.xml
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<document id="C7B1AC23BAA18D2B42725F80B0AA62D8" ID-CLB-Dataset="30126" ID-DOI="10.11646/zootaxa.4418.2.1" ID-GBIF-Dataset="743a03e6-c13d-416f-b2fe-77434c2ab1c8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242733" ID-ZooBank="9A816F34-E39A-4AC2-88A1-3D320A6C2848" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525676165405" checkinUser="plazi" docAuthor="Zhou, Hu-Ting, Lelej, Arkady S., Williams, Kevin A. &amp; Liu, Jing-Xian" docDate="2018" docId="A442F619FFC5CA68FF2CFB58739DFC7C" docLanguage="en" docName="zootaxa.4418.2.1.pdf.imf" docOrigin="Zootaxa 4418 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zavatilla Tsuneki 1993" docType="treatment" docVersion="8" lastPageNumber="103" masterDocId="587B8E61FFC4CA6AFFBBFF887243FFFC" masterDocTitle="Revision of the Oriental genus Zavatilla Tsuneki (Hymenoptera, Mutillidae, Trogaspidiini), with descriptions of two new species" masterLastPageNumber="120" masterPageNumber="101" pageNumber="102" updateTime="1698543474477" updateUser="ExternalLinkService">
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<mods:title id="17F996DFBB37AEAD94FC9B4877E35F77">Revision of the Oriental genus Zavatilla Tsuneki (Hymenoptera, Mutillidae, Trogaspidiini), with descriptions of two new species</mods:title>
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<mods:namePart id="134B7C401AA5EA4CF345EE4AD9976814">Zhou, Hu-Ting</mods:namePart>
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<mods:namePart id="5FCEF165697871E1E25BEA86069DF456">Lelej, Arkady S.</mods:namePart>
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<mods:namePart id="8DC47D45A111ECABDA8C63838DB1DD09">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="C9B473852BAA331E9FC4051BC1AE71DE">Liu, Jing-Xian</mods:namePart>
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<mods:date id="3089FDB95E4D97C2AD6C1D4711D53E45">2018</mods:date>
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<subSubSection id="64F11484FFC5CA6BFF2CFB58704EFB16" box="[151,525,1231,1258]" pageId="1" pageNumber="102" type="nomenclature">
<paragraph id="2C54470FFFC5CA6BFF2CFB58704EFB16" blockId="1.[151,525,1231,1258]" box="[151,525,1231,1258]" pageId="1" pageNumber="102">
<heading id="771CF063FFC5CA6BFF2CFB58704EFB16" bold="true" box="[151,525,1231,1258]" fontSize="11" level="1" pageId="1" pageNumber="102" reason="1">
<emphasis id="1E9F9B1DFFC5CA6BFF2CFB58704EFB16" bold="true" box="[151,525,1231,1258]" pageId="1" pageNumber="102">
Genus
<taxonomicName id="EBEB3C8CFFC5CA6BFF56FB47704EFB16" authority="Tsuneki, 1993" authorityName="Tsuneki" authorityYear="1993" box="[237,525,1231,1258]" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="1E9F9B1DFFC5CA6BFF56FB47731AFB15" bold="true" box="[237,345,1231,1257]" italics="true" pageId="1" pageNumber="102">Zavatilla</emphasis>
<bibRefCitation id="487A3AFEFFC5CA6BFEDAFB58704EFB16" author="Tsuneki, K." box="[353,525,1232,1258]" journalOrPublisher="Special Publications of the Japan Hymenopterists Association" pageId="1" pageNumber="102" pagination="39 50" part="41" refId="ref10470" refString="Tsuneki, K. (1993) On some Taiwanese Mutillidae, collected in 1976 by Mr. T. Murota, with descriptions of new taxa (Hymenoptera). Special Publications of the Japan Hymenopterists Association, 41, 39 - 50." title="On some Taiwanese Mutillidae, collected in 1976 by Mr. T. Murota, with descriptions of new taxa (Hymenoptera)" type="journal article" year="1993">Tsuneki, 1993</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="2C54470FFFC5CA6BFF2CFA9E7003FAB7" blockId="1.[151,1436,1302,1386]" pageId="1" pageNumber="102">
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<taxonomicName id="EBEB3C8CFFC5CA6BFF2CFA9E73EEFAD0" authority="Tsuneki, 1993: 42" authorityName="Tsuneki" authorityPageNumber="42" authorityYear="1993" box="[151,429,1302,1324]" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="1E9F9B1DFFC5CA6BFF2CFA9E72B0FAD0" box="[151,243,1302,1324]" italics="true" pageId="1" pageNumber="102">Zavatilla</emphasis>
<bibRefCitation id="487A3AFEFFC5CA6BFF43FA9E73C9FAD0" author="Tsuneki, K." box="[248,394,1302,1324]" journalOrPublisher="Special Publications of the Japan Hymenopterists Association" pageId="1" pageNumber="102" pagination="39 50" part="41" refId="ref10470" refString="Tsuneki, K. (1993) On some Taiwanese Mutillidae, collected in 1976 by Mr. T. Murota, with descriptions of new taxa (Hymenoptera). Special Publications of the Japan Hymenopterists Association, 41, 39 - 50." title="On some Taiwanese Mutillidae, collected in 1976 by Mr. T. Murota, with descriptions of new taxa (Hymenoptera)" type="journal article" year="1993">Tsuneki, 1993</bibRefCitation>
: 42
</taxonomicName>
(as subgenus of
<taxonomicName id="EBEB3C8CFFC5CA6BFDECFA9F71CFFAD0" authority="Thomson, 1870" authorityName="THOMSON" authorityYear="1870" box="[599,908,1302,1324]" class="Insecta" family="Mutillidae" genus="Smicromyrme" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="genus">
<emphasis id="1E9F9B1DFFC5CA6BFDECFA9F70A0FAD0" box="[599,739,1303,1324]" italics="true" pageId="1" pageNumber="102">Smicromyrme</emphasis>
<bibRefCitation id="487A3AFEFFC5CA6BFD52FA9E71CFFAD0" author="Thomson, C. G." box="[745,908,1302,1324]" journalOrPublisher="Opuscula Entomologica, 1" pageId="1" pageNumber="102" pagination="202 251" part="2" refId="ref10417" refString="Thomson, C. G. (1870) Ofversigt af Sveriges rofsteklar. Opuscula Entomologica, 1 (2), 202 - 251." title="Öfversigt af Sveriges rofsteklar" type="journal article" year="1870">Thomson, 1870</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="AD4A611EFFC5CA6BFC26FA9E7665FAD0" author="" box="[925,1062,1302,1324]" page="2" pageId="1" pageNumber="102">
<bibRefCitation id="487A3AFEFFC5CA6BFC26FA9E764EFAD0" author="Lelej" box="[925,1037,1302,1324]" pageId="1" pageNumber="102" refId="ref9603" refString="Lelej, A. S. (1995) A review of Palaearctic and Oriental species of genus Nemka Lelej with description of Oriental genus Mickelomyrme gen. n. (Hymenoptera, Mutillidae). Far Eastern Entomologist, 6, 1 - 20." type="journal article" year="1995">Lelej 1995</bibRefCitation>
: 2
</treatmentCitation>
; 1996: 8; 2005: 77;
<treatmentCitation id="AD4A611EFFC5CA6BFB4AFA9E7360FAB7" author="" page="66" pageId="1" pageNumber="102">
<bibRefCitation id="487A3AFEFFC5CA6BFB4AFA9E72BEFAB7" author="Lelej" pageId="1" pageNumber="102" refId="ref9815" refString="Lelej, A. S. &amp; Brothers, D. J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889, 1 - 79." type="journal article" year="2008">Lelej &amp; Brothers 2008</bibRefCitation>
: 66
</treatmentCitation>
;
<treatmentCitation id="AD4A611EFFC5CA6BFE96FABD7078FAB7" author="" box="[301,571,1333,1355]" page="97" pageId="1" pageNumber="102">
<bibRefCitation id="487A3AFEFFC5CA6BFE96FABD7056FAB7" author="Brothers" box="[301,533,1333,1355]" pageId="1" pageNumber="102" refId="ref9250" refString="Brothers, D. J. &amp; Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1 - 97." type="journal article" year="2017">Brothers &amp; Lelej 2017</bibRefCitation>
: 97
</treatmentCitation>
.
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</paragraph>
</subSubSection>
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<paragraph id="2C54470FFFC5CA6BFF2CFADD71CCFA96" blockId="1.[151,1436,1302,1386]" box="[151,911,1364,1386]" pageId="1" pageNumber="102">
Type species.
<taxonomicName id="EBEB3C8CFFC5CA6BFE9EFADC7036FA96" authority="Zavattari, 1913" authorityName="ZAVATTARI" authorityYear="1913" box="[293,629,1364,1386]" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="species" species="gutrunae">
<emphasis id="1E9F9B1DFFC5CA6BFE9EFADC7392FA96" box="[293,465,1364,1386]" italics="true" pageId="1" pageNumber="102">Mutilla gutrunae</emphasis>
<bibRefCitation id="487A3AFEFFC5CA6BFE63FADC7036FA96" author="Zavattari, E." box="[472,629,1364,1386]" journalOrPublisher="Archiv für Naturgeschichte, 79 A" pageId="1" pageNumber="102" pagination="19 42" part="3" refId="ref10751" refString="Zavattari, E. (1913) H. Sauter's Formosa-Ausbeute. Mutillidae (Hym.). Archiv fur Naturgeschichte, 79 A (3), 19 - 42." title="H. Sauters Formosa-Ausbeute. Mutillidae (Hym.)" type="journal article" year="1913">Zavattari, 1913</bibRefCitation>
</taxonomicName>
, ♂, by original designation.
</paragraph>
<paragraph id="2C54470FFFC5CA6BFF2CFA1E732CFA52" blockId="1.[151,1438,1430,2031]" box="[151,367,1430,1455]" pageId="1" pageNumber="102">
<emphasis id="1E9F9B1DFFC5CA6BFF2CFA1E72B5FA53" bold="true" box="[151,246,1430,1455]" pageId="1" pageNumber="102">Gender.</emphasis>
Feminine.
</paragraph>
</subSubSection>
<subSubSection id="64F11484FFC5CA68FF7CFA327381FEBC" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102" type="diagnosis">
<paragraph id="2C54470FFFC5CA68FF7CFA327724FF04" blockId="1.[151,1438,1430,2031]" lastBlockId="2.[151,1437,151,896]" lastPageId="2" lastPageNumber="103" pageId="1" pageNumber="102">
<emphasis id="1E9F9B1DFFC5CA6BFF7CFA327302FA2F" bold="true" box="[199,321,1466,1491]" pageId="1" pageNumber="102">Diagnosis.</emphasis>
MALE.
<materialsCitation id="9C834D52FFC5CA6BFE14FA33717AF9E7" ID-GBIF-Occurrence="1841174340" collectionCode="SMOOTH, SHINY, WITH, PART, APICALLY" location="Of" municipality="UPPeR HeAd" pageId="1" pageNumber="102" specimenCount="1">
Punctures
<location id="293411D4FFC5CA6BFD92FA327003FA2F" LSID="urn:lsid:plazi:treatment:A442F619FFC5CA68FF2CFB58739DFC7C:293411D4FFC5CA6BFD92FA327003FA2F" box="[553,576,1466,1491]" municipality="UPPeR HeAd" name="Of" pageId="1" pageNumber="102">of</location>
<collectingMunicipality id="CC30DD75FFC5CA6BFDF6FA33708EFA2F" box="[589,717,1466,1491]" pageId="1" pageNumber="102">upper head</collectingMunicipality>
and
<taxonomicName id="EBEB3C8CFFC5CA6BFCB6FA3371C0FA2F" box="[781,899,1467,1491]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="114" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
coarse, confluent, interspaces linear. Clypeus conspicuously concave, medially
<collectionCode id="4AFADFCAFFC5CA6BFDAFFA577024FA0B" box="[532,615,1503,1527]" pageId="1" pageNumber="102">smooth</collectionCode>
and
<collectionCode id="4AFADFCAFFC5CA6BFD24FA577155FA0A" box="[671,790,1503,1527]" pageId="1" pageNumber="102">shiny with</collectionCode>
transverse anterior sculptured
<collectionCode id="4AFADFCAFFC5CA6BFBC8FA5776DCFA0B" box="[1139,1183,1503,1527]" pageId="1" pageNumber="102">part</collectionCode>
and slight but distinct anteromedian emargination. Mandible bidentate
<collectionCode id="4AFADFCAFFC5CA6BFD65F98B717AF9E7" box="[734,825,1539,1563]" pageId="1" pageNumber="102">apically</collectionCode>
</materialsCitation>
, with moderate subbasal tooth beneath (
<figureCitation id="B4D05B8AFFC5CA6BFA8AF98B77D3F9E7" box="[1329,1424,1538,1563]" captionStart="FIGURES 2530" captionStartId="9.[151,264,1632,1655]" captionTargetBox="[151,1436,193,1611]" captionTargetId="figure@9.[151,1436,193,1611]" captionTargetPageId="9" captionText="FIGURES 2530. Zavatilla flavotegulata, ♂, Nanling. 25. Antenna; 26. Mandible; 27. Head, anterior view; 28, 29. Head and mesosoma (28, lateral view, 29, dorsal view); 30. Propodeum, dorsal view. (Photos H.-t. Zhou)." httpUri="https://zenodo.org/record/1242743/files/figure.png" pageId="1" pageNumber="102">Fig. 26</figureCitation>
). Flagellomere 1 length 1.01.2 × width, 0.80.95 × flagellomere 2 length. Scape with one strong ventral carina. Mesoscutum with inconspicuous lateral ridges. Mesoscutellum evenly gibbous, with median carina on anterior inclination. Tegula long, protruding over scuto-scutellar suture. Prospurs simple. Metasomal tergum 2 with long lateral felt line, 3.84.0 × length of that on metasomal sternum 2. Metasomal tergum 7 with median longitudinal smooth shiny line that widens and is gibbous posterad. Sternum 8 (hypopygium) transversely depressed at base, without strong lateral
<taxonomicName id="EBEB3C8CFFC5CA6BFE35F95373A0F90F" authorityName="" authorityYear="1992" box="[398,483,1755,1779]" class="Insecta" family="Eccoptarthridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="102" phylum="Arthropoda" rank="subFamily" subFamily="Carinae">carinae</taxonomicName>
. Penis valves symmetrical. Volsella with cuspis equal to or slightly longer than penis valves (
<figureCitation id="B4D05B8AFFC5CA6BFF55F97773EAF8EB" box="[238,425,1790,1815]" captionStart="FIGURES 4654" captionStartId="14.[151,263,1936,1959]" captionTargetBox="[224,1309,260,1902]" captionTargetId="figure@14.[168,1373,253,1905]" captionTargetPageId="14" captionText="FIGURES 4654. Zavatilla, ♂. 4648. Z. gutrunae; 4951. Z. flavotegulata, Nanling; 5254. Z. xuzaifui sp. nov., holotype. 46, 49, 52. Genitalia, dorsal view; 47. Scape; 50, 53. Genitalia, lateral view; 48, 51, 54. Genitalia, ventral view. (Photos H.-t. Zhou)." httpUri="https://zenodo.org/record/1242753/files/figure.png" pageId="1" pageNumber="102">Figs. 48, 51, 54</figureCitation>
), with digitus and distinct paracuspis. Parapenis much shorter than penis valve, apically thin, acuminated.
<materialsCitation id="9C834D52FFC5CA6BFECBF8AB7171F8A3" ID-GBIF-Occurrence="1841174343" collectionCode="TOOTH, SMALL, INCONSPICUOUS" location="Of" municipality="UPPeR HeAd" pageId="1" pageNumber="102" specimenCount="1">
FEMALE. Punctures
<location id="293411D4FFC5CA6BFDCEF8AA70CFF8C7" LSID="urn:lsid:plazi:treatment:A442F619FFC5CA68FF2CFB58739DFC7C:293411D4FFC5CA6BFDCEF8AA70CFF8C7" box="[629,652,1826,1851]" municipality="UPPeR HeAd" name="Of" pageId="1" pageNumber="102">of</location>
<collectingMunicipality id="CC30DD75FFC5CA6BFD27F8AB7163F8C7" box="[668,800,1826,1851]" pageId="1" pageNumber="102">upper head</collectingMunicipality>
and
<taxonomicName id="EBEB3C8CFFC5CA6BFCDCF8AB719EF8C7" box="[871,989,1827,1851]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="114" phylum="Arthropoda" rank="genus">mesosoma</taxonomicName>
coarse, confluent, interspaces linear. Mandible bidentate, upper
<collectionCode id="4AFADFCAFFC5CA6BFE74F8CF7013F8A3" box="[463,592,1863,1887]" pageId="1" pageNumber="102">tooth small</collectionCode>
and
<collectionCode id="4AFADFCAFFC5CA6BFD36F8CF7171F8A3" box="[653,818,1863,1887]" pageId="1" pageNumber="102">inconspicuous</collectionCode>
</materialsCitation>
. Dorsal transverse curved
<taxonomicName id="EBEB3C8CFFC5CA6BFBDDF8CF7688F8A3" authority="Of" authorityName="Of" box="[1126,1227,1862,1887]" class="Aves" family="Anatidae" genus="Carina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anseriformes" pageId="1" pageNumber="102" phylum="Chordata" rank="genus">carina of</taxonomicName>
clypeus with two tubercles that are larger or smaller than basomedian one. Flagellomere 1 length 1.331.45 × apical width, 1.541.6 × flagellomere 2 length, which is 0.810.83 × its width. Genal carina developed, ending anterad on hypostomal carina with tubercle.
<materialsCitation id="9C834D52FFC5CA6BFE2EF83B76E5F813" ID-GBIF-Occurrence="1841174327" collectionCode="WITH, CARINA" location="Of" pageId="1" pageNumber="102" specimenCount="1">
Scutellar scale wide, 0.640.72 × distance between scutellar scale and inner border
<location id="293411D4FFC5CA6BFA3EF83A77DFF837" LSID="urn:lsid:plazi:treatment:A442F619FFC5CA68FF2CFB58739DFC7C:293411D4FFC5CA6BFA3EF83A77DFF837" box="[1413,1436,1970,1995]" name="Of" pageId="1" pageNumber="102">of</location>
posterior spiracle. Metasomal sternum 1
<collectionCode id="4AFADFCAFFC5CA6BFDD2F85F70D8F812" box="[617,667,2007,2030]" pageId="1" pageNumber="102">with</collectionCode>
more or less emarginated longitudinal
<collectionCode id="4AFADFCAFFC5CA6BFBE6F85F76E5F813" box="[1117,1190,2007,2031]" pageId="1" pageNumber="102">carina</collectionCode>
</materialsCitation>
. Metasomal tergum 2 posteriorly with fringe of dense golden (whitish) or black decumbent setae. Metasomal tergum 3 with continuous band of dense golden or whitish decumbent setae. Pygidial plate wide, 0.60.7 × total width of metasomal tergum 6, rounded laterally, inconspicuously carinated laterally, finely granulate or longitudinally striate in basal half.
</paragraph>
<paragraph id="2C54470FFFC6CA68FF7CFE8C7381FEBC" blockId="2.[151,1437,151,896]" pageId="2" pageNumber="103">
<emphasis id="1E9F9B1DFFC6CA68FF7CFE8C73C0FEE1" bold="true" box="[199,387,260,285]" pageId="2" pageNumber="103">Sex association.</emphasis>
The female of the
<typeStatus id="F350F9ADFFC6CA68FDE5FE8D70CDFEE0" box="[606,654,261,284]" pageId="2" pageNumber="103">type</typeStatus>
species was originally described
<collectingCountry id="54FC079FFFC6CA68FBB5FE8C7666FEE0" box="[1038,1061,260,284]" name="American Samoa" pageId="2" pageNumber="103">as</collectingCountry>
<taxonomicName id="EBEB3C8CFFC6CA68FB96FE8C773BFEE0" authority="ZAVATTARI, 1913" authorityName="Zavattari" authorityYear="1913" box="[1069,1400,260,285]" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="species" species="logei">
<emphasis id="1E9F9B1DFFC6CA68FB96FE8C7683FEE1" box="[1069,1216,260,285]" italics="true" pageId="2" pageNumber="103">Mutilla logei</emphasis>
<bibRefCitation id="487A3AFEFFC6CA68FB72FE8C773BFEE0" author="Zavattari, E." box="[1225,1400,260,284]" journalOrPublisher="Archiv für Naturgeschichte, 79 A" pageId="2" pageNumber="103" pagination="19 42" part="3" refId="ref10751" refString="Zavattari, E. (1913) H. Sauter's Formosa-Ausbeute. Mutillidae (Hym.). Archiv fur Naturgeschichte, 79 A (3), 19 - 42." title="H. Sauters Formosa-Ausbeute. Mutillidae (Hym.)" type="journal article" year="1913">Zavattari, 1913</bibRefCitation>
</taxonomicName>
,
<collectingCountry id="54FC079FFFC6CA68FA3EFE8C77DFFEE0" box="[1413,1436,260,284]" name="American Samoa" pageId="2" pageNumber="103">as</collectingCountry>
suggested by
<bibRefCitation id="487A3AFEFFC6CA68FE94FEAF73FEFEBC" author="Lelej" box="[303,445,295,320]" pageId="2" pageNumber="103" refId="ref9725" refString="Lelej, A. S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp." type="book" year="2005">Lelej (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="64F11484FFC6CA68FF7CFEC47733FE2C" pageId="2" pageNumber="103" type="description">
<paragraph id="2C54470FFFC6CA68FF7CFEC47733FE2C" blockId="2.[151,1437,151,896]" pageId="2" pageNumber="103">
<emphasis id="1E9F9B1DFFC6CA68FF7CFEC473DBFE99" bold="true" box="[199,408,332,357]" pageId="2" pageNumber="103">Species included.</emphasis>
The genus includes four species (including two new ones described below):
<taxonomicName id="EBEB3C8CFFC6CA68FA8CFEC47387FE74" authority="(ZAVATTARI, 1913)" baseAuthorityName="ZAVATTARI" baseAuthorityYear="1913" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="105" phylum="Arthropoda" rank="species" species="gutrunae">
<emphasis id="1E9F9B1DFFC6CA68FA8CFEC472BFFE74" italics="true" pageId="2" pageNumber="103">Zavatilla gutrunae</emphasis>
(
<bibRefCitation id="487A3AFEFFC6CA68FEB2FEF873FFFE74" author="Zavattari, E." box="[265,444,368,392]" journalOrPublisher="Archiv für Naturgeschichte, 79 A" pageId="2" pageNumber="103" pagination="19 42" part="3" refId="ref10751" refString="Zavattari, E. (1913) H. Sauter's Formosa-Ausbeute. Mutillidae (Hym.). Archiv fur Naturgeschichte, 79 A (3), 19 - 42." title="H. Sauters Formosa-Ausbeute. Mutillidae (Hym.)" type="journal article" year="1913">Zavattari, 1913</bibRefCitation>
)
</taxonomicName>
, ♂, ♀ (China: Taiwan),
<taxonomicName id="EBEB3C8CFFC6CA68FD63FEF9766AFE74" authority="(CHeN, 1957)" authorityName="Zhou &amp; Lelej &amp; Williams &amp; Liu" authorityYear="2018" baseAuthorityName="CHeN" baseAuthorityYear="1957" box="[728,1065,367,392]" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="flavotegulata">
<emphasis id="1E9F9B1DFFC6CA68FD63FEF971C9FE74" box="[728,906,367,392]" italics="true" pageId="2" pageNumber="103">Z. flavotegulata</emphasis>
(
<bibRefCitation id="487A3AFEFFC6CA68FC23FEF87663FE74" author="Chen" box="[920,1056,367,392]" pageId="2" pageNumber="103" refId="ref9303" refString="Chen, C. W. (1957) A revision of the velvet ants or Mutillidae of China (Hymenoptera). Quarterly Journal of the Taiwan Museum, 10 (3 - 4), 135 - 224, 6 pls." type="journal article" year="1957">Chen, 1957</bibRefCitation>
)
</taxonomicName>
, ♂, ♀ (China: Zhejiang, Jiangxi, Fujian, Hunan, Guangdong, Hainan);
<taxonomicName id="EBEB3C8CFFC6CA68FDFBFE1D70F4FE50" authority="Zhou &amp; Lelej &amp; Williams &amp; Liu, 2018" authorityName="Zhou &amp; Lelej &amp; Williams &amp; Liu" authorityYear="2018" box="[576,695,404,428]" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="xuzaifui" status="sp. nov.">
<emphasis id="1E9F9B1DFFC6CA68FDFBFE1D70F4FE50" box="[576,695,404,428]" italics="true" pageId="2" pageNumber="103">Z. xuzaifui</emphasis>
</taxonomicName>
<emphasis id="1E9F9B1DFFC6CA68FD06FE1C7154FE51" bold="true" box="[701,791,404,429]" pageId="2" pageNumber="103">
<taxonomicNameLabel id="05AC2666FFC6CA68FD06FE1C7154FE51" box="[701,791,404,429]" pageId="2" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂, ♀ (China: Guangdong, Yunnan, Hainan; Vietnam);
<taxonomicName id="EBEB3C8CFFC6CA68FA3EFE1D734EFE2C" authority="Zhou &amp; Lelej &amp; Williams &amp; Liu, 2018" authorityName="Zhou &amp; Lelej &amp; Williams &amp; Liu" authorityYear="2018" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nepalensis" status="sp. nov.">
<emphasis id="1E9F9B1DFFC6CA68FA3EFE1D734EFE2C" italics="true" pageId="2" pageNumber="103">Z. nepalensis</emphasis>
</taxonomicName>
<emphasis id="1E9F9B1DFFC6CA68FEAFFE3F732DFE2C" bold="true" box="[276,366,439,464]" pageId="2" pageNumber="103">
<taxonomicNameLabel id="05AC2666FFC6CA68FEAFFE3F732DFE2C" box="[276,366,439,464]" pageId="2" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♀ (Nepal). Three species are known from both sexes and
<taxonomicName id="EBEB3C8CFFC6CA68FBBAFE3176D7FE2C" authorityName="Zhou &amp; Lelej &amp; Williams &amp; Liu" authorityYear="2018" box="[1025,1172,439,464]" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="1E9F9B1DFFC6CA68FBBAFE3176D7FE2C" box="[1025,1172,439,464]" italics="true" pageId="2" pageNumber="103">Z. nepalensis</emphasis>
</taxonomicName>
from females only.
</paragraph>
</subSubSection>
<subSubSection id="64F11484FFC6CA68FF7CFE5472A1FDE4" pageId="2" pageNumber="103" type="distribution">
<paragraph id="2C54470FFFC6CA68FF7CFE5472A1FDE4" blockId="2.[151,1437,151,896]" pageId="2" pageNumber="103">
<emphasis id="1E9F9B1DFFC6CA68FF7CFE547323FE09" bold="true" box="[199,352,476,501]" pageId="2" pageNumber="103">Distribution.</emphasis>
<collectingCountry id="54FC079FFFC6CA68FED6FE5473F3FE08" box="[365,432,476,500]" name="China" pageId="2" pageNumber="103">China</collectingCountry>
(
<collectingRegion id="EE2F89EDFFC6CA68FE7FFE54706CFE09" box="[452,559,476,501]" country="China" name="Zhejiang" pageId="2" pageNumber="103">Zhejiang</collectingRegion>
,
<collectingRegion id="EE2F89EDFFC6CA68FD84FE5470D7FE08" box="[575,660,476,500]" country="China" name="Jiangxi" pageId="2" pageNumber="103">Jiangxi</collectingRegion>
,
<collectingRegion id="EE2F89EDFFC6CA68FD1EFE5470B2FE09" box="[677,753,476,501]" country="China" name="Fujian" pageId="2" pageNumber="103">Fujian</collectingRegion>
,
<collectingCountry id="54FC079FFFC6CA68FCBAFE547114FE08" box="[769,855,476,500]" name="Taiwan" pageId="2" pageNumber="103">Taiwan</collectingCountry>
,
<collectingRegion id="EE2F89EDFFC6CA68FCDCFE5471F5FE08" box="[871,950,476,500]" country="China" name="Hunan" pageId="2" pageNumber="103">Hunan</collectingRegion>
,
<collectingRegion id="EE2F89EDFFC6CA68FC7CFE54760DFE09" box="[967,1102,476,501]" country="China" name="Guangdong" pageId="2" pageNumber="103">Guangdong</collectingRegion>
,
<collectingRegion id="EE2F89EDFFC6CA68FBE5FE5476FBFE08" box="[1118,1208,476,500]" country="China" name="Yunnan" pageId="2" pageNumber="103">Yunnan</collectingRegion>
,
<collectingRegion id="EE2F89EDFFC6CA68FB73FE547763FE08" box="[1224,1312,476,500]" country="China" name="Hainan" pageId="2" pageNumber="103">Hainan</collectingRegion>
),
<collectingCountry id="54FC079FFFC6CA68FA8DFE5477DAFE08" box="[1334,1433,476,500]" name="Vietnam" pageId="2" pageNumber="103">Vietnam</collectingCountry>
,
<collectingCountry id="54FC079FFFC6CA68FF2CFD88729DFDE4" box="[151,222,512,536]" name="Nepal" pageId="2" pageNumber="103">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="64F11484FFC6CA68FF7CFDAC739DFC7C" pageId="2" pageNumber="103" type="discussion">
<paragraph id="2C54470FFFC6CA68FF7CFDAC739DFC7C" blockId="2.[151,1437,151,896]" pageId="2" pageNumber="103">
<emphasis id="1E9F9B1DFFC6CA68FF7CFDAC7378FDC1" bold="true" box="[199,315,548,573]" pageId="2" pageNumber="103">Remarks.</emphasis>
The male of the genus
<taxonomicName id="EBEB3C8CFFC6CA68FDE6FDAC7081FDC1" authorityName="Tsuneki" authorityYear="1993" box="[605,706,548,573]" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="1E9F9B1DFFC6CA68FDE6FDAC7081FDC1" box="[605,706,548,573]" italics="true" pageId="2" pageNumber="103">Zavatilla</emphasis>
</taxonomicName>
differs from other Oriental genera of tribe Trogaspidiini with symmetric penis valves (former tribe Petersenidiini) by having the following set of characters: scape with one ventral carina, volsella with paracuspis, flagellomeres short (flagellomere 1 length 1.01.2 × width, 0.80.95 × flagellomere 2 length), and metasomal sternum 2 with short lateral felt line. The female of the genus
<taxonomicName id="EBEB3C8CFFC6CA68FA8CFD0777DFFD54" authorityName="Tsuneki" authorityYear="1993" box="[1335,1436,655,680]" class="Insecta" family="Mutillidae" genus="Zavatilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="1E9F9B1DFFC6CA68FA8CFD0777DFFD54" box="[1335,1436,655,680]" italics="true" pageId="2" pageNumber="103">Zavatilla</emphasis>
</taxonomicName>
differs from other Oriental genera of tribe Trogaspidiini with propodeum not wider than pronotum (former tribe Petersenidiini) by having the following set of characters: transverse curved
<taxonomicName id="EBEB3C8CFFC6CA68FC4CFD507618FD0C" authority="Of" authorityName="Of" box="[1015,1115,727,752]" class="Aves" family="Anatidae" genus="Carina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anseriformes" pageId="2" pageNumber="103" phylum="Chordata" rank="genus">carina of</taxonomicName>
clypeus with two sublateral tubercles, which can be smaller or larger than basal median clypeal tubercle; scutellar scale wide (0.640.72 × distance between scutellar scale and inner border of posterior spiracle); and pygidial plate wide (0.60.7 × total width of metasomal tergum 6), carinated laterally, finely granulated or longitudinally striated in basal part and mostly smooth in apical part.
</paragraph>
</subSubSection>
</treatment>
</document>