169 lines
16 KiB
XML
169 lines
16 KiB
XML
<document id="1059F70F5A7FAB080026ABB13FFF2C87" ID-CLB-Dataset="28831" ID-DOI="http://dx.doi.org/10.3897/zookeys.796.21877" ID-GBIF-Dataset="0476ce18-4e9c-4ee6-8d6e-99d1146cdbf4" ID-PMC="PMC6256130" ID-Pensoft-Pub="1313-2970-796-215" ID-PubMed="30505157" ID-ZooBank="CC96882F71234A2D83457C5E36F817C1" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-796-215" ModsDocOrigin="ZooKeys 796" ModsDocTitle="Review of the subgenus Plumiger of Myrmecophyes, with description of a new species (Heteroptera, Miridae, Halticini)" checkinTime="1542332380955" checkinUser="pensoft" docAuthor="Konstantinov, Fedor V. & Simov, Nikolay" docDate="2018" docId="8746D900922F3744799FF829719148A8" docLanguage="en" docName="ZooKeys 796: 215-239" docOrigin="ZooKeys 796" docSource="http://dx.doi.org/10.3897/zookeys.796.21877" docTitle="Myrmecophyes nasutus Drapolyuk 1989" docType="treatment" docVersion="7" lastPageNumber="230" masterDocId="6D0C0038FFD8FFBFFFD8867C7F0D2327" masterDocTitle="Review of the subgenus Plumiger of Myrmecophyes, with description of a new species (Heteroptera, Miridae, Halticini)" masterLastPageNumber="239" masterPageNumber="215" originalUpdateDomain="Boston" originalUpdateTime="1542332380955" originalUpdateUser="pensoft" pageNumber="229" updateTime="1701382271246" updateUser="ExternalLinkService">
|
||
<mods:mods id="79D3C2AFACC4C538C3ED3E89A98868D3" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="E036CD4A148D74E67B8F37C84DA69722">
|
||
<mods:title id="A5918BEDFDFC12322D779FC51F2FF6EB">Review of the subgenus Plumiger of Myrmecophyes, with description of a new species (Heteroptera, Miridae, Halticini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="E24BA66D38D19307B3F4EE21F37F9AA7" type="personal">
|
||
<mods:role id="C658FF9F4C8B1F57E74A6D8836B6C39A">
|
||
<mods:roleTerm id="BFBED5D981965E7849184355D4C55C37">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0C7D0FC423C019D0042025C69DC794C1">Konstantinov, Fedor V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="69EC80C908787A1AA3C0B69E163BA33F" type="personal">
|
||
<mods:role id="E8184FA0719AB5E73EBA8B7B55FF0CE3">
|
||
<mods:roleTerm id="83C3B0D3FBC98134F10F28255339AC6D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0E01D6356597EF286907929EB888DE44">Simov, Nikolay</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="14A04BA0A309C05EFBB4B3D1301CD832">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F2CDF879DF22494E2D896B04DFA13813" type="host">
|
||
<mods:titleInfo id="EDAEA4D4C39135B5DBCE1587AB1A41D9">
|
||
<mods:title id="44D430349CAC6F0B3C074F3AB7051AAE">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="6ACF9F31B6545570B6A0DBDC8C7D8C31">
|
||
<mods:date id="2E39C49D9C654B9E3A76FC5234307FF5">2018</mods:date>
|
||
<mods:detail id="780591E2BFD34FBF0700EE3F974270D3" type="volume">
|
||
<mods:number id="67F209A75F9BFC411336F7A9779784B7">796</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="045C67097B25D503278D2B8E5B12DEEF" unit="page">
|
||
<mods:start id="66DC4AFA01BA366D79840792D7B4002D">215</mods:start>
|
||
<mods:end id="4B61A8746931955B17AF262DC8B3B6B9">239</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="2BCBB7B153F6FFE9BD3A840C1700368F">
|
||
<mods:url id="9E136F8DF504F87FB44E4285E9C56E61">http://dx.doi.org/10.3897/zookeys.796.21877</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="7FA3E3BB6E8FD5B646CE645445CF93B6">journal article</mods:classification>
|
||
<mods:identifier id="945F6D7D7E4DCAF4DD5BBCFB5CF92DA4" type="DOI">http://dx.doi.org/10.3897/zookeys.796.21877</mods:identifier>
|
||
<mods:identifier id="A477B3FC68C59ECC54982E7E110C9ADA" type="Pensoft-Pub">1313-2970-796-215</mods:identifier>
|
||
<mods:identifier id="D01D80813AFDDD9441125C5C779B2CF3" type="ZooBank">CC96882F71234A2D83457C5E36F817C1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="8746D900922F3744799FF829719148A8" ID-GBIF-Taxon="149667424" LSID="urn:lsid:plazi:treatment:8746D900922F3744799FF829719148A8" httpUri="http://treatment.plazi.org/id/8746D900922F3744799FF829719148A8" lastPageId="15" lastPageNumber="230" pageId="14" pageNumber="229">
|
||
<subSubSection id="C2EAD4DD9643D7B79C78F246FDAC8D85" pageId="14" pageNumber="229" type="nomenclature">
|
||
<paragraph id="99A1C815D50ACC49D2DBFFEFF719912F" pageId="14" pageNumber="229">
|
||
<taxonomicName id="DBA05E5C3DD3753A42278C7E30F58991" ID-CoL="45B8N" authority="Drapolyuk, 1989" authorityName="Drapolyuk" authorityYear="1989" class="Insecta" family="Miridae" genus="Myrmecophyes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmecophyes nasutus" order="Hemiptera" pageId="14" pageNumber="229" phylum="Arthropoda" rank="species" species="nasutus">Myrmecophyes nasutus Drapolyuk, 1989</taxonomicName>
|
||
Figs 11-13, 27, 41-43, 47, 48, 54, 57, 61, 64
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F1DD6620B3A3F256F989869E854301B4" pageId="14" pageNumber="229" type="reference_group">
|
||
<paragraph id="1F4AEC0935CF3AF82F1BD2B6D2DFD807" pageId="14" pageNumber="229">
|
||
<taxonomicName id="5D0D502969680D61C56D08542E8E3B93" class="Insecta" family="Miridae" genus="Myrmecophyes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Myrmecophyes (Plumiger) nasutus" order="Hemiptera" pageId="14" pageNumber="229" phylum="Arthropoda" rank="species" species="nasutus" subGenus="Plumiger">Myrmecophyes (Plumiger) nasutus</taxonomicName>
|
||
Drapolyuk, 1989: 125; figs 40-45, 52, 53.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="230B3A09288567CF53C1F76EB297887B" pageId="14" pageNumber="229" type="materials_examined">
|
||
<paragraph id="0DE7EEA20560D67200C252D2F906331F" pageId="14" pageNumber="229">Material examined.</paragraph>
|
||
<paragraph id="845CB79A9BF3283E762AE835C3B46FAA" pageId="14" pageNumber="229">
|
||
Holotype: GEORGIA: Racha-Lechkhumi and Kvemo Svaneti:
|
||
<normalizedToken id="91549F0A2ACB6498D6CEAE593DEBECFF" originalValue="Korel’dash">Korel'dash</normalizedToken>
|
||
,
|
||
<geoCoordinate id="16D5246039D5C926BCEAAD4F0D0E9399" direction="north" orientation="latitude" precision="5" value="42.9183">42.9183°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3ADA9076119DF05D46BFD00D59E1F048" direction="east" orientation="longitude" precision="55" value="43.144">43.144°E</geoCoordinate>
|
||
, 26 Jul 1957, Akramovskaya, 1♂ (AMNH_PBI 00261441) (ZISP). Paratypes: GEORGIA: Racha-Lechkhumi and Kvemo Svaneti:
|
||
<normalizedToken id="040C4A87EDC92A66D711D4967109B392" originalValue="Korel’dash">Korel'dash</normalizedToken>
|
||
,
|
||
<geoCoordinate id="EC819FF4C5BF1DA2583B3B766DAAAFB9" direction="north" orientation="latitude" precision="5" value="42.9183">42.9183°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D187878C328702D17D9D75F235700AD6" direction="east" orientation="longitude" precision="55" value="43.144">43.144°E</geoCoordinate>
|
||
, 26 Jul 1957, Akramovskaya, 2♀ (AMNH_PBI 00271200, AMNH_PBI 00271201) (ZISP); 30 Jul 1957, Akramovskaya, 1♂ (AMNH_PBI 00261442), 23♀ (AMNH_PBI 00261639-AMNH_PBI 00261661) (ZISP). Lentekhi, Svanetia,
|
||
<geoCoordinate id="8A6EA3818AAB2170B93BBA95D36E425C" direction="north" orientation="latitude" precision="1" value="42.78333">42.78333°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="DB55B8858A338333B7F4536CB5BD55F3" direction="east" orientation="longitude" precision="5555" value="42.7">42.7°E</geoCoordinate>
|
||
, 10 Aug 1957, Akramovskaya, 3♀ (AMNH_PBI 00261674-AMNH_PBI 00261676) (ZISP). Tsena, Svanetia,
|
||
<geoCoordinate id="FE5963BD671BEFF55BB56374DF370136" direction="north" orientation="latitude" precision="5" value="42.8789">42.8789°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="8876CADD78C3C20724FE512D7BA43F98" direction="east" orientation="longitude" precision="5" value="43.1519">43.1519°E</geoCoordinate>
|
||
, 25 Jul 1957, Akramovskaya, 15♀ (AMNH_PBI 00271197-AMNH_PBI 00271199, AMNH_PBI 00261662-AMNH_PBI 00261673) (ZISP). Zarsky Pass, Lentekhi Distr.,
|
||
<geoCoordinate id="89E9DD7ACF17B9ED446242D21831D830" direction="north" orientation="latitude" precision="1" value="42.91666">42.91666°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D31FB86C328972775D1A5D3B191CB265" direction="east" orientation="longitude" precision="1" value="43.16666">43.16666°E</geoCoordinate>
|
||
, 27 Jul 1957, Akramovskaya, 3♀ (AMNH_PBI 00261677-AMNH_PBI 00261679) (ZISP). Mtskheta-Mtianeti: Kazbek Mt.,
|
||
<geoCoordinate id="871BFC6545BF56464193B65D45B1F488" direction="north" orientation="latitude" precision="1" value="42.68333">42.68333°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="647228A4C21737B132FD2C7283872D42" direction="east" orientation="longitude" precision="1" value="44.46666">44.46666°E</geoCoordinate>
|
||
, 17 Jul 1910, Unknown collector, 1♀ (AMNH_PBI 00261685) (ZISP).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="51DCC0CAA68CAEDE90179EE48C372213" pageId="14" pageNumber="229" type="diagnosis">
|
||
<paragraph id="FD8816E1ED6CAD78C4B45DAEC2E1C6D6" pageId="14" pageNumber="229">Diagnosis.</paragraph>
|
||
<paragraph id="C839D1CFA8251FBEF2F2529B410C31C2" pageId="14" pageNumber="229">
|
||
Distinguished from other
|
||
<taxonomicName id="11DC1AF35601C14194207E3A18CF9FA5" genus="Plumiger" lsidName="Plumiger" pageId="14" pageNumber="229" rank="genus">Plumiger</taxonomicName>
|
||
spp. by the larger size (Figs 11-13), distinctly bulging clypeus (Figure 27), non-shiny head and abdomen, harpoon-shaped left paramere with two subapical recurved denticles (Figs 41, 42), and long, narrow, distinctly attenuate sclerotized rings of dorsal labiate plate (Figs 54, 57). The diagnosis and remarks for
|
||
<taxonomicName id="3AFB0653D831F33D221DC1AC4B4C50FC" lsidName="M. tomi" pageId="14" pageNumber="229" rank="species" species="tomi">M. tomi</taxonomicName>
|
||
include a discussion of the distinctive features.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5A1A468F0D2F3343D3E1F2C8FE61037" pageId="14" pageNumber="229" type="redescription">
|
||
<paragraph id="8FDDE3C4CB559464E6CED0DA577F197F" pageId="14" pageNumber="229">Redescription.</paragraph>
|
||
<paragraph id="130DFB60D42ADB12AA26346948B421CF" pageId="14" pageNumber="229">
|
||
Male. Total length 3.8-3.9. Coloration (Figure 11): Black, with pale brown apices of femora and dirty white transverse stripe along apex of wing pad. Surface and vestiture: As in generic diagnosis. Structure: Body 4.2-4.6
|
||
<normalizedToken id="466A67A8548F1C12E5840061979AC91D" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Head: Clypeus strongly bulging apically, concave and apically rectangular in lateral view (Figure 27), apically dilated, broadly rounded in frontal view; vertex 2.1-2.2
|
||
<normalizedToken id="A4C29C3E843575851694FB89D92299F1" originalValue="×">x</normalizedToken>
|
||
as wide as eye; antennal segment I strongly swollen, relatively long, 1.3-1.4
|
||
<normalizedToken id="78EA292A56270BEE4CFFCE19587A33DB" originalValue="×">x</normalizedToken>
|
||
as long as length of pronotum, segment II gradually curved along entire length and somewhat flattened apically, long, 1.8-1.9
|
||
<normalizedToken id="1539596242A479865FD6CBA7A5A80CB3" originalValue="×">x</normalizedToken>
|
||
as long as I, 1.8-1.9
|
||
<normalizedToken id="139C3328F622640E88C444EBFBB55CB0" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle, and 1.2-1.3
|
||
<normalizedToken id="D209F6C9E2E0E5AC3CC15CC0B27E0194" originalValue="×">x</normalizedToken>
|
||
as long as width of head. Thorax: Pronotum at middle 1.3-1.5
|
||
<normalizedToken id="0FF1265E03770ECD45C3862901889AC8" originalValue="×">x</normalizedToken>
|
||
as wide as long, 0.7
|
||
<normalizedToken id="F8EE836C3B0C49943C4EE3C307EE865D" originalValue="×">x</normalizedToken>
|
||
as wide as head; hind tibia 3.8-4.2
|
||
<normalizedToken id="3C67582A6F67C0352105137860C64777" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Genitalia: genital capsule about 0.4 of abdomen; left paramere as in
|
||
<taxonomicName id="01E3EC9D0E4501A31B49F8025D971762" lsidName="M. tomi" pageId="14" pageNumber="229" rank="species" species="tomi">M. tomi</taxonomicName>
|
||
but slightly larger, with two subapical recurved denticles (Figs 41, 42); right paramere similar to that of
|
||
<taxonomicName id="E8305982E5C32B33EFF07C81FCE6F21A" lsidName="M. tomi" pageId="14" pageNumber="229" rank="species" species="tomi">M. tomi</taxonomicName>
|
||
but larger and equipped with larger apical tooth on inner margin (Figure 43); endosoma of aedeagus voluminous, membranous, with three large, folded, minutely dentate and slightly sclerotized eversible lobes (Figs 47, 48).
|
||
</paragraph>
|
||
<paragraph id="4E1F21F7797D6D8502E0E7EE8347188E" pageId="14" pageNumber="229">
|
||
Female. Body larger on average, total length 4.2-4.5. Coloration: Similar to male, but antennal segment I ranging from dirty orange to entirely black, coxae usually entirely or at least apically dirty orange, femora from entirely dirty orange to black, with apices dirty orange (Figs 12, 13). Surface and vestiture: As in
|
||
<taxonomicName id="6D7FD0C280262C70F70CAB6C6C9E972C" lsidName="M. tomi" pageId="14" pageNumber="229" rank="species" species="tomi">M. tomi</taxonomicName>
|
||
sp. n. Structure: Body 4.1-4.6
|
||
<normalizedToken id="7D6ECBE3B598ED5375146E5F9D603E0A" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Head: Clypeus convex, not bulging apically; vertex 2.0-2.4
|
||
<normalizedToken id="8FC6EBF36898D054150BD1774C60546C" originalValue="×">x</normalizedToken>
|
||
as wide as eye; antennal segment I slightly and uniformly swollen, short, equal to length of pronotum, segment II thin, straight and rod-shaped, 2.2-2.4
|
||
<normalizedToken id="2189631431B4B31E401DA7B48CB843B3" originalValue="×">x</normalizedToken>
|
||
as long as I, 1.4-1.6
|
||
<normalizedToken id="5D5FD91EEB9E8B5811D1932B8711FC06" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle, and 1.1-1.2
|
||
<normalizedToken id="B9896A04A7608E22CA8E742273783023" originalValue="×">x</normalizedToken>
|
||
as long as width of head. Thorax: Pronotum at middle 1.5
|
||
<normalizedToken id="7E170B47DDEA1F8D82AA69790B1BF297" originalValue="×">x</normalizedToken>
|
||
as wide as long, 0.7-.0.8
|
||
<normalizedToken id="9D51F599F6EF79F3F1492F16E4323980" originalValue="×">x</normalizedToken>
|
||
as wide as head; hind tibia 3.2-3.3
|
||
<normalizedToken id="A77D784EE8BF797C36379BBEE18EB56E" originalValue="×">x</normalizedToken>
|
||
as long as width of pronotum at middle. Genitalia: Sclerotized rings of dorsal labiate plate long and narrow, gradually curved, with apices gradually attenuated (Figs 54, 61); interramal sclerites flat, finely striated (Figure 57).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3720DA7F1E74D5086BED925CBB6CAADE" pageId="15" pageNumber="230" type="distribution">
|
||
<paragraph id="5E99580DB5BDEB719E9313DC0D692A90" pageId="15" pageNumber="230">
|
||
<pageBreakToken id="DC16D14283473058E149F0B37E080FBE" pageId="15" pageNumber="230" start="start">Distribution</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="43083E053F6511FB704BD8A34D65B64A" pageId="15" pageNumber="230">Known from several localities in northern Georgia along the southern slopes of the Greater Caucasus Mountain range (Figure 64).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F3538110A434CF822C1A0D85BCF69B90" pageId="15" pageNumber="230" type="discussion">
|
||
<paragraph id="87AF7D9BB68A8AD33C7B635AF5374FCC" pageId="15" pageNumber="230">Discussion.</paragraph>
|
||
<paragraph id="DF8E1E1A5EA1B9CBBB81B205EF98CF61" pageId="15" pageNumber="230">
|
||
<bibRefCitation id="AD83DD34C98C97DC719969AEFDB8050B" author="Drapolyuk, IS" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="20" pageNumber="235" pagination="125 - 135" title="Species of the genus Myrmecophyes Fieb. (Heteroptera, Miridae) from the Caucasus." volume="68" year="1989">Drapolyuk (1989)</bibRefCitation>
|
||
diagnosed
|
||
<taxonomicName id="B334A1CB4C6B790DEB993ABCC6D775F2" lsidName="M. nasutus" pageId="15" pageNumber="230" rank="species" species="nasutus">M. nasutus</taxonomicName>
|
||
as having the aedeagus with a single spicula, although her figure 45 shows a membranous and finely dentate lobe of the endosoma. We observed that the eversible lobes of the endosoma in
|
||
<taxonomicName id="6E08363D29D642771F4000708F93826E" lsidName="M. nasutus" pageId="15" pageNumber="230" rank="species" species="nasutus">M. nasutus</taxonomicName>
|
||
and
|
||
<taxonomicName id="E8237B719CA1CC9416929A899A52ED34" lsidName="M. tomi" pageId="15" pageNumber="230" rank="species" species="tomi">M. tomi</taxonomicName>
|
||
sp. n. are faintly sclerotized but effectively devoid of spiculae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |