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<document id="04D407B17C0D9A6A5FF73924EFBE3863" ID="10.11646/zootaxa.4168.2.12" ID-DOI="10.11646/zootaxa.4168.2.12" ID-GBIF-Dataset="ec03921e-011a-4dfa-8384-7a6e0ab50ade" ID-ISSN="1175-5326" ID-Zenodo-Dep="257193" ID-ZooBank="D6AE160B-7442-47D5-B57F-584CC4BBB1F2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473925729460" checkinUser="plazi" docAuthor="Bartlett, Justin S. &amp; Gerstmeier, Roland" docDate="2016" docId="997887F56A56FFC9FF0040D9D1116DAD" docLanguage="en" docName="zootaxa.4168.2.12.pdf" docOrigin="Zootaxa 4168 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Menieroclerus Bartlett &amp; Gerstmeier, gen. nov." docType="treatment" docVersion="7" lastPageNumber="398" masterDocId="6541FF8D6A57FFCAFF97416DD0066B71" masterDocTitle="Menieroclerus gen. nov.: A new genus for Thanasimodes nigropiceus (Kuwert, 1893) (Coleoptera: Cleridae: Clerinae)" masterLastPageNumber="400" masterPageNumber="395" pageNumber="396" updateTime="1698385950029" updateUser="plazi">
<mods:mods id="923A2717B79DA45E1D53C12A192D2B3F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="69C43AAA0A6A2EB255BFD214A138DA1D">Menieroclerus gen. nov.: A new genus for Thanasimodes nigropiceus (Kuwert, 1893) (Coleoptera: Cleridae: Clerinae)</mods:title>
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<mods:namePart id="CD7AC3B861E47D59D865869D01DE40B4">Bartlett, Justin S.</mods:namePart>
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<mods:namePart id="61A08B51AB68BC876BD8BC6C3ED09102">Gerstmeier, Roland</mods:namePart>
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<mods:title id="CDFCDEFA313EAE8DF69011F0DB358E95">Zootaxa</mods:title>
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<mods:part id="DC63BDB7B1BF80DAB630D074CB545CE7">
<mods:date id="87391F4C3442FD77C52113BEB18C0AC5">2016</mods:date>
<mods:detail id="9D622F295A67A29EB5F80F7717E3E436" type="volume">
<mods:number id="C805889D531B9524E50DDC17F57542D3">4168</mods:number>
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<mods:detail id="6D08667A53D7368F9E4B138ED8253449" type="issue">
<mods:number id="753F1F88DB639B9B1AA3B8197948117F">2</mods:number>
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<mods:identifier id="F4778A5B26CF36BD51D8F3632DADCA60" type="DOI">10.11646/zootaxa.4168.2.12</mods:identifier>
<mods:identifier id="2AD411AF35945BE9C2758768D5E1DEE6" type="GBIF-Dataset">ec03921e-011a-4dfa-8384-7a6e0ab50ade</mods:identifier>
<mods:identifier id="93BE6306D6CCB0B508D5020AEBE082C8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5A392743C97D78DFD17528050C1A1CF6" type="Zenodo-Dep">257193</mods:identifier>
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<treatment id="997887F56A56FFC9FF0040D9D1116DAD" ID-DOI="http://doi.org/10.5281/zenodo.6079839" ID-GBIF-Taxon="124551846" ID-Zenodo-Dep="6079839" LSID="urn:lsid:plazi:treatment:997887F56A56FFC9FF0040D9D1116DAD" httpUri="http://treatment.plazi.org/id/997887F56A56FFC9FF0040D9D1116DAD" lastPageId="3" lastPageNumber="398" pageId="1" pageNumber="396">
<subSubSection id="59CB65686A56FFCBFF0040D9D2D86ABE" box="[151,734,436,463]" pageId="1" pageNumber="396" type="nomenclature">
<paragraph id="116E36E36A56FFCBFF0040D9D2D86ABE" blockId="1.[151,734,436,463]" box="[151,734,436,463]" pageId="1" pageNumber="396">
<heading id="4A26818F6A56FFCBFF0040D9D2D86ABE" bold="true" box="[151,734,436,463]" fontSize="11" level="1" pageId="1" pageNumber="396" reason="1">
<emphasis id="23A5EAF16A56FFCBFF0040D9D2D86ABE" bold="true" box="[151,734,436,463]" pageId="1" pageNumber="396">
<taxonomicName id="D6D14D606A56FFCBFF0040D9D2656ABE" authority="Bartlett &amp; Gerstmeier" authorityName="Bartlett &amp; Gerstmeier" box="[151,611,436,463]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="23A5EAF16A56FFCBFF0040D9D1416ABF" bold="true" box="[151,327,436,462]" italics="true" pageId="1" pageNumber="396">Menieroclerus</emphasis>
Bartlett &amp; Gerstmeier
</taxonomicName>
,
<taxonomicNameLabel id="3896578A6A56FFCBFDE740D8D2D86ABE" box="[624,734,437,463]" pageId="1" pageNumber="396" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="59CB65686A56FFCBFF004091D3756963" box="[151,883,508,530]" pageId="1" pageNumber="396" type="materials_examined">
<paragraph id="116E36E36A56FFCBFF004091D3756963" blockId="1.[151,883,508,530]" box="[151,883,508,530]" pageId="1" pageNumber="396">
<emphasis id="23A5EAF16A56FFCBFF004091D1246963" bold="true" box="[151,290,508,530]" pageId="1" pageNumber="396">Type species.</emphasis>
<taxonomicName id="D6D14D606A56FFCBFEBE4091D2746963" authority="Kuwert, 1893" authorityName="Kuwert" authorityYear="1893" box="[297,626,508,530]" class="Insecta" family="Cleridae" genus="Opilo" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="nigropiceus">
<emphasis id="23A5EAF16A56FFCBFEBE4091D1E66963" box="[297,480,508,530]" italics="true" pageId="1" pageNumber="396">Opilo nigropiceus</emphasis>
<bibRefCitation id="75404B126A56FFCBFE714090D2746963" author="Kuwert" box="[486,626,508,530]" pageId="1" pageNumber="396" refString="Kuwert, A. (1893) Neue und alte Africanische Cleriden. Annales de la Societe entomologique de Belgique, 37, 467 - 479." type="journal article" year="1893">Kuwert, 1893</bibRefCitation>
</taxonomicName>
(by present designation).
</paragraph>
</subSubSection>
<subSubSection id="59CB65686A56FFCBFF004353D1776892" pageId="1" pageNumber="396" type="diagnosis">
<paragraph id="116E36E36A56FFCBFF004353D284687A" blockId="1.[151,1437,574,1931]" pageId="1" pageNumber="396">
<emphasis id="23A5EAF16A56FFCBFF004353D10C6926" bold="true" box="[151,266,574,599]" pageId="1" pageNumber="396">Diagnosis</emphasis>
. The strongly bilobate condition of the labrum (
<figureCitation id="89EA2A666A56FFCBFCA14353D3816926" box="[822,903,574,599]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 11</figureCitation>
) is possibly unique among
<taxonomicName id="D6D14D606A56FFCBFB514353D52E6926" box="[1222,1320,574,599]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="family">Cleridae</taxonomicName>
; however, even if it is not unique then the following characters in combination are diagnostic for
<taxonomicName id="D6D14D606A56FFCBFB02430FD531690A" box="[1173,1335,610,635]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="23A5EAF16A56FFCBFB02430FD531690A" box="[1173,1335,610,635]" italics="true" pageId="1" pageNumber="396">Menieroclerus</emphasis>
</taxonomicName>
<emphasis id="23A5EAF16A56FFCBFAD5430FD593690A" bold="true" box="[1346,1429,610,635]" pageId="1" pageNumber="396">
<taxonomicNameLabel id="3896578A6A56FFCBFAD5430FD593690A" box="[1346,1429,610,635]" pageId="1" pageNumber="396" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
: terminal maxillary palpomeres securiform (
<figureCitation id="89EA2A666A56FFCBFD0643EBD2EF69EE" box="[657,745,646,671]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 13</figureCitation>
); gular sutures very weakly divergent (appearing parallel); elytron with ten rows of deep, laterally nodulate punctures (
<figureCitation id="89EA2A666A56FFCBFCC643C7D3A169B2" box="[849,935,682,707]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 12</figureCitation>
b); tibiae longitudinally carinate; tibial spur formula 2–2–2; tarsal pad formula 4–4–4 (basitarsal pads substantially smaller than those of tarsomeres 2–4), tarsal pads not divided into paired lobes (
<figureCitation id="89EA2A666A56FFCBFDB6439FD272687A" box="[545,628,754,779]" captionStart="FIGURES 14 – 16" captionStartId="3.[151,264,1165,1187]" captionTargetBox="[153,1436,193,1137]" captionTargetId="figure@3.[151,1436,193,1144]" captionTargetPageId="3" captionText="FIGURES 14 – 16. Menieroclerus nigropiceus: (14) female specimen (scale = 1 mm); (15) female mesotarsus showing nonbifid tarsal pads (scale = 0.5 mm); (16) distribution." httpUri="https://zenodo.org/record/257195/files/figure.png" pageId="1" pageNumber="396">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="116E36E36A56FFCBFF50427BD1776892" blockId="1.[151,1437,574,1931]" pageId="1" pageNumber="396">
<taxonomicName id="D6D14D606A56FFCBFF50427BD16C685E" box="[199,362,790,815]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="23A5EAF16A56FFCBFF50427BD16C685E" box="[199,362,790,815]" italics="true" pageId="1" pageNumber="396">Menieroclerus</emphasis>
</taxonomicName>
<emphasis id="23A5EAF16A56FFCBFEE5427BD1C2685E" bold="true" box="[370,452,790,815]" pageId="1" pageNumber="396">
<taxonomicNameLabel id="3896578A6A56FFCBFEE5427BD1C2685E" box="[370,452,790,815]" pageId="1" pageNumber="396" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="D6D14D606A56FFCBFDCA427BD2F9685E" box="[605,767,790,815]" class="Insecta" family="Cleridae" genus="Thanasimodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="23A5EAF16A56FFCBFDCA427BD2F9685E" box="[605,767,790,815]" italics="true" pageId="1" pageNumber="396">Thanasimodes</emphasis>
</taxonomicName>
(= ‘T’) in the following character states: Labrum bilobate (emarginate in ‘T’); gular sutures parallel (convergent in ‘T’); procoxal cavities posteriorly broadly open (narrowly open to completely closed in ‘T’); tarsal pads not incised (incised in ‘T’); two protibial spurs (none in ‘T’); pretarsal claws appendiculate (simple in ‘T’); elytral punctation nodulate (without nodules in ‘T’); base of CuA1+2 of hindwing detached from CuA (attached to CuA in ‘T’); tegminal phallobasic struts not fused to apodeme (fused to apodeme in ‘T’).
</paragraph>
</subSubSection>
<subSubSection id="59CB65686A56FFC9FF504283D1116DAD" lastPageId="3" lastPageNumber="398" pageId="1" pageNumber="396" type="description">
<paragraph id="116E36E36A56FFCBFF504283D14B6D1A" blockId="1.[151,1437,574,1931]" pageId="1" pageNumber="396">
<emphasis id="23A5EAF16A56FFCBFF504283D1576F76" bold="true" box="[199,337,1006,1031]" pageId="1" pageNumber="396">Description</emphasis>
.
<emphasis id="23A5EAF16A56FFCBFEF54283D1A06F76" box="[354,422,1006,1031]" italics="true" pageId="1" pageNumber="396">Head.</emphasis>
Eyes large, coarsely facetted, frontal rim strongly margined, emarginate above antennal socket; antennae with 11 antennomeres, flagellomeres increasingly dilated towards
<collectingRegion id="D315F8016A56FFCBFBC0457ED46B6F5A" box="[1111,1133,1043,1067]" country="Nigeria" name="Rivers" pageId="1" pageNumber="396">A</collectingRegion>
9,
<collectingRegion id="D315F8016A56FFCBFB1D457ED4996F5A" box="[1162,1183,1043,1067]" country="Nigeria" name="Rivers" pageId="1" pageNumber="396">A</collectingRegion>
9–10 cupuliform,
<collectingRegion id="D315F8016A56FFCBFAFA457ED5856F5A" box="[1389,1411,1043,1067]" country="Nigeria" name="Rivers" pageId="1" pageNumber="396">A</collectingRegion>
11 subovate; terminal labial palpomere securiform, maxillary palpomere securiform (
<figureCitation id="89EA2A666A56FFCBFBFC455BD4C36F3E" box="[1131,1221,1078,1103]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 13</figureCitation>
); labrum strongly bilobate, lobes extending over mandibles (
<figureCitation id="89EA2A666A56FFCBFDEF4537D2CD6F02" box="[632,715,1114,1139]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 11</figureCitation>
); gular sutures very weakly divergent (appearing parallel), post gular process a short, slender, bifurcated central sclerite with paired, weakly defined, setose lobes.
<emphasis id="23A5EAF16A56FFCBFB484513D53E6FE6" box="[1247,1336,1150,1175]" italics="true" pageId="1" pageNumber="396">Thorax.</emphasis>
Pronotal disc without median sulcus, subapical depression reasonably well-defined; postero-lateral pits present (
<figureCitation id="89EA2A666A56FFCBFABB45CFD5866FCA" box="[1324,1408,1186,1211]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 12</figureCitation>
a); procoxal cavities broadly open, prosternal process with small terminal expansion; mesoventrite with rounded anterior process; elytron elongate, disc with ten rows of deep punctures, punctures laterally nodulate (
<figureCitation id="89EA2A666A56FFCBFABC4587D5796E72" box="[1323,1407,1258,1283]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 12</figureCitation>
b), discal vestiture composed of paired sensilla (i.e., each long seta with minute accompanying sensillum at its base), each stria with a single setal pair on the upper anterior rim of each puncture, each interstria with single row of setal pairs; metathoracic wing with CuA1+2 vein narrowly separated from CuA (may appear connected due to continuation of pigmentation to CuA; see
<figureCitation id="89EA2A666A56FFCBFDE44417D2B26EE2" box="[627,692,1402,1427]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 1</figureCitation>
); metendosternite as in
<figureCitation id="89EA2A666A56FFCBFC524417D4336EE2" box="[965,1077,1402,1427]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Figure 10</figureCitation>
.
<emphasis id="23A5EAF16A56FFCBFBD74411D47B6EE2" box="[1088,1149,1404,1427]" italics="true" pageId="1" pageNumber="396">Legs.</emphasis>
Lateral surfaces of tibiae longitudinally carinate; tibial spur formula 2–2–2; tarsomere formula 5–5–5; all basitarsi reduced in length (no visible dorsal surface); tarsal pad formula 4–4–4, basitarsal pads extremely small, pads of T2–4 conspicuous, increasing is size towards T4, pads not divided into paired lobes (i.e., apical margins not divided or emarginate) (
<figureCitation id="89EA2A666A56FFCBFF084767D0F06D52" box="[159,246,1546,1571]" captionStart="FIGURES 14 – 16" captionStartId="3.[151,264,1165,1187]" captionTargetBox="[153,1436,193,1137]" captionTargetId="figure@3.[151,1436,193,1144]" captionTargetPageId="3" captionText="FIGURES 14 – 16. Menieroclerus nigropiceus: (14) female specimen (scale = 1 mm); (15) female mesotarsus showing nonbifid tarsal pads (scale = 0.5 mm); (16) distribution." httpUri="https://zenodo.org/record/257195/files/figure.png" pageId="1" pageNumber="396">Fig. 15</figureCitation>
); each side of pretarsal claw with a triangular basal denticle.
<emphasis id="23A5EAF16A56FFCBFC594767D4466D52" box="[974,1088,1546,1571]" italics="true" pageId="1" pageNumber="396">Abdomen.</emphasis>
Six ventrites.
<emphasis id="23A5EAF16A56FFCBFB7A4767D59A6D52" box="[1261,1436,1546,1571]" italics="true" pageId="1" pageNumber="396">Male genitalia.</emphasis>
Tegmen with phallobasic struts not fused to apodeme (
<figureCitation id="89EA2A666A56FFCBFC814743D38E6D36" box="[790,904,1582,1607]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Figs. 2, 3</figureCitation>
); phallus relatively simple with slender distal process (
<figureCitation id="89EA2A666A56FFCBFF6D473FD1396D1A" box="[250,319,1618,1643]" captionStart="FIGURES 1 – 13" captionStartId="2.[151,264,1825,1847]" captionTargetBox="[162,1436,198,1789]" captionTargetId="figure@2.[162,1447,193,1790]" captionTargetPageId="2" captionText="FIGURES 1 – 13. Menieroclerus nigropiceus: (1) hind wing (‘ a’ showing basal limit of CuA 1 + 2); (2 – 3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view) — (12 a) arrow indicating location of baso-lateral pronotal pit, (12 b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2 – 12), 0.25 mm (Fig. 13)]." httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="1" pageNumber="396">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="116E36E36A56FFCBFF50471BD1AC6DFE" blockId="1.[151,1437,574,1931]" box="[199,426,1654,1679]" pageId="1" pageNumber="396">
<emphasis id="23A5EAF16A56FFCBFF50471BD1266DFE" bold="true" box="[199,288,1654,1679]" pageId="1" pageNumber="396">Gender</emphasis>
. Masculine.
</paragraph>
<paragraph id="116E36E36A56FFC9FF5047F7D4C26E3E" blockId="1.[151,1437,574,1931]" lastBlockId="3.[151,1436,1262,1756]" lastPageId="3" lastPageNumber="398" pageId="1" pageNumber="396">
<emphasis id="23A5EAF16A56FFCBFF5047F7D1CD6DC2" bold="true" box="[199,459,1690,1715]" pageId="1" pageNumber="396">Systematic placement</emphasis>
. By process of elimination,
<taxonomicName id="D6D14D606A56FFCBFC9B47F7D43F6DC2" box="[780,1081,1690,1715]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="species" species="nigropiceus" status="comb. nov.">
<emphasis id="23A5EAF16A56FFCBFC9B47F7D43F6DC2" box="[780,1081,1690,1715]" italics="true" pageId="1" pageNumber="396">Menieroclerus nigropiceus</emphasis>
</taxonomicName>
<emphasis id="23A5EAF16A56FFCBFBD547F7D4AF6DC2" bold="true" box="[1090,1193,1690,1715]" pageId="1" pageNumber="396">
<taxonomicNameLabel id="3896578A6A56FFCBFBD547F7D4AF6DC2" box="[1090,1193,1690,1715]" pageId="1" pageNumber="396">comb. n.</taxonomicNameLabel>
</emphasis>
is negatively defined as a member of the subfamily
<taxonomicName id="D6D14D606A56FFCBFE7C47D3D24C6DA6" box="[491,586,1726,1751]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="subFamily" subFamily="Clerinae">Clerinae</taxonomicName>
simply due to the absence of the following morphological attributes: fourth tarsosomeres minute and hidden between lobes of third tarsomeres (=
<taxonomicName id="D6D14D606A56FFCBFC5F478FD4546D8A" box="[968,1106,1762,1787]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="subFamily" subFamily="Korynetinae">Korynetinae</taxonomicName>
<emphasis id="23A5EAF16A56FFCBFBCB4789D4D66D8A" box="[1116,1232,1762,1787]" italics="true" pageId="1" pageNumber="396">sensu lato</emphasis>
); first tarsomeres well developed, dorsally visible between second tarsomere and tibial distal margin and pretarsal claw tripartite (=
<taxonomicName id="D6D14D606A56FFCBFF004647D0F46C32" box="[151,242,1834,1859]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="subFamily" subFamily="Tillinae">Tillinae</taxonomicName>
); apodeme of male tegmen long and membranously connected to phallobase (= Epiclininae); labrum large, generally not emarginate and eyes with ocular notch absent or minutely developed (=
<taxonomicName id="D6D14D606A56FFCBFBC24623D4FE6C16" box="[1109,1272,1870,1895]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="subFamily" subFamily="Hydnocerinae">Hydnocerinae</taxonomicName>
). No formallyestablished genus-groupings (i.e., tribes) have been proposed for
<taxonomicName id="D6D14D606A56FFCBFC03461FD3F06CFA" box="[916,1014,1906,1931]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="subFamily" subFamily="Clerinae">Clerinae</taxonomicName>
. Several informal clerine sub-groups, however, have been preliminarily proposed: the
<taxonomicName id="D6D14D606A54FFC9FD574583D30D6E76" box="[704,779,1262,1287]" class="Insecta" family="Cleridae" genus="Clerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="23A5EAF16A54FFC9FD574583D30D6E76" box="[704,779,1262,1287]" italics="true" pageId="3" pageNumber="398">Clerus</emphasis>
</taxonomicName>
-series (containing
<taxonomicName id="D6D14D606A54FFC9FC714583D46B6E76" box="[998,1133,1262,1287]" class="Insecta" family="Cleridae" genus="Thanasimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="23A5EAF16A54FFC9FC714583D46B6E76" box="[998,1133,1262,1287]" italics="true" pageId="3" pageNumber="398">Thanasimus</emphasis>
</taxonomicName>
,
<taxonomicName id="D6D14D606A54FFC9FBEA4582D4FD6E77" box="[1149,1275,1263,1286]" class="Insecta" family="Cleridae" genus="Stigmatium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="23A5EAF16A54FFC9FBEA4582D4FD6E77" box="[1149,1275,1263,1286]" italics="true" pageId="3" pageNumber="398">Stigmatium</emphasis>
</taxonomicName>
and
<taxonomicName id="D6D14D606A54FFC9FAA04583D59A6E76" box="[1335,1436,1262,1287]" class="Insecta" family="Cleridae" genus="Omadius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="23A5EAF16A54FFC9FAA04583D59A6E76" box="[1335,1436,1262,1287]" italics="true" pageId="3" pageNumber="398">Omadius</emphasis>
</taxonomicName>
genus-groups), and the
<taxonomicName id="D6D14D606A54FFC9FE39447ED2086E5D" box="[430,526,1299,1324]" class="Insecta" family="Cleridae" genus="Orthrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus">
<emphasis id="23A5EAF16A54FFC9FE39447ED2086E5D" box="[430,526,1299,1324]" italics="true" pageId="3" pageNumber="398">Orthrius</emphasis>
</taxonomicName>
-group (
<bibRefCitation id="75404B126A54FFC9FDFA447ED3356E5A" author="Gerstmeier" box="[621,819,1299,1324]" pageId="3" pageNumber="398" refString="Gerstmeier, R. (2002) Generic concept of clerid taxa related to Clerus (Coleoptera: Cleridae: Clerinae). Entomological Problems, 32 (2), 99 - 111." type="journal article" year="2002">Gerstmeier 2002</bibRefCitation>
;
<bibRefCitation id="75404B126A54FFC9FCD3447ED4876E5A" author="Gerstmeier" box="[836,1153,1299,1324]" pageId="3" pageNumber="398" refString="Gerstmeier, R. &amp; Eberle, J. (2011) Definition and revision of the Orthrius - group of genera (Coleoptera, Cleridae, Clerinae). ZooKeys, 92, 35 - 60. http: // dx. doi. org / 10.3897 / zookeys. 92.1157" type="journal article" year="2011">Gerstmeier &amp; Eberle 2011</bibRefCitation>
), though the securiform terminal maxillary palpomeres of
<taxonomicName id="D6D14D606A54FFC9FD834455D2BA6E3E" box="[532,700,1336,1359]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="nigropiceus" status="comb. nov.">
<emphasis id="23A5EAF16A54FFC9FD834455D2BA6E3E" box="[532,700,1336,1359]" italics="true" pageId="3" pageNumber="398">M. nigropiceus</emphasis>
</taxonomicName>
<emphasis id="23A5EAF16A54FFC9FD54445BD32C6E3E" bold="true" box="[707,810,1334,1359]" pageId="3" pageNumber="398">
<taxonomicNameLabel id="3896578A6A54FFC9FD54445BD32C6E3E" box="[707,810,1334,1359]" pageId="3" pageNumber="398">comb. n.</taxonomicNameLabel>
</emphasis>
prevent its inclusion in any of them.
</paragraph>
<caption id="45AE666B6A55FFC8FF00464CD3CD6CA0" httpUri="https://zenodo.org/record/257194/files/figure.png" pageId="2" pageNumber="397" targetBox="[162,1436,198,1789]" targetPageId="2">
<paragraph id="116E36E36A55FFC8FF00464CD3CD6CA0" blockId="2.[151,1436,1825,2001]" pageId="2" pageNumber="397">
<emphasis id="23A5EAF16A55FFC8FF00464CD1456C46" bold="true" box="[151,323,1825,1847]" pageId="2" pageNumber="397">FIGURES 1–13</emphasis>
.
<taxonomicName id="D6D14D606A55FFC8FEC5464CD2656C46" box="[338,611,1825,1847]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="species" species="nigropiceus">
<emphasis id="23A5EAF16A55FFC8FEC5464CD2656C46" box="[338,611,1825,1847]" italics="true" pageId="2" pageNumber="397">Menieroclerus nigropiceus</emphasis>
</taxonomicName>
: (1) hind wing (‘a’ showing basal limit of CuA1+2); (2–3) male tegmen (ps = phallobasic strut, pa = phallobasic apodeme), (2) dorsal, (3) lateral; (4) male phallus; (5) male spicular fork; (6) male pygidium; (7) female pygidium; (8) male sixth ventrite; (9) female sixth ventrite; (10) metendosternite; (11) male head with arrows indicating each side of bifurcated labrum; (12) male pronotal and elytral base (lateral view)—(12a) arrow indicating location of baso-lateral pronotal pit, (12b) arrow pointing to laterally binodulate elytral puncture; (13) securiform terminal maxillary palpomere [scale bars = 1 mm (Fig. 1), 0.5 mm (Figs. 2–12), 0.25 mm (Fig. 13)].
</paragraph>
</caption>
<caption id="45AE666B6A54FFC9FF0045E0D2A36FB3" httpUri="https://zenodo.org/record/257195/files/figure.png" pageId="3" pageNumber="398" targetBox="[153,1436,193,1137]" targetPageId="3">
<paragraph id="116E36E36A54FFC9FF0045E0D2A36FB3" blockId="3.[151,1435,1165,1218]" pageId="3" pageNumber="398">
<emphasis id="23A5EAF16A54FFC9FF0045E0D1496FD2" bold="true" box="[151,335,1165,1187]" pageId="3" pageNumber="398">FIGURES 14–16</emphasis>
.
<taxonomicName id="D6D14D606A54FFC9FEC945E0D2766FD5" box="[350,624,1165,1188]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="species" species="nigropiceus">
<emphasis id="23A5EAF16A54FFC9FEC945E0D2766FD5" box="[350,624,1165,1188]" italics="true" pageId="3" pageNumber="398">Menieroclerus nigropiceus</emphasis>
</taxonomicName>
: (14) female specimen (scale = 1 mm); (15) female mesotarsus showing nonbifid tarsal pads (scale = 0.5 mm); (16) distribution.
</paragraph>
</caption>
<paragraph id="116E36E36A54FFC9FF504436D45A6DE5" blockId="3.[151,1436,1262,1756]" pageId="3" pageNumber="398">
The taxon-diverse subfamily
<taxonomicName id="D6D14D606A54FFC9FD8A4436D27A6E05" box="[541,636,1371,1396]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Clerinae">Clerinae</taxonomicName>
is yet to be adequately defined morphologically or established as a monophylum (
<bibRefCitation id="75404B126A54FFC9FEAC4413D1DE6EE6" author="Kolibac" box="[315,472,1406,1431]" pageId="3" pageNumber="398" refString="Kolibac, J. (1997) Classification of the subfamilies of Cleridae (Coleoptera: Cleroidea). Acta Musei Moraviae, Scientiae Naturales, 81, 307 - 361." type="journal article" year="1997">Kolibáč 1997</bibRefCitation>
;
<bibRefCitation id="75404B126A54FFC9FE744413D2606EE6" author="Opitz" box="[483,614,1406,1431]" pageId="3" pageNumber="398" refString="Opitz, W. (2010) Classification, natural history, phylogeny, and subfamily composition of the Cleridae and generic content of the subfamilies (Coleoptera: Cleroidea). Entomologica Basiliensia et Collectionis Frey, 32, 31 - 128." type="journal article" year="2010">Opitz 2010</bibRefCitation>
).
<collectingRegion id="D315F8016A54FFC9FDEF4412D28A6EE6" box="[632,652,1407,1431]" country="Nigeria" name="Rivers" pageId="3" pageNumber="398">A</collectingRegion>
preliminary molecular phylogeny-estimate of
<taxonomicName id="D6D14D606A54FFC9FB004413D4F06EE6" box="[1175,1270,1406,1431]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="family">Cleridae</taxonomicName>
suggested that the exclusion of
<taxonomicName id="D6D14D606A54FFC9FEC244CED1F46ECD" box="[341,498,1443,1468]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Hydnocerinae">Hydnocerinae</taxonomicName>
from
<taxonomicName id="D6D14D606A54FFC9FDAE44CED29E6ECD" box="[569,664,1443,1468]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Clerinae">Clerinae</taxonomicName>
renders the latter paraphyletic (
<bibRefCitation id="75404B126A54FFC9FB9544CED4DF6ECA" author="Gunter" box="[1026,1241,1443,1468]" pageId="3" pageNumber="398" refString="Gunter, N. L., Leavengood, J. M., Bartlett, J. S., Chapman, E. G. &amp; Cameron, S. L. (2013) A molecular phylogeny of the checkered beetles and a description of Epiclininae a new subfamily (Coleoptera: Cleroidea: Cleridae). Systematic Entomology, 38 (3), 626 - 636. http: // dx. doi. org / 10.1111 / syen. 12019" type="journal article" year="2013">
Gunter
<emphasis id="23A5EAF16A54FFC9FBCD44C9D4926ECD" box="[1114,1172,1443,1468]" italics="true" pageId="3" pageNumber="398">et al.</emphasis>
2013
</bibRefCitation>
).
<collectingRegion id="D315F8016A54FFC9FB7A44CED5076ECA" box="[1261,1281,1443,1467]" country="Nigeria" name="Rivers" pageId="3" pageNumber="398">A</collectingRegion>
morphologybased character analysis of clerine and hydnocerine (plus anthicoclerine) genera could be a first step in understanding if/how the genera of
<taxonomicName id="D6D14D606A54FFC9FDB44486D2846D75" box="[547,642,1515,1540]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Clerinae">Clerinae</taxonomicName>
and
<taxonomicName id="D6D14D606A54FFC9FD204486D3526D75" box="[695,852,1515,1540]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Hydnocerinae">Hydnocerinae</taxonomicName>
are related. The threefold usefulness of such a study would be to: a) test the monophyly of
<taxonomicName id="D6D14D606A54FFC9FDD94763D2AB6D56" box="[590,685,1550,1575]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Clerinae">Clerinae</taxonomicName>
in relation to the inclusion/exclusion of
<taxonomicName id="D6D14D606A54FFC9FBEA4763D5186D56" box="[1149,1310,1550,1575]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Hydnocerinae">Hydnocerinae</taxonomicName>
; b) test the monophyly of previously proposed informal clerine groupings; and, c) potentially identify genus-groupings among genera not yet arranged into informal groupings.
<collectingRegion id="D315F8016A54FFC9FD5F473AD2DA6D1E" box="[712,732,1623,1647]" country="Nigeria" name="Rivers" pageId="3" pageNumber="398">A</collectingRegion>
moderately large scale study such as this would be necessary before the systematic position of
<taxonomicName id="D6D14D606A54FFC9FD9C4716D2A86DE5" box="[523,686,1659,1684]" class="Insecta" family="Cleridae" genus="Menieroclerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="23A5EAF16A54FFC9FD9C4716D2A86DE5" box="[523,686,1659,1684]" italics="true" pageId="3" pageNumber="398">Menieroclerus</emphasis>
</taxonomicName>
<emphasis id="23A5EAF16A54FFC9FD234716D3026DE5" bold="true" box="[692,772,1659,1684]" pageId="3" pageNumber="398">
<taxonomicNameLabel id="3896578A6A54FFC9FD234716D3026DE5" box="[692,772,1659,1684]" pageId="3" pageNumber="398" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
within
<taxonomicName id="D6D14D606A54FFC9FCCE4716D3BE6DE5" box="[857,952,1659,1684]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subFamily" subFamily="Clerinae">Clerinae</taxonomicName>
is understood.
</paragraph>
<paragraph id="116E36E36A54FFC9FF5047F3D1116DAD" blockId="3.[151,1436,1262,1756]" pageId="3" pageNumber="398">
<emphasis id="23A5EAF16A54FFC9FF5047F3D1BC6DC6" bold="true" box="[199,442,1694,1719]" pageId="3" pageNumber="398">Known distribution</emphasis>
(
<figureCitation id="89EA2A666A54FFC9FE5847F3D22F6DC6" box="[463,553,1694,1719]" captionStart="FIGURES 14 – 16" captionStartId="3.[151,264,1165,1187]" captionTargetBox="[153,1436,193,1137]" captionTargetId="figure@3.[151,1436,193,1144]" captionTargetPageId="3" captionText="FIGURES 14 – 16. Menieroclerus nigropiceus: (14) female specimen (scale = 1 mm); (15) female mesotarsus showing nonbifid tarsal pads (scale = 0.5 mm); (16) distribution." httpUri="https://zenodo.org/record/257195/files/figure.png" pageId="3" pageNumber="398">Fig. 16</figureCitation>
). Ethiopia, Kenya, Namibia, Republic of South Africa, Tanzania, Yemen, Zimbabwe.
</paragraph>
</subSubSection>
</treatment>
</document>
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