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<mods:title id="0F4E1AAA4571AF923D41AF0348E56310">Redescription of the genus Cheleocloeon Wuillot &amp; Gillies 1993 (Ephemeroptera: Baetidae) with descriptions of three new species from Zambia and Uganda</mods:title>
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Genus 
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<emphasis id="FDC181224F29FF8D21CF0F4FFE7CED07" bold="true" box="[238,403,1842,1868]" italics="true" pageId="1" pageNumber="136">Cheleocloeon</emphasis>
<bibRefCitation id="AB2420C14F29FF8D20BB0F4EFD5EED06" author="Wuillot" box="[410,689,1843,1869]" pageId="1" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot &amp; Gillies 1993</bibRefCitation>
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(
<figureCitation id="578E41B54F29FF8D21BE0F28FEF1ED25" box="[159,286,1877,1902]" captionStart-0="FIGURES 1 – 6. 1 – 5" captionStart-1="FIGURES 7 – 18. 7 – 15" captionStart-10="FIGURES 99 – 102" captionStart-11="FIGURES 103 – 106" captionStart-12="FIGURES 107 – 112. 107 – 109" captionStart-13="FIGURES 113 – 116" captionStart-14="FIGURES 117 – 119" captionStart-15="FIGURES 120 – 121" captionStart-16="FIGURES 122 – 127" captionStart-17="FIGURES 128 – 133" captionStart-2="FIGURES 19 – 29. 19 – 23" captionStart-3="FIGURES 30 – 57" captionStart-4="FIGURES 58 – 63" captionStart-5="FIGURES 64 – 70" captionStart-6="FIGURES 71 – 78" captionStart-7="FIGURES 79 – 86. 79 – 85" captionStart-8="FIGURES 87 – 94" captionStart-9="FIGURES 95 – 98. 95 – 96" captionStartId-0="4.[151,264,1942,1965]" captionStartId-1="5.[151,264,1436,1459]" captionStartId-10="22.[151,264,1439,1462]" captionStartId-11="23.[151,264,1757,1780]" captionStartId-12="24.[151,264,1860,1883]" captionStartId-13="25.[151,264,1652,1675]" captionStartId-14="26.[151,264,2002,2025]" captionStartId-15="27.[151,264,1018,1041]" captionStartId-16="28.[151,264,1786,1809]" captionStartId-17="30.[151,264,1616,1639]" captionStartId-2="7.[151,264,1883,1906]" captionStartId-3="8.[151,264,1554,1577]" captionStartId-4="11.[151,264,1736,1759]" captionStartId-5="12.[151,264,1899,1922]" captionStartId-6="16.[151,264,1837,1860]" captionStartId-7="18.[151,264,1620,1643]" captionStartId-8="20.[151,264,1528,1551]" captionStartId-9="21.[151,264,1779,1802]" captionTargetBox-0="[168,1406,201,1903]" captionTargetBox-1="[226,1346,181,1410]" captionTargetBox-10="[162,1435,193,1418]" captionTargetBox-11="[227,1341,194,1740]" captionTargetBox-12="[177,1387,202,1819]" captionTargetBox-13="[151,1435,193,1630]" captionTargetBox-14="[151,1436,193,1980]" captionTargetBox-15="[151,1436,193,997]" captionTargetBox-16="[150,1435,303,1764]" captionTargetBox-17="[177,1402,195,1589]" captionTargetBox-2="[156,1432,199,1855]" captionTargetBox-3="[153,1432,201,1529]" captionTargetBox-4="[151,1434,176,1715]" captionTargetBox-5="[150,1436,193,1878]" captionTargetBox-6="[151,1434,193,1816]" captionTargetBox-7="[151,1435,193,1599]" captionTargetBox-8="[167,1412,193,1510]" captionTargetBox-9="[254,1331,471,1750]" captionTargetId-0="figure@4.[168,1407,193,1913]" captionTargetId-1="figure@5.[226,1360,169,1415]" captionTargetId-10="figure@22.[151,1436,193,1419]" captionTargetId-11="figure@23.[220,1347,193,1741]" captionTargetId-12="figure@24.[177,1400,193,1824]" captionTargetId-13="figure@25.[151,1437,193,1632]" captionTargetId-14="figure@26.[151,1436,193,1981]" captionTargetId-15="figure@27.[151,1436,193,997]" captionTargetId-16="figure@28.[150,1436,303,1764]" captionTargetId-17="figure@30.[177,1402,195,1589]" captionTargetId-2="figure@7.[151,1436,193,1862]" captionTargetId-3="figure@8.[151,1436,193,1533]" captionTargetId-4="figure@11.[151,1435,159,1715]" captionTargetId-5="figure@12.[150,1436,193,1878]" captionTargetId-6="figure@16.[151,1435,193,1816]" captionTargetId-7="figure@18.[151,1435,193,1600]" captionTargetId-8="figure@20.[167,1413,193,1511]" captionTargetId-9="figure@21.[240,1347,467,1758]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="22" captionTargetPageId-11="23" captionTargetPageId-12="24" captionTargetPageId-13="25" captionTargetPageId-14="26" captionTargetPageId-15="27" captionTargetPageId-16="28" captionTargetPageId-17="30" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="11" captionTargetPageId-5="12" captionTargetPageId-6="16" captionTargetPageId-7="18" captionTargetPageId-8="20" captionTargetPageId-9="21" captionText-0="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." captionText-1="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." captionText-10="FIGURES 99 – 102. Genitals of Cheleocloeon clavifolium sp. n. 99 – 100, cuticle treated with alkali, ventral view: 99, with focus on gonovectes; 100, with focus on unistyligers. 101, cuticle of left half of penis treated by alkali; 102, left half of penis with muscles and gonoduct. Abbreviations: ap, apodeme served for attachment of gonovectal muscle; gd, gonoduct; gv, gonovectis; pb, penial bridge." captionText-11="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." captionText-12="FIGURES 107 – 112. 107 – 109, Cheleocloeon lancetofolium sp. n.; 110 – 112, Ch. truncifolium sp. n. 107, 111, subimaginal exuviae of genitals; 108, 110, imaginal genitals, ventral view (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines; see Fig. 105); 109, 112, hind wing." captionText-13="FIGURES 113 – 116. Cheleocloeon clavifolium sp. n.: 113, 114, male imagoes; 115, abdominal segments II – V of male imago; 116, female imago." captionText-14="FIGURES 117 – 119. Cheleocloeon lancetofolium sp. n.: 117, 118, male imagoes; 119, female imago." captionText-15="FIGURES 120 – 121. Cheleocloeon truncifolium sp. n.: 120, head and thorax of male imago (holotype); 121, female subimago." captionText-16="FIGURES 122 – 127. Eggs. 122 – 123, Cheleocloeon yolandae; 124 – 125, Cheleocloeon clavifolium sp. n., 126 – 127, Cheleocloeon lancetofolium sp. n." captionText-17="FIGURES 128 – 133. Eggs. 128 – 131, Cheleocloeon truncifolium sp. n.; 132 – 133, Cheleocloeon sp. 2." captionText-2="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." captionText-3="FIGURES 30 – 57. Tergalii I – VII: 30 – 36, Cheleocloeon clavifolium sp. n.; 37 – 43, Ch. lancetofolium sp. n.; 44 – 50, Ch. truncifolium sp. n. (male, holotype); 51 – 57, Ch. truncifolium sp. n. (female)." captionText-4="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." captionText-5="FIGURES 64 – 70. Details of exuviae of last instar larva (64 – 69 under the same magnification): 64 – 66, Cheleocloeon yolandae; 67 – 70, Ch. carinatum." captionText-6="FIGURES 71 – 78. Details of exuviae of last instar larvae of Cheleocloeon clavifolium sp. n. under the same magnification (72, middle leg; 77, abdomen of holotype)." captionText-7="FIGURES 79 – 86. 79 – 85, details of exuviae of last instar larvae under the same magnification: 79 – 81, Cheleocloeon lancetofolium sp. n.; 82 – 84, Ch. truncifolium sp. n. (83, holotype); 85, Cheleocloeon sp. 2; 86, abdomen of female subimago of the same specimen." captionText-8="FIGURES 87 – 94. Details of exuviae of subimago. 87 – 89, Cheleocloeon clavifolium sp. n.: 87 – 88, left half of mesonotum of different specimens; 89, pterothorax sterna and pleura. 90 – 92, Ch. truncifolium sp. n.: 90, left half of mesonotum (holotype); 91, abdominal segments IV – V; 92, left pleuron. 93, Ch. yolandae, left half of mesonotum; 94, Cheleocloeon sp. 1, abdominal segments III – V. Abbreviations: lpnc, lateropostnotal crest; psa, postsubalar sclerite; sa, subalar sclerite." captionText-9="FIGURES 95 – 98. 95 – 96, Cheleocloeon clavifolium sp. n.: 95, larval protogonostyli and subimaginal gonostyli developing under larval cuticle; 96, subimaginal gonostyli of mature larva. 97 – 98, Cheleocloeon sp. 1: 97, subimaginal exuviae of genitals, 98, genitals of male imago (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles and gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri-0="https://zenodo.org/record/270597/files/figure.png" httpUri-1="https://zenodo.org/record/270598/files/figure.png" httpUri-10="https://zenodo.org/record/270607/files/figure.png" httpUri-11="https://zenodo.org/record/270608/files/figure.png" httpUri-12="https://zenodo.org/record/270609/files/figure.png" httpUri-13="https://zenodo.org/record/270610/files/figure.png" httpUri-14="https://zenodo.org/record/270611/files/figure.png" httpUri-15="https://zenodo.org/record/270612/files/figure.png" httpUri-16="https://zenodo.org/record/270613/files/figure.png" httpUri-17="https://zenodo.org/record/270614/files/figure.png" httpUri-2="https://zenodo.org/record/270599/files/figure.png" httpUri-3="https://zenodo.org/record/270600/files/figure.png" httpUri-4="https://zenodo.org/record/270601/files/figure.png" httpUri-5="https://zenodo.org/record/270602/files/figure.png" httpUri-6="https://zenodo.org/record/270603/files/figure.png" httpUri-7="https://zenodo.org/record/270604/files/figure.png" httpUri-8="https://zenodo.org/record/270605/files/figure.png" httpUri-9="https://zenodo.org/record/270606/files/figure.png" pageId="1" pageNumber="136">Figs 1–133</figureCitation>
)
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Group 
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<emphasis id="FDC181224F29FF8D21FF0FE0FE8AEDF8" box="[222,357,1949,1971]" italics="true" pageId="1" pageNumber="136">dimorphicum</emphasis>
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of the genus 
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<emphasis id="FDC181224F29FF8D20CC0FE0FDB5EDF8" box="[493,602,1949,1971]" italics="true" pageId="1" pageNumber="136">Afroptilum</emphasis>
: 
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<bibRefCitation id="AB2420C14F29FF8D23460FE0FD07EDF8" author="Gillies" box="[615,744,1949,1971]" pageId="1" pageNumber="167" refString="Gillies, M. T. (1990) A revision of the African species of Centroptilum Eaton (Baetidae, Ephemeroptera). Aquatic Insects, 12 (2), 97 - 128. http: // dx. doi. org / 10.1080 / 01650429009361395" type="journal article" year="1990">Gillies 1990</bibRefCitation>
: 99
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, 100, 119. Genus 
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<emphasis id="FDC181224F29FF8D21FF0FC1FE86ED99" box="[222,361,1980,2002]" italics="true" pageId="1" pageNumber="136">Cheleocloeon</emphasis>
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.
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Hierarchical name: 
<taxonomicName id="08B526B34F2AFF8E207F08E5FE1EEAE5" ID-CoL="3MMB" box="[350,497,152,174]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">Cheleocloeon</taxonomicName>
/g1.
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</subSubSection>
<subSubSection id="87AF0EBB4F2AFF8E21B608CAFDA3E9D2" pageId="2" pageNumber="137" type="materials_examined">
<paragraph id="CF0A5D304F2AFF8E21B608CAFC67EA87" blockId="2.[151,904,152,204]" box="[151,904,182,204]" pageId="2" pageNumber="137">
<typeStatus id="100EE3924F2AFF8E21B608CAFF26EA87" box="[151,201,183,204]" pageId="2" pageNumber="137">Type</typeStatus>
species: 
<taxonomicName id="08B526B34F2AFF8E200708CBFD88EA87" authority="Wuillot" authorityName="Wuillot" box="[294,615,182,204]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2AFF8E200708CBFDFBEA87" box="[294,532,182,204]" italics="true" pageId="2" pageNumber="137">Cheleocloeon yolandae</emphasis>
Wuillot
</taxonomicName>
(in 
<bibRefCitation id="AB2420C14F2AFF8E23B108CBFC6BEA87" author="Wuillot" box="[656,900,182,204]" pageId="2" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot &amp; Gillies) 1993</bibRefCitation>
.
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21B60884FCF1EB35" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21B60884FE68EB59" bold="true" box="[151,391,249,274]" pageId="2" pageNumber="137">Systematic position.</emphasis>
Following the formal ranks of classification, the genus 
<taxonomicName id="08B526B34F2AFF8E25260884FB4EEB5A" box="[1031,1185,249,273]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E25260884FB4EEB5A" box="[1031,1185,249,273]" italics="true" pageId="2" pageNumber="137">Cheleocloeon</emphasis>
</taxonomicName>
belongs to the family 
<taxonomicName id="08B526B34F2AFF8E21B60960FE6AEB7D" authority="Leach 1815" authorityName="Leach" authorityYear="1815" box="[151,389,285,310]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="family">Baetidae Leach 1815</taxonomicName>
. In the rank-free classification developed by 
<bibRefCitation id="AB2420C14F2AFF8E22590960FBE1EB7D" author="Kluge" box="[888,1038,285,310]" pageId="2" pageNumber="167" refString="Kluge, N. J. (2004) The phylogenetic system of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, 456 pp." type="book" year="2004">Kluge (2004)</bibRefCitation>
the taxon 
<taxonomicName id="08B526B34F2AFF8E25A00960FACEEB7D" box="[1153,1313,285,310]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">Cheleocloeon</taxonomicName>
/g1 belongs to the following subsequently subordinated taxa: Tetramerotarsata 
<bibRefCitation id="AB2420C14F2AFF8E2286093CFBDFEB12" author="Kluge" box="[935,1072,321,346]" pageId="2" pageNumber="167" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
&gt; Liberevenata 
<bibRefCitation id="AB2420C14F2AFF8E25D6093CFA6FEB12" author="Kluge" box="[1271,1408,321,346]" pageId="2" pageNumber="167" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
&gt; Turbanoculata 
<bibRefCitation id="AB2420C14F2AFF8E201F0918FE2DEB35" author="Kluge" box="[318,450,357,382]" pageId="2" pageNumber="167" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
&gt; Anteropatellata 
<bibRefCitation id="AB2420C14F2AFF8E23B20918FCF5EB35" author="Kluge" box="[659,794,357,382]" pageId="2" pageNumber="167" refString="Kluge, N. J. (1997) Classification and phylogeny of the Baetidae (Ephemeroptera) with description of the new species from the Upper Cretaceous resins of Taimyr. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera. Biology-Ecology- Systematics. Mauron + Tinguely &amp; Lacht SA, Fribourg, pp. 527 - 535." type="book chapter" year="1997">Kluge 1997</bibRefCitation>
.
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E609F7FAEBEB8D" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
In all species of 
<taxonomicName id="08B526B34F2AFF8E20A409F4FDF0EBEA" box="[389,543,393,417]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E20A409F4FDF0EBEA" box="[389,543,393,417]" italics="true" pageId="2" pageNumber="137">Cheleocloeon</emphasis>
</taxonomicName>
, the patella-tibial suture is equally developed in the fore and other legs of the larva, and female subimago and imago, that is characteristic for Anteropatellata (
<figureCitation id="578E41B54F2AFF8E250009D0FB17EB8D" box="[1057,1272,429,454]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="2" pageNumber="137">Figs 19–22, 25, 26</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E609ACFDFDE919" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
The taxon Anteropatellata is divided into the following taxa: (1) Baetovectata 
<bibRefCitation id="AB2420C14F2AFF8E256D09ACFA88EBA2" author="Kluge" box="[1100,1383,465,490]" pageId="2" pageNumber="167" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
(the largest baetid taxon with world-wide distribution); (2) 
<taxonomicName id="08B526B34F2AFF8E22350988FC85E845" box="[788,874,501,526]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1, or tribe Cloeonini Newman 1853 (the large taxon distributed in Holarctic, Afrotropical, Oriental and Australian Regions); (3) 
<taxonomicName id="08B526B34F2AFF8E25670A64FB0CE879" box="[1094,1251,537,562]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">Centroptilum</taxonomicName>
/g1, or the genus 
<taxonomicName id="08B526B34F2AFF8E21B60A40FE5BE81D" authority="Eaton 1869" authorityName="Eaton" authorityYear="1869" box="[151,436,573,598]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E21B60A40FEC4E81E" box="[151,299,573,597]" italics="true" pageId="2" pageNumber="137">Centroptilum</emphasis>
Eaton 1869
</taxonomicName>
[with a singe Holarctic species 
<taxonomicName id="08B526B34F2AFF8E22000A40FB22E81D" authority="Muller 1776" authorityName="Muller" authorityYear="1776" box="[801,1229,573,598]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="luteolum">
<emphasis id="FDC181224F2AFF8E22000A40FBCFE81E" box="[801,1056,573,597]" italics="true" pageId="2" pageNumber="137">Centroptilum luteolum</emphasis>
(Müller 1776)
</taxonomicName>
]; (4) 
<taxonomicName id="08B526B34F2AFF8E24280A43FA97E81D" box="[1289,1400,574,598]" class="Insecta" family="Baetidae" genus="Baetopus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">Baetopus</taxonomicName>
/g1, or the genus 
<taxonomicName id="08B526B34F2AFF8E20180A1FFD67E831" authority="Keffermuller 1960" authorityName="Keffermuller" authorityYear="1960" box="[313,648,609,634]" class="Insecta" family="Baetidae" genus="Baetopus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E20180A1FFE4EE832" box="[313,417,610,633]" italics="true" pageId="2" pageNumber="137">Baetopus</emphasis>
Keffermüller 1960
</taxonomicName>
, s.l. (with several species in Holarctic and Oriental Regions); (5) 
<taxonomicName id="08B526B34F2AFF8E21B60AF8FED9E8D5" box="[151,310,645,670]" class="Insecta" family="Baetidae" genus="Afrobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">Afrobaetodes</taxonomicName>
/g1, or the genus 
<taxonomicName id="08B526B34F2AFF8E20D20AF8FCD1E8D5" authority="Demoulin 1970" authorityName="Demoulin" authorityYear="1970" box="[499,830,645,670]" class="Insecta" family="Baetidae" genus="Afrobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E20D20AF8FD68E8D6" box="[499,647,645,669]" italics="true" pageId="2" pageNumber="137">Afrobaetodes</emphasis>
Demoulin 1970
</taxonomicName>
(with several species in Afrotropical Region) and (6) 
<taxonomicName id="08B526B34F2AFF8E21B60AD4FED6E889" box="[151,313,681,706]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">Cheleocloeon</taxonomicName>
/g1, or the genus 
<taxonomicName id="08B526B34F2AFF8E20D40AD4FC74E889" authority="Wuillot &amp; Gillies 1993" authorityName="Wuillot &amp; Gillies" authorityYear="1993" box="[501,923,681,706]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E20D40AD4FD60E88A" box="[501,655,681,705]" italics="true" pageId="2" pageNumber="137">Cheleocloeon</emphasis>
<bibRefCitation id="AB2420C14F2AFF8E23B60AD4FC74E889" author="Wuillot" box="[663,923,681,706]" pageId="2" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot &amp; Gillies 1993</bibRefCitation>
</taxonomicName>
(with several species in Afrotropical Region, northern Africa and Arabia). The genus 
<taxonomicName id="08B526B34F2AFF8E23580AB0FC4DE8AD" authority="Kluge 2012" authorityName="Kluge" authorityYear="2012" box="[633,930,717,742]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E23580AB0FCE2E8AE" box="[633,781,717,741]" italics="true" pageId="2" pageNumber="137">Anafroptilum</emphasis>
Kluge 2012
</taxonomicName>
(with several species in the Holarctic and Oriental Regions) has questionable systematic position, because its patella-tibial suture is always absent on fore legs, as in true Protopatellata 
<bibRefCitation id="AB2420C14F2AFF8E20FE0B68FD1BE965" author="Kluge" box="[479,756,789,814]" pageId="2" pageNumber="167" refString="Kluge, N. J. &amp; Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge &amp; Novikova 2011</bibRefCitation>
, but other characters agree with that of the taxon 
<taxonomicName id="08B526B34F2AFF8E24360B68FA82E965" box="[1303,1389,789,814]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1, which belongs to Anteropatellata.
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E60B23FDA3E9D2" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
In 
<taxonomicName id="08B526B34F2AFF8E21C40B20FE90E93E" box="[229,383,861,885]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E21C40B20FE90E93E" box="[229,383,861,885]" italics="true" pageId="2" pageNumber="137">Cheleocloeon</emphasis>
</taxonomicName>
, as well as in all other Turbanoculata except for Baetovectata, the fore wing has no more than one marginal intercalary in each space.
</paragraph>
</subSubSection>
<subSubSection id="87AF0EBB4F2AFF8521E60BD8FE35ED31" lastPageId="9" lastPageNumber="144" pageId="2" pageNumber="137" type="description">
<paragraph id="CF0A5D304F2AFF8E21E60BD8FDEDEE61" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60BD8FE2CE9F5" bold="true" box="[199,451,933,958]" pageId="2" pageNumber="137">Annotated diagnosis.</emphasis>
Male imago of 
<taxonomicName id="08B526B34F2AFF8E23580BD8FCFCE9F6" box="[633,787,933,957]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E23580BD8FCFCE9F6" box="[633,787,933,957]" italics="true" pageId="2" pageNumber="137">Cheleocloeon</emphasis>
</taxonomicName>
has a peculiar structure of the penis, which represents an autapomorphy of this taxon [see (23)]; other characters are either plesiomorphies, or characters which could repeatedly appear in some non-related taxa, so that the larval stage of 
<taxonomicName id="08B526B34F2AFF8E22FC0B90FB98EE4E" box="[989,1143,1005,1029]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E22FC0B90FB98EE4E" box="[989,1143,1005,1029]" italics="true" pageId="2" pageNumber="137">Cheleocloeon</emphasis>
</taxonomicName>
is characterized only by combination of these characters.
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E60C48FEB6EE39" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60C48FE65EE05" bold="true" box="[199,394,1077,1102]" pageId="2" pageNumber="137">(1) Larval frons.</emphasis>
Bases of antennae are widely separated, frons between them is flat (
<figureCitation id="578E41B54F2AFF8E25A10C48FB2CEE05" box="[1152,1219,1077,1102]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="2" pageNumber="137">Fig. 9</figureCitation>
) (unlike carinate in some other taxa).
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E60C00FF5AEEF1" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60C00FE1DEEDD" bold="true" box="[199,498,1149,1174]" pageId="2" pageNumber="137">(2) Larval frontal suture.</emphasis>
Frontal suture is nearly semicircular, with angle blunt and arms evenly arched (
<figureCitation id="578E41B54F2AFF8E244E0C00FF48EEF1" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="2" pageNumber="137">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E60CB8FA98EF26" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60CB8FEB6EE95" bold="true" box="[199,345,1221,1246]" pageId="2" pageNumber="137">(3) Labrum.</emphasis>
Labrum has an usual form, outer surface with submedian pair of long setae and a few (often two) antero-lateral setae on each side (
<figureCitation id="578E41B54F2AFF8E23340C94FDB5EF49" box="[533,602,1257,1282]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="2" pageNumber="137">Fig. 1</figureCitation>
; 
<bibRefCitation id="AB2420C14F2AFF8E23480C94FCB5EF49" author="Gattolliat" box="[617,858,1257,1282]" pageId="2" pageNumber="167" refString="Gattolliat, J. - L., Barber-James, H. M. &amp; Monaghan, M. T. (2009) New species and generic delimitation of the Afrotropical genera Bugilliesia Lugo-Ortiz &amp; McCafferty, 1996; Cheleocloeon Wuillot &amp; Gillies, 1993 and Delouardus Lugo-Ortiz &amp; McCafferty, 1999 (Ephemeroptera: Baetidae). Aquatic Insects, 31 (3), 167 - 186. http: // dx. doi. org / 10.1080 / 01650420902782987" type="journal article" year="2009">
Gattolliat 
<emphasis id="FDC181224F2AFF8E23FD0C97FCF9EF4A" box="[732,790,1257,1281]" italics="true" pageId="2" pageNumber="137">et al.</emphasis>
2009
</bibRefCitation>
: 
<figureCitation id="578E41B54F2AFF8E22470C94FC24EF49" box="[870,971,1257,1282]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="2" pageNumber="137">Figs 1–3</figureCitation>
, 
<figureCitation id="578E41B54F2AFF8E22F80C94FC17EF49" box="[985,1016,1257,1282]" captionStart="FIGURES 30 – 57" captionStartId="8.[151,264,1554,1577]" captionTargetBox="[153,1432,201,1529]" captionTargetId="figure@8.[151,1436,193,1533]" captionTargetPageId="8" captionText="FIGURES 30 – 57. Tergalii I – VII: 30 – 36, Cheleocloeon clavifolium sp. n.; 37 – 43, Ch. lancetofolium sp. n.; 44 – 50, Ch. truncifolium sp. n. (male, holotype); 51 – 57, Ch. truncifolium sp. n. (female)." httpUri="https://zenodo.org/record/270600/files/figure.png" pageId="2" pageNumber="137">35</figureCitation>
); in some individuals these setae are absent. Smaller setae are irregularly scattered of outer surface of labrum; some of them can be subequal to the submedian and antero-lateral setae. Therefore the structure of the labrum is not a good diagnostic character for 
<taxonomicName id="08B526B34F2AFF8E21B60D28FEDEEF26" box="[151,305,1365,1389]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E21B60D28FEDEEF26" box="[151,305,1365,1389]" italics="true" pageId="2" pageNumber="137">Cheleocloeon</emphasis>
</taxonomicName>
, but it allows it to be distinguished from other taxa, whose antero-lateral setae form regular rows.
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E60D04FDEAECFA" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60D04FE82EFD9" bold="true" box="[199,365,1401,1426]" pageId="2" pageNumber="137">(4) Mandibles</emphasis>
(
<figureCitation id="578E41B54F2AFF8E205D0D04FDD8EFD9" box="[380,567,1401,1426]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="2" pageNumber="137">Figs 7, 8, 16–18</figureCitation>
, 
<figureCitation id="578E41B54F2AFF8E23620D04FD62EFD9" box="[579,653,1401,1426]" captionStart="FIGURES 58 – 63" captionStartId="11.[151,264,1736,1759]" captionTargetBox="[151,1434,176,1715]" captionTargetId="figure@11.[151,1435,159,1715]" captionTargetPageId="11" captionText="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." httpUri="https://zenodo.org/record/270601/files/figure.png" pageId="2" pageNumber="137">58, 59</figureCitation>
; 
<bibRefCitation id="AB2420C14F2AFF8E23B80D04FC4EEFD9" author="Wuillot" box="[665,929,1401,1426]" pageId="2" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot &amp; Gillies 1993</bibRefCitation>
: 
<figureCitation id="578E41B54F2AFF8E228D0D04FBFFEFD9" box="[940,1040,1401,1426]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="2" pageNumber="137">Figs 4, 5</figureCitation>
, 
<figureCitation id="578E41B54F2AFF8E253C0D07FB88EFD9" box="[1053,1127,1402,1426]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="2" pageNumber="137">13, 14</figureCitation>
; 
<bibRefCitation id="AB2420C14F2AFF8E25520D04FA8EEFD9" author="Gattolliat" box="[1139,1377,1401,1426]" pageId="2" pageNumber="167" refString="Gattolliat, J. - L., Barber-James, H. M. &amp; Monaghan, M. T. (2009) New species and generic delimitation of the Afrotropical genera Bugilliesia Lugo-Ortiz &amp; McCafferty, 1996; Cheleocloeon Wuillot &amp; Gillies, 1993 and Delouardus Lugo-Ortiz &amp; McCafferty, 1999 (Ephemeroptera: Baetidae). Aquatic Insects, 31 (3), 167 - 186. http: // dx. doi. org / 10.1080 / 01650420902782987" type="journal article" year="2009">
Gattolliat 
<emphasis id="FDC181224F2AFF8E25C50D07FAF2EFDA" box="[1252,1309,1401,1425]" italics="true" pageId="2" pageNumber="137">et al.</emphasis>
2009
</bibRefCitation>
: 
<figureCitation id="578E41B54F2AFF8E244D0D04FECFEFFD" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="2" pageNumber="137">Figs 7–9, 13–15</figureCitation>
). On both mandibles, incisor and kinetodontium are moderately fused together. Left incisor is terminated by 3 prominent denticles with distal denticle the longest, without small ventral distal denticle. Left kinetodontium has 1st (most distal) denticle the longest, 2nd shorter than 3rd denticle. Left prostheca is terminated by 3–4 stout short denticles and 2–3 slender longer denticles (individual variability). Right incisor is terminated by 4 prominent denticles, with or without a small ventral distal denticle. Right kinetodontium is terminated by 4 denticles gradually getting shorter from most distal to most proximal. Right prostheca is stick-like, directed medially, apically either narrowed, or widened (species-specific character). Margin between prostheca and mola bears dense setae-like processes.
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E60EC0FC8BEDC1" blockId="2.[151,1437,249,2038]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60EC0FEB4EC9D" bold="true" box="[199,347,1725,1750]" pageId="2" pageNumber="137">(5) Maxillae</emphasis>
(
<figureCitation id="578E41B54F2AFF8E204F0EC0FE33EC9D" box="[366,476,1725,1750]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="2" pageNumber="137">Figs 2, 3</figureCitation>
). Maxilla of the « 
<taxonomicName id="08B526B34F2AFF8E23970EC0FCE8EC9E" box="[694,775,1725,1749]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E23970EC0FCE8EC9E" box="[694,775,1725,1749]" italics="true" pageId="2" pageNumber="137">Cloeon</emphasis>
</taxonomicName>
- 
<typeStatus id="100EE3924F2AFF8E222E0EC3FCABEC9D" box="[783,836,1726,1750]" pageId="2" pageNumber="137">type</typeStatus>
», i.e. with long slender canines and dentisetae; 1st dentiseta is simple, 2nd and 3rd dentisetae are bifid; 1st and 2nd dentisetae are pressed to canines, 3rd dentiseta is separated. Medio-dorsal setal row, continued proximal of dentisetae, consists of 4–6 setae, which are either all simple, or some of them are bifid (individual variability); in some individuals the first seta of this row, being adjacent to the 3rd dentiseta, is bifid, thicker than others and resembles dentiseta, so that maxilla appears to have 4 dentisetae (while only 3 true dentisetae are present in 
<taxonomicName id="08B526B34F2AFF8E23D00F0CFCB7EDC1" box="[753,856,1905,1930]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
).
</paragraph>
<paragraph id="CF0A5D304F2AFF8E21E60FE8FB22EDE5" blockId="2.[151,1437,249,2038]" box="[199,1229,1941,1966]" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60FE8FE47EDE5" bold="true" box="[199,424,1941,1966]" pageId="2" pageNumber="137">(6) Maxillary palp.</emphasis>
Maxillary palp is 2-segmented, with 2nd segment often arched (
<figureCitation id="578E41B54F2AFF8E255B0FE8FB2FEDE5" box="[1146,1216,1941,1966]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="2" pageNumber="137">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2AFF8F21E60FC4FC99EB56" blockId="2.[151,1437,249,2038]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="138" pageId="2" pageNumber="137">
<emphasis id="FDC181224F2AFF8E21E60FC4FEBFED99" bold="true" box="[199,336,1977,2002]" pageId="2" pageNumber="137">(7) Labium</emphasis>
(
<figureCitation id="578E41B54F2AFF8E20430FC4FE24ED99" box="[354,459,1977,2002]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="2" pageNumber="137">Figs 4–6</figureCitation>
). Labium of the « 
<taxonomicName id="08B526B34F2AFF8E23BF0FC4FD00ED9A" box="[670,751,1977,2001]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2AFF8E23BF0FC4FD00ED9A" box="[670,751,1977,2001]" italics="true" pageId="2" pageNumber="137">Cloeon</emphasis>
</taxonomicName>
- 
<typeStatus id="100EE3924F2AFF8E23D60FC7FCC3ED99" box="[759,812,1978,2002]" pageId="2" pageNumber="137">type</typeStatus>
», i.e., glossa and paraglossa have subequal width and length, paraglossa is crescent-shaped. Glossa has the following setal rows: median row (regular row of spine-like setae along median margin); dorso-lateral row (regular row along lateral margin, with setal bases visible in dorsal view). Ventral side of glossa bears setae directed ventrally; they are located irregularly in the proximal part of the glossa and form a ventro-median row (sparse row parallel to median margin). The median, the dorso-lateral and the ventro-medial rows stretch along most of the length of the glossa.
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E6095AFD9CEB9B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">Paraglossa has following setation: latero-apical setae (longest ones) form a regular sparse row along lateral and apical margin and irregularly located on the dorsal side; ventro-median row (regular row of long slender straight setae attached on ventral side and directed mainly medially); dorso-median row (regular row of thicker setae attached on dorsal side and directed medially-distally). The ventro-median and the dorso-median rows are stretched along most of the length of the paraglossa.</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E609A1FD77E853" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
Similar labial structure occurs in some other baetid taxa, both among Protopatellata and Anteropatellata (e.g., most Cloeonini, 
<taxonomicName id="08B526B34F2BFF8F20700A7DFE0AE853" box="[337,485,512,536]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F20700A7DFE0AE853" box="[337,485,512,536]" italics="true" pageId="3" pageNumber="138">Centroptilum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F2BFF8F20D20A7DFD68E853" box="[499,647,512,536]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F20D20A7DFD68E853" box="[499,647,512,536]" italics="true" pageId="3" pageNumber="138">Anafroptilum</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60A59FAD8E97C" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
<emphasis id="FDC181224F2BFF8F21E60A59FE6FE876" bold="true" box="[199,384,548,573]" pageId="3" pageNumber="138">(8) Labial palp</emphasis>
(
<figureCitation id="578E41B54F2BFF8F20B50A59FDEDE876" box="[404,514,548,573]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="3" pageNumber="138">Figs 4, 5</figureCitation>
). Shape of the labial palp is characteristic for 
<taxonomicName id="08B526B34F2BFF8F250A0A59FB2AE877" box="[1067,1221,548,572]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F250A0A59FB2AE877" box="[1067,1221,548,572]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
: 2nd segment has prominent disto-median projection with distal margin convex. 2nd segment retains muscle running from its base to base of 3rd segment (shown by dotted stripe on figures by 
<bibRefCitation id="AB2420C14F2BFF8F221E0A11FB1DE8CE" author="Wuillot" box="[831,1266,620,645]" pageId="3" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">
Wuillot and Gillies (1993: 
<figureCitation id="578E41B54F2BFF8F25520A11FB52E8CE" box="[1139,1213,620,645]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="3" pageNumber="138">Figs 6</figureCitation>
, 
<figureCitation id="578E41B54F2BFF8F25EA0A11FB05E8CE" box="[1227,1258,620,645]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="3" pageNumber="138">15</figureCitation>
)
</bibRefCitation>
. Two forms of labial palp are distinguishable among 
<taxonomicName id="08B526B34F2BFF8F236A0AEDFD0AE8E3" box="[587,741,656,680]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F236A0AEDFD0AE8E3" box="[587,741,656,680]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
: (1) with apex of projection of 2nd segment pointed (in 
<taxonomicName id="08B526B34F2BFF8F24550AEDFF11E887" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2BFF8F24550AEDFF11E887" italics="true" pageId="3" pageNumber="138">Ch. yolandae</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F2BFF8F202A0AC9FE42E887" box="[267,429,692,716]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F2BFF8F202A0AC9FE42E887" box="[267,429,692,716]" italics="true" pageId="3" pageNumber="138">Ch. carinatum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F2BFF8F209B0AC9FDACE887" box="[442,579,692,716]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="littorale">
<emphasis id="FDC181224F2BFF8F209B0AC9FDACE887" box="[442,579,692,716]" italics="true" pageId="3" pageNumber="138">Ch. littorale</emphasis>
</taxonomicName>
and 
<emphasis id="FDC181224F2BFF8F23580AC9FD75E887" box="[633,666,692,716]" italics="true" pageId="3" pageNumber="138">Ch</emphasis>
. sp. 3) (
<bibRefCitation id="AB2420C14F2BFF8F23D20AC9FB9BE886" author="Wuillot" box="[755,1140,692,717]" pageId="3" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">
Wuillot &amp; Gillies 1993: 
<figureCitation id="578E41B54F2BFF8F25240AC9FBA3E886" box="[1029,1100,692,717]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="3" pageNumber="138">Figs 6</figureCitation>
, 
<figureCitation id="578E41B54F2BFF8F25760AC9FB9BE886" box="[1111,1140,692,717]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="3" pageNumber="138">15</figureCitation>
</bibRefCitation>
) and (2) rounded (in other species— 
<taxonomicName id="08B526B34F2BFF8F20250AA5FE5AE8BB" box="[260,437,728,752]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2BFF8F20250AA5FE5AE8BB" box="[260,437,728,752]" italics="true" pageId="3" pageNumber="138">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2BFF8F20E00AAAFDE9E8BB" box="[449,518,727,752]" pageId="3" pageNumber="138" rank="species">
<emphasis id="FDC181224F2BFF8F20E00AAAFDE9E8BB" bold="true" box="[449,518,727,752]" pageId="3" pageNumber="138">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F2BFF8F23390AA5FD09E8BB" box="[536,742,728,752]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F2BFF8F23390AA5FD09E8BB" box="[536,742,728,752]" italics="true" pageId="3" pageNumber="138">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2BFF8F23D30AAAFCD9E8BB" box="[754,822,727,752]" pageId="3" pageNumber="138" rank="species">
<emphasis id="FDC181224F2BFF8F23D30AAAFCD9E8BB" bold="true" box="[754,822,727,752]" pageId="3" pageNumber="138">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F2BFF8F22680AA5FBE8E8BB" box="[841,1031,728,752]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F2BFF8F22680AA5FBE8E8BB" box="[841,1031,728,752]" italics="true" pageId="3" pageNumber="138">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2BFF8F25320AAAFBB8E8BB" box="[1043,1111,727,752]" pageId="3" pageNumber="138" rank="species">
<emphasis id="FDC181224F2BFF8F25320AAAFBB8E8BB" bold="true" box="[1043,1111,727,752]" pageId="3" pageNumber="138">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F2BFF8F254B0AA5FA8EE8BB" box="[1130,1377,728,752]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="madagascariense">
<emphasis id="FDC181224F2BFF8F254B0AA5FA8EE8BB" box="[1130,1377,728,752]" italics="true" pageId="3" pageNumber="138">Ch. madagascariense</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F2BFF8F24550AA5FEC3E95F" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="dimorphicum">
<emphasis id="FDC181224F2BFF8F24550AA5FEC3E95F" italics="true" pageId="3" pageNumber="138">Ch. dimorphicum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F2BFF8F201D0A81FE29E95F" box="[316,454,764,788]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F2BFF8F201D0A81FE29E95F" box="[316,454,764,788]" italics="true" pageId="3" pageNumber="138">Ch. excisum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F2BFF8F20F40A81FDB7E95F" box="[469,600,764,788]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F2BFF8F20F40A81FDB7E95F" box="[469,600,764,788]" italics="true" pageId="3" pageNumber="138">Ch. soldani</emphasis>
</taxonomicName>
and 
<emphasis id="FDC181224F2BFF8F23B00A81FD56E95F" box="[657,697,764,788]" italics="true" pageId="3" pageNumber="138">Ch.</emphasis>
sp. 2) (
<figureCitation id="578E41B54F2BFF8F22300A81FC96E95E" box="[785,889,764,789]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="3" pageNumber="138">Figs 4, 5</figureCitation>
). Some non-related taxa also have disto-median projections of 2nd segment of maxillary palp, but their shape is often different from that of 
<taxonomicName id="08B526B34F2BFF8F25B70B5DFADFE973" box="[1174,1328,800,824]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F25B70B5DFADFE973" box="[1174,1328,800,824]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60B39FC76E983" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
<emphasis id="FDC181224F2BFF8F21E60B39FE37E916" bold="true" box="[199,472,836,861]" pageId="3" pageNumber="138">(9) Postsubalar sclerite</emphasis>
(
<figureCitation id="578E41B54F2BFF8F20C70B39FD88E917" box="[486,615,836,861]" captionStart="FIGURES 87 – 94" captionStartId="20.[151,264,1528,1551]" captionTargetBox="[167,1412,193,1510]" captionTargetId="figure@20.[167,1413,193,1511]" captionTargetPageId="20" captionText="FIGURES 87 – 94. Details of exuviae of subimago. 87 – 89, Cheleocloeon clavifolium sp. n.: 87 – 88, left half of mesonotum of different specimens; 89, pterothorax sterna and pleura. 90 – 92, Ch. truncifolium sp. n.: 90, left half of mesonotum (holotype); 91, abdominal segments IV – V; 92, left pleuron. 93, Ch. yolandae, left half of mesonotum; 94, Cheleocloeon sp. 1, abdominal segments III – V. Abbreviations: lpnc, lateropostnotal crest; psa, postsubalar sclerite; sa, subalar sclerite." httpUri="https://zenodo.org/record/270605/files/figure.png" pageId="3" pageNumber="138">Figs 89, 92</figureCitation>
). Postsubalare (which is present in imago and subimago and in both these stages has contrasting brown cuticular coloration) has posterior-dorsal corner stretched to a rather wide process with shallowly convex dorsal margin. This is different from Cloeonini and some other taxa, whose posterior-dorsal postsubalare process is small or narrow, with concave dorsal margin.
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60BA9FED8EE7E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
<emphasis id="FDC181224F2BFF8F21E60BA9FE32E9A6" bold="true" box="[199,477,980,1005]" pageId="3" pageNumber="138">(10) Larval leg setation</emphasis>
(
<figureCitation id="578E41B54F2BFF8F20CC0BA9FD73E9A6" box="[493,668,980,1005]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Figs 21, 25, 26</figureCitation>
). Outer side of femur bears sparse row of small stout setae, apically terminated by 2 small stout setae located close together. The same in many other baetid taxa; possible plesiomorphy.
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60C42FC46EE36" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
While setation of legs is rather uniform among 
<taxonomicName id="08B526B34F2BFF8F22200C3DFC74EE13" box="[769,923,1088,1112]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F22200C3DFC74EE13" box="[769,923,1088,1112]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
, leg shape varies from slender, with femur parallel-sided (
<figureCitation id="578E41B54F2BFF8F201F0C19FE7FEE37" box="[318,400,1124,1149]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Fig. 21</figureCitation>
) to rather stout, with femur thickened (
<figureCitation id="578E41B54F2BFF8F22690C19FC73EE36" box="[840,924,1124,1149]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60CFAFD4CEEA3" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
<emphasis id="FDC181224F2BFF8F21E60CFAFE03EEEB" bold="true" box="[199,492,1159,1184]" pageId="3" pageNumber="138">(11) Patella-tibial suture</emphasis>
(
<figureCitation id="578E41B54F2BFF8F20DD0CFAFD36EEEB" box="[508,729,1159,1184]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Figs 19–22, 25, 26</figureCitation>
). This suture is equally developed on all legs of larva of both sexes and on all legs of female subimago and imago, being absent only on fore legs of male subimago and imago (i.e., the typical condition for Anteropatellata).
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60C89FD4EEC43" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
This character is important here, because 
<taxonomicName id="08B526B34F2BFF8F23880C89FCACEF47" box="[681,835,1268,1292]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F23880C89FCACEF47" box="[681,835,1268,1292]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
was confused with both some Anteropatellata (e.g., 
<taxonomicName id="08B526B34F2BFF8F21B60D65FEC4EF7B" box="[151,299,1304,1328]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F21B60D65FEC4EF7B" box="[151,299,1304,1328]" italics="true" pageId="3" pageNumber="138">Centroptilum</emphasis>
</taxonomicName>
) and with some Protopatellata (e.g., 
<taxonomicName id="08B526B34F2BFF8F23EB0D65FCADEF7B" box="[714,834,1304,1328]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F23EB0D65FCADEF7B" box="[714,834,1304,1328]" italics="true" pageId="3" pageNumber="138">Afroptilum</emphasis>
</taxonomicName>
and 
<taxonomicName id="08B526B34F2BFF8F225B0D65FC1CEF7B" box="[890,1011,1304,1328]" class="Insecta" family="Baetidae" genus="Bugilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F225B0D65FC1CEF7B" box="[890,1011,1304,1328]" italics="true" pageId="3" pageNumber="138">Bugilliesia</emphasis>
</taxonomicName>
). Gillies wrote that 
<taxonomicName id="08B526B34F2BFF8F25F40D65FA80EF7B" box="[1237,1391,1304,1328]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F25F40D65FA80EF7B" box="[1237,1391,1304,1328]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
has patella-tibial suture on middle and hind leg only (
<bibRefCitation id="AB2420C14F2BFF8F23F90D41FC77EF1F" author="Gillies" box="[728,920,1340,1365]" pageId="3" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">Gillies 2001: 32</bibRefCitation>
); this conclusion was based on 
<taxonomicName id="08B526B34F2BFF8F24280D41FA7AEF1F" box="[1289,1429,1340,1364]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="sigiense">
<emphasis id="FDC181224F2BFF8F24280D41FA7AEF1F" box="[1289,1429,1340,1364]" italics="true" pageId="3" pageNumber="138">Ch. sigiense</emphasis>
</taxonomicName>
, which does not belong to 
<taxonomicName id="08B526B34F2BFF8F20ED0D1DFD89EF33" box="[460,614,1376,1400]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F20ED0D1DFD89EF33" box="[460,614,1376,1400]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
(see below). 
<bibRefCitation id="AB2420C14F2BFF8F22280D22FBFFEF33" box="[777,1040,1375,1400]" pageId="3" pageNumber="167" refString="Gattolliat, J. - L., Barber-James, H. M. &amp; Monaghan, M. T. (2009) New species and generic delimitation of the Afrotropical genera Bugilliesia Lugo-Ortiz &amp; McCafferty, 1996; Cheleocloeon Wuillot &amp; Gillies, 1993 and Delouardus Lugo-Ortiz &amp; McCafferty, 1999 (Ephemeroptera: Baetidae). Aquatic Insects, 31 (3), 167 - 186. http: // dx. doi. org / 10.1080 / 01650420902782987" type="journal article">
Gattolliat 
<emphasis id="FDC181224F2BFF8F225F0D1CFC55EF33" box="[894,954,1376,1400]" italics="true" pageId="3" pageNumber="138">et al.</emphasis>
(2009)
</bibRefCitation>
wrote in their larval diagnosis of 
<taxonomicName id="08B526B34F2BFF8F21B60DF9FEDEEFD7" box="[151,305,1412,1436]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F21B60DF9FEDEEFD7" box="[151,305,1412,1436]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
, that the patella-tibial suture is absent. Probably, this statement was caused by the fact that larval legs of 
<taxonomicName id="08B526B34F2BFF8F219C0DD5FEB8EF8B" box="[189,343,1448,1472]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2BFF8F219C0DD5FEB8EF8B" box="[189,343,1448,1472]" italics="true" pageId="3" pageNumber="138">Cheleocloeon</emphasis>
</taxonomicName>
have colorless translucent cuticle, so the patella-tibial suture can be poorly visible on a background of living tissues; it is well visible on empty larval exuviae. The anteropatellate condition of patellatibial suture is present in all species examined.
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60E69FA98ECF7" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
When 
<bibRefCitation id="AB2420C14F2BFF8F20350E69FDABEC67" author="Wuillot" box="[276,580,1556,1581]" pageId="3" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot and Gillies (1993)</bibRefCitation>
described 
<taxonomicName id="08B526B34F2BFF8F23E20E69FCB4EC67" box="[707,859,1556,1580]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2BFF8F23E20E69FCB4EC67" box="[707,859,1556,1580]" italics="true" pageId="3" pageNumber="138">Ch. yolandae</emphasis>
</taxonomicName>
and 
<taxonomicName id="08B526B34F2BFF8F22B90E69FBD3EC67" box="[920,1084,1556,1580]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F2BFF8F22B90E69FBD3EC67" box="[920,1084,1556,1580]" italics="true" pageId="3" pageNumber="138">Ch. carinatum</emphasis>
</taxonomicName>
, they did not know about this character and did not pay special attention to it. Distal border of dark macula, shown by dots on fore legs of both these species (
<bibRefCitation id="AB2420C14F2BFF8F20620E21FD04EC3F" author="Wuillot" box="[323,747,1628,1653]" pageId="3" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">
Wuillot &amp; Gillies 1993: 
<figureCitation id="578E41B54F2BFF8F234E0E21FD04EC3F" box="[623,747,1628,1653]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="3" pageNumber="138">Figs 8, 17</figureCitation>
</bibRefCitation>
) correspond to the patella-tibial suture. According to my examination of larval exuviae from 
<collectingCountry id="B7A21DA04F2BFF8F231F0E02FD78ECD3" box="[574,663,1663,1688]" name="Zambia" pageId="3" pageNumber="138">Zambia</collectingCountry>
, in both species patella-tibial suture is present on all legs; female subimago and imago of 
<taxonomicName id="08B526B34F2BFF8F20840ED9FDD4ECF7" box="[421,571,1700,1724]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2BFF8F20840ED9FDD4ECF7" box="[421,571,1700,1724]" italics="true" pageId="3" pageNumber="138">Ch. yolandae</emphasis>
</taxonomicName>
also have patella-tibial suture equally developed on all legs (
<figureCitation id="578E41B54F2BFF8F25C60ED9FA84ECF7" box="[1255,1387,1700,1725]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Figs 19–22</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60EB5FBC9ED4E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
In the descriptions of 
<taxonomicName id="08B526B34F2BFF8F20E00EB5FDA4EC94" box="[449,587,1736,1760]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F2BFF8F20E00EB5FDA4EC94" box="[449,587,1736,1760]" italics="true" pageId="3" pageNumber="138">Ch. excisum</emphasis>
</taxonomicName>
(
<bibRefCitation id="AB2420C14F2BFF8F237A0EBAFD14ECAB" author="Barnard" box="[603,763,1735,1760]" pageId="3" pageNumber="166" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. http: // dx. doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
; 
<bibRefCitation id="AB2420C14F2BFF8F22290EBAFB92ECAB" author="Lugo-Ortiz" box="[776,1149,1735,1760]" pageId="3" pageNumber="167" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1997 a) Contribution to the systematics of the genus Cheleocloeon (Ephemeroptera: Baetidae). Entomological News, 108 (4), 283 - 289" type="journal article" year="1997" yearSuffix="a">Lugo-Ortiz &amp; McCafferty 1997a</bibRefCitation>
) patella-tibial suture was not mentioned; actually it is equally developed on all legs of larva (as in 
<figureCitation id="578E41B54F2BFF8F22E70E91FBF6ED4E" box="[966,1049,1772,1797]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60F72FC35ED06" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
According to the original description of 
<taxonomicName id="08B526B34F2BFF8F23B10F6DFCFCED63" box="[656,787,1808,1832]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F2BFF8F23B10F6DFCFCED63" box="[656,787,1808,1832]" italics="true" pageId="3" pageNumber="138">Ch. soldani</emphasis>
</taxonomicName>
, “tibio-patellar suture absent” (Gattolliat &amp; Sartori 2008: 52). Actually this suture is equally developed on all legs of larva (
<figureCitation id="578E41B54F2BFF8F225B0F49FC21ED06" box="[890,974,1844,1869]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2BFF8F21E60F2AFEEFED96" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="138">
All three new species, 
<taxonomicName id="08B526B34F2BFF8F20E50F25FD9FED3B" box="[452,624,1880,1904]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2BFF8F20E50F25FD9FED3B" box="[452,624,1880,1904]" italics="true" pageId="3" pageNumber="138">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2BFF8F23560F2AFD59ED3B" box="[631,694,1879,1904]" pageId="3" pageNumber="138" rank="species">
<emphasis id="FDC181224F2BFF8F23560F2AFD59ED3B" bold="true" box="[631,694,1879,1904]" pageId="3" pageNumber="138">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F2BFF8F23E50F25FC63ED3B" box="[708,908,1880,1904]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F2BFF8F23E50F25FC63ED3B" box="[708,908,1880,1904]" italics="true" pageId="3" pageNumber="138">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2BFF8F22B20F2AFC3DED3B" box="[915,978,1879,1904]" pageId="3" pageNumber="138" rank="species">
<emphasis id="FDC181224F2BFF8F22B20F2AFC3DED3B" bold="true" box="[915,978,1879,1904]" pageId="3" pageNumber="138">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="08B526B34F2BFF8F25260F25FB2FED3B" box="[1031,1216,1880,1904]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F2BFF8F25260F25FB2FED3B" box="[1031,1216,1880,1904]" italics="true" pageId="3" pageNumber="138">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2BFF8F25E60F2AFAE9ED3B" box="[1223,1286,1879,1904]" pageId="3" pageNumber="138" rank="species">
<emphasis id="FDC181224F2BFF8F25E60F2AFAE9ED3B" bold="true" box="[1223,1286,1879,1904]" pageId="3" pageNumber="138">sp. n.</emphasis>
</taxonomicNameLabel>
have patellatibial suture equally developed on all legs of larva and equally developed on all legs of female subimago and imago. Larvae of unnamed species reported below, also have patella-tibial suture equally developed on all legs (
<figureCitation id="578E41B54F2BFF8F21BE0FB9FF1BED96" box="[159,244,1988,2013]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="3" pageNumber="138">Fig. 26</figureCitation>
).
</paragraph>
<caption id="9BCA0DB84F2CFF8821B60FEBFBA3EDA1" httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="4" pageNumber="139" targetBox="[168,1406,201,1903]" targetPageId="4">
<paragraph id="CF0A5D304F2CFF8821B60FEBFBA3EDA1" blockId="4.[151,1436,1942,2026]" pageId="4" pageNumber="139">
<emphasis id="FDC181224F2CFF8821B60FEBFED4EDE7" bold="true" box="[151,315,1942,1965]" pageId="4" pageNumber="139">FIGURES 1–6.</emphasis>
1–5, 
<taxonomicName id="08B526B34F2CFF8820550FEBFD97EDE7" box="[372,632,1942,1964]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2CFF8820550FEBFD97EDE7" box="[372,632,1942,1964]" italics="true" pageId="4" pageNumber="139">Cheleocloeon clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2CFF88235E0FEAFD57EDE7" box="[639,696,1943,1964]" pageId="4" pageNumber="139" rank="species">
<emphasis id="FDC181224F2CFF88235E0FEAFD57EDE7" bold="true" box="[639,696,1943,1964]" pageId="4" pageNumber="139">sp. n.</emphasis>
</taxonomicNameLabel>
: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, 
<taxonomicName id="08B526B34F2CFF8824590FC8FECDEDA1" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F2CFF8824590FC8FECDEDA1" italics="true" pageId="4" pageNumber="139">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2CFF8820090FA9FE8EEDA2" box="[296,353,2004,2025]" pageId="4" pageNumber="139" rank="species">
<emphasis id="FDC181224F2CFF8820090FA9FE8EEDA2" bold="true" box="[296,353,2004,2025]" pageId="4" pageNumber="139">sp. n.</emphasis>
</taxonomicNameLabel>
(holotype): glossa, ventral view. Abbreviations: ds1, ds2, ds3, dentisetae.
</paragraph>
</caption>
<caption id="9BCA0DB84F2DFF8921B60DE1FE0FEC21" httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="5" pageNumber="140" targetBox="[226,1346,181,1410]" targetPageId="5">
<paragraph id="CF0A5D304F2DFF8921B60DE1FE0FEC21" blockId="5.[151,1436,1436,1642]" pageId="5" pageNumber="140">
<emphasis id="FDC181224F2DFF8921B60DE1FEA8EFFA" bold="true" box="[151,327,1436,1459]" pageId="5" pageNumber="140">FIGURES 7–18.</emphasis>
7–15, 
<taxonomicName id="08B526B34F2DFF8920AD0DE1FD60EFF9" box="[396,655,1436,1458]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2DFF8920AD0DE1FD60EFF9" box="[396,655,1436,1458]" italics="true" pageId="5" pageNumber="140">Cheleocloeon clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2DFF8923B40DE0FD21EFF9" box="[661,718,1437,1458]" pageId="5" pageNumber="140" rank="species">
<emphasis id="FDC181224F2DFF8923B40DE0FD21EFF9" bold="true" box="[661,718,1437,1458]" pageId="5" pageNumber="140">sp. n.</emphasis>
</taxonomicNameLabel>
: 7–8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10–11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12–13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, 
<taxonomicName id="08B526B34F2DFF8924590D85FECDEC66" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F2DFF8924590D85FECDEC66" italics="true" pageId="5" pageNumber="140">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2DFF89200C0E6AFE84EC67" box="[301,363,1559,1580]" pageId="5" pageNumber="140" rank="species">
<emphasis id="FDC181224F2DFF89200C0E6AFE84EC67" bold="true" box="[301,363,1559,1580]" pageId="5" pageNumber="140">sp. n.</emphasis>
</taxonomicNameLabel>
: incisor, kinetodontium and prostheca of right mandible; 17, 
<taxonomicName id="08B526B34F2DFF8925240E6AFB5EEC66" box="[1029,1201,1559,1581]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F2DFF8925240E6AFB5EEC66" box="[1029,1201,1559,1581]" italics="true" pageId="5" pageNumber="140">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2DFF89259D0E6AFB15EC67" box="[1212,1274,1559,1580]" pageId="5" pageNumber="140" rank="species">
<emphasis id="FDC181224F2DFF89259D0E6AFB15EC67" bold="true" box="[1212,1274,1559,1580]" pageId="5" pageNumber="140">sp. n.</emphasis>
</taxonomicNameLabel>
, the same; 18, 
<taxonomicName id="08B526B34F2DFF8921B60E48FECDEC00" box="[151,290,1589,1611]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2DFF8921B60E48FECDEC00" box="[151,290,1589,1611]" italics="true" pageId="5" pageNumber="140">Cheleocloeon</emphasis>
</taxonomicName>
sp. 3, the same. Abbreviations: in1, in2, in3, in4, denticles of incisor; in-V, ventral denticle of incisor; kd1, kd2, kd3, denticles of kinetodontium.
</paragraph>
</caption>
<paragraph id="CF0A5D304F2DFF8921E60EECFC38ED5D" blockId="5.[151,1436,1681,2030]" pageId="5" pageNumber="140">
<emphasis id="FDC181224F2DFF8921E60EECFD93ECE1" bold="true" box="[199,636,1681,1706]" pageId="5" pageNumber="140">(12) Imaginal and subimaginal tarsi.</emphasis>
Tarsus of middle and hind leg is relatively long, about 0.7–0.9 of tibia length; proximal tarsal segment (result of complete fusion of primary 1st+2nd tarsomeres) is much longer than others, about 0.4–0.5 of total tarsal length. All examined species have 2 apical spines on middle and hind legs of both sexes and on fore leg of female (on 2nd and 3rd primary tarsomeres).
</paragraph>
<paragraph id="CF0A5D304F2DFF8921E60F5CFCD8EDA5" blockId="5.[151,1436,1681,2030]" pageId="5" pageNumber="140">
<emphasis id="FDC181224F2DFF8921E60F5CFD72ED71" bold="true" box="[199,669,1825,1850]" pageId="5" pageNumber="140">(13) Microlepides on subimaginal tarsi.</emphasis>
All tarsal segments of all legs of both sexes are covered by pointed microlepides (examined for both sexes of 
<taxonomicName id="08B526B34F2DFF8923510F38FCF2ED16" box="[624,797,1861,1885]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2DFF8923510F38FCF2ED16" box="[624,797,1861,1885]" italics="true" pageId="5" pageNumber="140">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2DFF8922040F38FC8AED15" box="[805,869,1861,1886]" pageId="5" pageNumber="140" rank="species">
<emphasis id="FDC181224F2DFF8922040F38FC8AED15" bold="true" box="[805,869,1861,1886]" pageId="5" pageNumber="140">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F2DFF8922520F38FBD3ED16" box="[883,1084,1861,1885]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F2DFF8922520F38FBD3ED16" box="[883,1084,1861,1885]" italics="true" pageId="5" pageNumber="140">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2DFF8925620F38FB6CED15" box="[1091,1155,1861,1886]" pageId="5" pageNumber="140" rank="species">
<emphasis id="FDC181224F2DFF8925620F38FB6CED15" bold="true" box="[1091,1155,1861,1886]" pageId="5" pageNumber="140">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="08B526B34F2DFF89259B0F38FA9CED16" box="[1210,1395,1861,1885]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F2DFF89259B0F38FA9CED16" box="[1210,1395,1861,1885]" italics="true" pageId="5" pageNumber="140">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2DFF89245B0F38FF41EDC9" pageId="5" pageNumber="140" rank="species">
<emphasis id="FDC181224F2DFF89245B0F38FF41EDC9" bold="true" pageId="5" pageNumber="140">sp. n.</emphasis>
</taxonomicNameLabel>
, males of 
<taxonomicName id="08B526B34F2DFF8920110F17FE59EDC9" box="[304,438,1898,1922]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F2DFF8920110F17FE59EDC9" box="[304,438,1898,1922]" italics="true" pageId="5" pageNumber="140">Ch. soldani</emphasis>
</taxonomicName>
and 
<emphasis id="FDC181224F2DFF8920D40F17FDF9EDC9" box="[501,534,1898,1922]" italics="true" pageId="5" pageNumber="140">Ch</emphasis>
. sp. 1, females of 
<taxonomicName id="08B526B34F2DFF8923DB0F17FC7BEDC9" box="[762,916,1898,1922]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2DFF8923DB0F17FC7BEDC9" box="[762,916,1898,1922]" italics="true" pageId="5" pageNumber="140">Ch. yolandae</emphasis>
</taxonomicName>
and 
<emphasis id="FDC181224F2DFF8922F20F17FC1BEDC9" box="[979,1012,1898,1922]" italics="true" pageId="5" pageNumber="140">Ch</emphasis>
. sp. 2). The same in Protopatellata, Cloeonini, 
<taxonomicName id="08B526B34F2DFF8920350FF0FD2EEDEE" authority="Baetopus, Afrobaetodes" authorityName="Baetopus, Afrobaetodes" box="[276,705,1933,1957]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2DFF8920350FF0FE47EDEE" box="[276,424,1933,1957]" italics="true" pageId="5" pageNumber="140">Centroptilum</emphasis>
, 
<emphasis id="FDC181224F2DFF8920970FF3FDF1EDEE" box="[438,542,1934,1957]" italics="true" pageId="5" pageNumber="140">Baetopus</emphasis>
, 
<emphasis id="FDC181224F2DFF89230C0FF0FD2EEDEE" box="[557,705,1933,1957]" italics="true" pageId="5" pageNumber="140">Afrobaetodes</emphasis>
</taxonomicName>
and some Baetovectata; in contrast to this, in some Baetovectata tarsi are partly or completely covered by blunt microlepides (Kluge &amp; Novikova 2014: 
<figureCitation id="578E41B54F2DFF8925450FCCFB0BED81" box="[1124,1252,1969,1994]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="5" pageNumber="140">Figs 11–17</figureCitation>
). Such character pattern testifies that this is a plesiomorphy of 
<taxonomicName id="08B526B34F2DFF8923B70FABFCDFEDA5" box="[662,816,2006,2030]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2DFF8923B70FABFCDFEDA5" box="[662,816,2006,2030]" italics="true" pageId="5" pageNumber="140">Cheleocloeon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E608EAFCE4EB0B" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<emphasis id="FDC181224F2EFF8A21E608EAFE62EAFB" bold="true" box="[199,397,151,176]" pageId="6" pageNumber="141">(14) Larval claw</emphasis>
(
<figureCitation id="578E41B54F2EFF8A20BC08EAFD97EAFB" box="[413,632,151,176]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="6" pageNumber="141">Figs 23, 24, 27–29</figureCitation>
). Claw is slender, slightly bent, with 2 rows of denticles, among which minute denticles are located close to base of claw and larger denticles, if present, occupy more distal part of claw. In some species only minute proximal denticles are present. Larger denticles can be either well-expressed, or very delicate, thin-walled, translucent and poorly visible; they can be absent on individual claws. This is true for all species of 
<taxonomicName id="08B526B34F2EFF8A202C0955FCE8EB0B" authority="Gattolliat &amp; Sartori 2008: 52" authorityName="Gattolliat &amp; Sartori" authorityPageNumber="52" authorityYear="2008" box="[269,775,295,320]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A202C0955FE48EB0B" box="[269,423,296,320]" italics="true" pageId="6" pageNumber="141">Cheleocloeon</emphasis>
(Gattolliat &amp; Sartori 2008: 52)
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60930FC5CEB9B" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
In the generic diagnosis of 
<taxonomicName id="08B526B34F2EFF8A20D90931FD7DEB2F" box="[504,658,332,356]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A20D90931FD7DEB2F" box="[504,658,332,356]" italics="true" pageId="6" pageNumber="141">Cheleocloeon</emphasis>
</taxonomicName>
, based on two species, 
<taxonomicName id="08B526B34F2EFF8A22BA0931FBDEEB2F" box="[923,1073,332,356]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2EFF8A22BA0931FBDEEB2F" box="[923,1073,332,356]" italics="true" pageId="6" pageNumber="141">Ch. yolandae</emphasis>
</taxonomicName>
and 
<taxonomicName id="08B526B34F2EFF8A25460931FEDBEBC3" authority="Wuillot and Gillies (1993)" authorityName="Wuillot and Gillies" authorityYear="1993" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F2EFF8A25460931FAE6EB2F" box="[1127,1289,332,356]" italics="true" pageId="6" pageNumber="141">Ch. carinatum</emphasis>
, 
<bibRefCitation id="AB2420C14F2EFF8A24360931FEDBEBC3" author="Wuillot" pageId="6" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot and Gillies (1993)</bibRefCitation>
</taxonomicName>
wrote that “claws long with minute teeth at base”. Actually, on the larval exuviae of 
<taxonomicName id="08B526B34F2EFF8A2424090DFA73EBC3" box="[1285,1436,368,392]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2EFF8A2424090DFA73EBC3" box="[1285,1436,368,392]" italics="true" pageId="6" pageNumber="141">Ch. yolandae</emphasis>
</taxonomicName>
examined by me, all six claws have, besides two rows of minute denticles near base, 2+2 larger denticles (
<figureCitation id="578E41B54F2EFF8A241A09E9FA61EBE7" box="[1339,1422,404,429]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="6" pageNumber="141">Fig. 23</figureCitation>
); they are so delicate and translucent, that they can be easily overlooked.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E609A1FF09E853" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
On the larval exuviae of 
<taxonomicName id="08B526B34F2EFF8A20D509A1FD74EBBF" box="[500,667,476,500]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F2EFF8A20D509A1FD74EBBF" box="[500,667,476,500]" italics="true" pageId="6" pageNumber="141">Ch. carinatum</emphasis>
</taxonomicName>
examined, only minute denticles are present, larger denticles are absent.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60A58FE9DE8E3" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
In the descriptions and figures of 
<taxonomicName id="08B526B34F2EFF8A231E0A59FD27E877" box="[575,712,548,572]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F2EFF8A231E0A59FD27E877" box="[575,712,548,572]" italics="true" pageId="6" pageNumber="141">Ch. excisum</emphasis>
</taxonomicName>
by 
<bibRefCitation id="AB2420C14F2EFF8A23D30A59FBAEE876" author="Barnard" box="[754,1089,548,573]" pageId="6" pageNumber="166" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. http: // dx. doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932" yearSuffix="k">
Barnard (1932: 226, 
<figureCitation id="578E41B54F2EFF8A22FA0A59FBC6E876" box="[987,1065,548,573]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 15</figureCitation>
k)
</bibRefCitation>
and 
<bibRefCitation id="AB2420C14F2EFF8A25590A59FE99E82B" author="Lugo-Ortiz" pageId="6" pageNumber="167" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1997 a) Contribution to the systematics of the genus Cheleocloeon (Ephemeroptera: Baetidae). Entomological News, 108 (4), 283 - 289" type="journal article" year="1997" yearSuffix="a">
Lugo-Ortiz &amp; McCafferty (1997a: 286, 
<figureCitation id="578E41B54F2EFF8A200A0A3AFE82E82B" box="[299,365,583,608]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 9</figureCitation>
)
</bibRefCitation>
, the claws are described as “with three to four poorly developed denticles”. Actually each claw has two rows of denticles, each row consists of minute denticles proximally and 4–7 larger denticles more distally (similar to 
<figureCitation id="578E41B54F2EFF8A20330AF2FE8AE8E3" box="[274,357,655,680]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="6" pageNumber="141">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60AC9FA84E8BB" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
According to the original description of 
<taxonomicName id="08B526B34F2EFF8A23AE0AC9FCFEE887" box="[655,785,692,716]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F2EFF8A23AE0AC9FCFEE887" box="[655,785,692,716]" italics="true" pageId="6" pageNumber="141">Ch. soldani</emphasis>
</taxonomicName>
, claw “with one row of 6 reduced teeth proximally and 7 stout teeth apically” (Gattolliat &amp; Sartori 2008: 52). Actually it has two equal rows of such denticles (
<figureCitation id="578E41B54F2EFF8A242B0AAAFAB1E8BB" box="[1290,1374,727,752]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="6" pageNumber="141">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60A81FCABE973" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
All three new species, 
<taxonomicName id="08B526B34F2EFF8A20E40A81FD9DE95F" box="[453,626,764,788]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2EFF8A20E40A81FD9DE95F" box="[453,626,764,788]" italics="true" pageId="6" pageNumber="141">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2EFF8A23580A81FD57E95E" box="[633,696,764,789]" pageId="6" pageNumber="141" rank="species">
<emphasis id="FDC181224F2EFF8A23580A81FD57E95E" bold="true" box="[633,696,764,789]" pageId="6" pageNumber="141">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F2EFF8A23E70A81FC61E95F" box="[710,910,764,788]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F2EFF8A23E70A81FC61E95F" box="[710,910,764,788]" italics="true" pageId="6" pageNumber="141">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2EFF8A22B40A81FC3AE95E" box="[917,981,764,789]" pageId="6" pageNumber="141" rank="species">
<emphasis id="FDC181224F2EFF8A22B40A81FC3AE95E" bold="true" box="[917,981,764,789]" pageId="6" pageNumber="141">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="08B526B34F2EFF8A252A0A81FB2BE95F" box="[1035,1220,764,788]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F2EFF8A252A0A81FB2BE95F" box="[1035,1220,764,788]" italics="true" pageId="6" pageNumber="141">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2EFF8A25EA0A81FAE4E95E" box="[1227,1291,764,789]" pageId="6" pageNumber="141" rank="species">
<emphasis id="FDC181224F2EFF8A25EA0A81FAE4E95E" bold="true" box="[1227,1291,764,789]" pageId="6" pageNumber="141">sp. n.</emphasis>
</taxonomicNameLabel>
have at least two rows of minute denticles near base of claw (
<figureCitation id="578E41B54F2EFF8A23940B62FCD6E973" box="[693,825,799,824]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="6" pageNumber="141">Figs 27–29</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60B39FB3CE9EF" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<emphasis id="FDC181224F2EFF8A21E60B39FE92E916" bold="true" box="[199,381,836,861]" pageId="6" pageNumber="141">
(15) 
<collectingCountry id="B7A21DA04F2EFF8A21DF0B39FED4E916" box="[254,315,836,861]" name="India" pageId="6" pageNumber="141">Hind</collectingCountry>
wing
</emphasis>
(
<figureCitation id="578E41B54F2EFF8A20AD0B39FDE0E917" box="[396,527,836,861]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Figs 10–13</figureCitation>
, 
<figureCitation id="578E41B54F2EFF8A233D0B39FD6DE917" box="[540,642,836,860]" captionStart="FIGURES 107 – 112. 107 – 109" captionStartId="24.[151,264,1860,1883]" captionTargetBox="[177,1387,202,1819]" captionTargetId="figure@24.[177,1400,193,1824]" captionTargetPageId="24" captionText="FIGURES 107 – 112. 107 – 109, Cheleocloeon lancetofolium sp. n.; 110 – 112, Ch. truncifolium sp. n. 107, 111, subimaginal exuviae of genitals; 108, 110, imaginal genitals, ventral view (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines; see Fig. 105); 109, 112, hind wing." httpUri="https://zenodo.org/record/270609/files/figure.png" pageId="6" pageNumber="141">109, 112</figureCitation>
). 
<collectingCountry id="B7A21DA04F2EFF8A23B60B39FD20E916" box="[663,719,836,861]" name="India" pageId="6" pageNumber="141">Hind</collectingCountry>
wing, if present, is of the « 
<taxonomicName id="08B526B34F2EFF8A252A0B39FB70E917" box="[1035,1183,836,860]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A252A0B39FB70E917" box="[1035,1183,836,860]" italics="true" pageId="6" pageNumber="141">Centroptilum</emphasis>
</taxonomicName>
- 
<typeStatus id="100EE3924F2EFF8A25860B38FB32E916" box="[1191,1245,837,861]" pageId="6" pageNumber="141">type</typeStatus>
»: narrow, with 2 longitudinal veins only, with hooked costal projection (
<figureCitation id="578E41B54F2EFF8A22270B1AFCB7E9CB" box="[774,856,871,896]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 11</figureCitation>
); larval hind protopteron is narrow, slightly S-bent, narrowed toward apex (
<figureCitation id="578E41B54F2EFF8A20810BF1FE1BE9EF" box="[416,500,908,933]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 10</figureCitation>
). Females never have hind wings; males either have them, or not.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60BCDFBF3EE13" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<taxonomicName id="08B526B34F2EFF8A21E60BCDFE99E983" box="[199,374,944,968]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2EFF8A21E60BCDFE99E983" box="[199,374,944,968]" italics="true" pageId="6" pageNumber="141">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2EFF8A20A10BD2FE2CE983" box="[384,451,943,968]" pageId="6" pageNumber="141" rank="species">
<emphasis id="FDC181224F2EFF8A20A10BD2FE2CE983" bold="true" box="[384,451,943,968]" pageId="6" pageNumber="141">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F2EFF8A20F20BCDFD4FE983" box="[467,672,944,968]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F2EFF8A20F20BCDFD4FE983" box="[467,672,944,968]" italics="true" pageId="6" pageNumber="141">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2EFF8A23880BD2FD04E983" box="[681,747,943,968]" pageId="6" pageNumber="141" rank="species">
<emphasis id="FDC181224F2EFF8A23880BD2FD04E983" bold="true" box="[681,747,943,968]" pageId="6" pageNumber="141">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="08B526B34F2EFF8A22060BCDFC0CE983" box="[807,995,944,968]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F2EFF8A22060BCDFC0CE983" box="[807,995,944,968]" italics="true" pageId="6" pageNumber="141">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2EFF8A22CC0BD2FBC0E983" box="[1005,1071,943,968]" pageId="6" pageNumber="141" rank="species">
<emphasis id="FDC181224F2EFF8A22CC0BD2FBC0E983" bold="true" box="[1005,1071,943,968]" pageId="6" pageNumber="141">sp. n.</emphasis>
</taxonomicNameLabel>
have hind wings in male; their female imagoes have vestiges of hind wings, which represent very small immovable membranous projections, whose length is equal to length of vestige of larval protopteron (
<figureCitation id="578E41B54F2EFF8A22A00B8AFC39EE5B" box="[897,982,1015,1040]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 13</figureCitation>
); vestige of larval protopteron is rather wide and projected posteriad of posterior margin of metanotum (
<figureCitation id="578E41B54F2EFF8A22530C61FC28EE7F" box="[882,967,1052,1077]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 12</figureCitation>
). The same vestiges of larval protopteron are present in female larvae of 
<taxonomicName id="08B526B34F2EFF8A20D00C3DFD95EE1C" box="[497,634,1088,1112]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F2EFF8A20D00C3DFD95EE1C" box="[497,634,1088,1112]" italics="true" pageId="6" pageNumber="141">Ch. excisum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F2EFF8A23A90C3DFCAFEE13" authority="Ch." authorityName="Ch." box="[648,832,1088,1112]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F2EFF8A23A90C3DFCE6EE13" box="[648,777,1088,1112]" italics="true" pageId="6" pageNumber="141">Ch. soldani</emphasis>
, 
<emphasis id="FDC181224F2EFF8A22360C3DFCAFEE13" box="[791,832,1088,1112]" italics="true" pageId="6" pageNumber="141">Ch.</emphasis>
</taxonomicName>
sp. 2 and 
<emphasis id="FDC181224F2EFF8A22900C3DFC36EE13" box="[945,985,1088,1112]" italics="true" pageId="6" pageNumber="141">Ch.</emphasis>
sp. 3.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60C19FE1CEE8F" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<taxonomicName id="08B526B34F2EFF8A21E60C19FE93EE37" box="[199,380,1124,1148]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="dimorphicum">
<emphasis id="FDC181224F2EFF8A21E60C19FE93EE37" box="[199,380,1124,1148]" italics="true" pageId="6" pageNumber="141">C. dimorphicum</emphasis>
</taxonomicName>
, according to the original description, also has hind wings in male, but its female imago has no vestiges of hind wings, and larval protoptera are “absent or considerably reduced, empty and rudimentary” (
<bibRefCitation id="AB2420C14F2EFF8A246C0CFAFE09EE8F" author="Soldan" pageId="6" pageNumber="167" refString="Soldan, T. &amp; Thomas, A. G. B. (1985) Centroptilum dimorphicum sp. n., a new species of mayfly (Ephemeroptera, Baetidae) from Algeria. Acta Entomologica Bohemoslovaca, 82, 180 - 186." type="journal article" year="1985">
Soldán &amp; Thomas 1985: 182, 
<figureCitation id="578E41B54F2EFF8A20B50CD1FE09EE8F" box="[404,486,1196,1221]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 10</figureCitation>
</bibRefCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60CADFEFDEF47" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<taxonomicName id="08B526B34F2EFF8A21E60CADFEB0EEA3" box="[199,351,1232,1256]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2EFF8A21E60CADFEB0EEA3" box="[199,351,1232,1256]" italics="true" pageId="6" pageNumber="141">Ch. yolandae</emphasis>
</taxonomicName>
and 
<taxonomicName id="08B526B34F2EFF8A20B80CADFDD2EEAC" box="[409,573,1232,1256]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F2EFF8A20B80CADFDD2EEAC" box="[409,573,1232,1256]" italics="true" pageId="6" pageNumber="141">Ch. carinatum</emphasis>
</taxonomicName>
have no hind wings in male; at least females have no any vestiges of hind protoptera.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60D65FE76EF8B" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
Contrary to the statement that females of 
<taxonomicName id="08B526B34F2EFF8A23890D65FCADEF7B" box="[680,834,1304,1328]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A23890D65FCADEF7B" box="[680,834,1304,1328]" italics="true" pageId="6" pageNumber="141">Cheleocloeon</emphasis>
</taxonomicName>
never have developed hind wings, two species with hind wings in the female were attributed to 
<taxonomicName id="08B526B34F2EFF8A23B30D41FCC3EF1F" box="[658,812,1340,1364]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A23B30D41FCC3EF1F" box="[658,812,1340,1364]" italics="true" pageId="6" pageNumber="141">Cheleocloeon</emphasis>
</taxonomicName>
. One of them, 
<taxonomicName id="08B526B34F2EFF8A22FC0D41FB27EF1F" authority="Crass 1947" authorityName="Crass" authorityYear="1947" box="[989,1224,1340,1364]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="falcatum">
<emphasis id="FDC181224F2EFF8A22FC0D41FBD0EF1F" box="[989,1087,1340,1364]" italics="true" pageId="6" pageNumber="141">falcatum</emphasis>
<bibRefCitation id="AB2420C14F2EFF8A25680D41FB27EF1F" author="Crass" box="[1097,1224,1340,1364]" pageId="6" pageNumber="167" refString="Crass, R. S. (1947) The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum, 11 (1), 37 - 100." type="journal article" year="1947">Crass 1947</bibRefCitation>
</taxonomicName>
[
<taxonomicName id="08B526B34F2EFF8A25FA0D41FA80EF1F" box="[1243,1391,1340,1364]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A25FA0D41FA80EF1F" box="[1243,1391,1340,1364]" italics="true" pageId="6" pageNumber="141">Centroptilum</emphasis>
</taxonomicName>
], is known from imagoes only, and does not belong to 
<taxonomicName id="08B526B34F2EFF8A23C90D1DFC6DEF33" box="[744,898,1376,1400]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A23C90D1DFC6DEF33" box="[744,898,1376,1400]" italics="true" pageId="6" pageNumber="141">Cheleocloeon</emphasis>
</taxonomicName>
(see below). Another species, 
<taxonomicName id="08B526B34F2EFF8A25C90D1DFAA2EF33" box="[1256,1357,1376,1400]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="mirandei">
<emphasis id="FDC181224F2EFF8A25C90D1DFAA2EF33" box="[1256,1357,1376,1400]" italics="true" pageId="6" pageNumber="141">mirandei</emphasis>
</taxonomicName>
Lugo- Ortiz &amp; McCafferty 1997 [
<taxonomicName id="08B526B34F2EFF8A20EA0DF9FD8AEFD7" box="[459,613,1412,1436]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A20EA0DF9FD8AEFD7" box="[459,613,1412,1436]" italics="true" pageId="6" pageNumber="141">Cheleocloeon</emphasis>
</taxonomicName>
] is known as a single female larva; recently it was moved to 
<taxonomicName id="08B526B34F2EFF8A24020DF9FE7AEF8B" authority="Gattolliat et al. 2009" authorityName="Gattolliat et al." authorityYear="2009" class="Insecta" family="Baetidae" genus="Bugilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A24020DF9FA73EFD7" box="[1315,1436,1412,1436]" italics="true" pageId="6" pageNumber="141">Bugilliesia</emphasis>
(
<bibRefCitation id="AB2420C14F2EFF8A21BF0DDAFE62EF8B" author="Gattolliat" box="[158,397,1447,1472]" pageId="6" pageNumber="167" refString="Gattolliat, J. - L., Barber-James, H. M. &amp; Monaghan, M. T. (2009) New species and generic delimitation of the Afrotropical genera Bugilliesia Lugo-Ortiz &amp; McCafferty, 1996; Cheleocloeon Wuillot &amp; Gillies, 1993 and Delouardus Lugo-Ortiz &amp; McCafferty, 1999 (Ephemeroptera: Baetidae). Aquatic Insects, 31 (3), 167 - 186. http: // dx. doi. org / 10.1080 / 01650420902782987" type="journal article" year="2009">
Gattolliat 
<emphasis id="FDC181224F2EFF8A20300DD4FEA6EF8B" box="[273,329,1448,1472]" italics="true" pageId="6" pageNumber="141">et al.</emphasis>
2009
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60DB1FE85ECF7" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<emphasis id="FDC181224F2EFF8A21E60DB1FD8DEFAE" bold="true" box="[199,610,1484,1509]" pageId="6" pageNumber="141">(16) Denticles on larval abdomen.</emphasis>
Posterior margins of abdominal terga (at least terga II–X) bear pointed denticles, which can be larger (
<figureCitation id="578E41B54F2EFF8A232E0D92FD89EC43" box="[527,614,1519,1544]" captionStart="FIGURES 58 – 63" captionStartId="11.[151,264,1736,1759]" captionTargetBox="[151,1434,176,1715]" captionTargetId="figure@11.[151,1435,159,1715]" captionTargetPageId="11" captionText="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." httpUri="https://zenodo.org/record/270601/files/figure.png" pageId="6" pageNumber="141">Fig. 62</figureCitation>
) or smaller. In 
<taxonomicName id="08B526B34F2EFF8A22060D8DFC21EC4C" box="[807,974,1520,1544]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F2EFF8A22060D8DFC21EC4C" box="[807,974,1520,1544]" italics="true" pageId="6" pageNumber="141">Ch. carinatum</emphasis>
</taxonomicName>
, whose abdominal terga bear unpaired projections, these denticles border the margin of each projection (
<figureCitation id="578E41B54F2EFF8A22A50E69FC35EC67" box="[900,986,1556,1581]" captionStart="FIGURES 64 – 70" captionStartId="12.[151,264,1899,1922]" captionTargetBox="[150,1436,193,1878]" captionTargetId="figure@12.[150,1436,193,1878]" captionTargetPageId="12" captionText="FIGURES 64 – 70. Details of exuviae of last instar larva (64 – 69 under the same magnification): 64 – 66, Cheleocloeon yolandae; 67 – 70, Ch. carinatum." httpUri="https://zenodo.org/record/270602/files/figure.png" pageId="6" pageNumber="141">Fig. 70</figureCitation>
). Posterolateral spines are absent on all abdominal segments (as in most 
<taxonomicName id="08B526B34F2EFF8A23010E4AFD68EC1B" box="[544,647,1591,1616]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
). Lateral margins of all abdominal segments have no denticles (in contrast to Cloeoninae and 
<taxonomicName id="08B526B34F2EFF8A20F00E21FD8AEC3F" box="[465,613,1628,1652]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A20F00E21FD8AEC3F" box="[465,613,1628,1652]" italics="true" pageId="6" pageNumber="141">Anafroptilum</emphasis>
</taxonomicName>
). Posterior margin of tenth abdominal tergum is semicircular with even denticles, as on previous terga (lateral denticles are not enlarged, in contrast to 
<taxonomicName id="08B526B34F2EFF8A25280EFDFB92ECD3" box="[1033,1149,1664,1688]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A25280EFDFB92ECD3" box="[1033,1149,1664,1688]" italics="true" pageId="6" pageNumber="141">Procloeon</emphasis>
</taxonomicName>
). Paraprocts bear pointed denticles (
<figureCitation id="578E41B54F2EFF8A202B0ED9FEB2ECF7" box="[266,349,1700,1725]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60EBAFD87ED07" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<emphasis id="FDC181224F2EFF8A21E60EBAFDDAECAB" bold="true" box="[199,565,1735,1760]" pageId="6" pageNumber="141">(17) Scales of larval abdomen.</emphasis>
Abdominal terga, sterna and caudalii bear translucent scales; socket of such scale is W-shaped, i.e. with two concavities; each concavity is covered by a short cover, which is not projected distad of the concavity (
<figureCitation id="578E41B54F2EFF8A20870F72FE12ED63" box="[422,509,1807,1832]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="6" pageNumber="141">Figs 15</figureCitation>
, 
<figureCitation id="578E41B54F2EFF8A23280F72FDBBED63" box="[521,596,1807,1832]" captionStart="FIGURES 58 – 63" captionStartId="11.[151,264,1736,1759]" captionTargetBox="[151,1434,176,1715]" captionTargetId="figure@11.[151,1435,159,1715]" captionTargetPageId="11" captionText="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." httpUri="https://zenodo.org/record/270601/files/figure.png" pageId="6" pageNumber="141">62, 63</figureCitation>
). Scales with two covers are present in various taxa of 
<taxonomicName id="08B526B34F2EFF8A25E10F72FACAED63" box="[1216,1317,1807,1832]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
, but not in Cloeonini, 
<taxonomicName id="08B526B34F2EFF8A20320F49FE48ED07" box="[275,423,1844,1868]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A20320F49FE48ED07" box="[275,423,1844,1868]" italics="true" pageId="6" pageNumber="141">Centroptilum</emphasis>
</taxonomicName>
or 
<taxonomicName id="08B526B34F2EFF8A20EC0F49FD8EED07" box="[461,609,1844,1868]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2EFF8A20EC0F49FD8EED07" box="[461,609,1844,1868]" italics="true" pageId="6" pageNumber="141">Anafroptilum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F2EFF8A21E60F2AFC16EDDE" blockId="6.[151,1437,151,1941]" pageId="6" pageNumber="141">
<emphasis id="FDC181224F2EFF8A21E60F2AFDD4ED3B" bold="true" box="[199,571,1879,1904]" pageId="6" pageNumber="141">(18) Setae of larval abdomen.</emphasis>
Abdominal terga and sterna, besides scales [see (17)] have only sparse, irregularly situated simple setae, which do not form rows or groups (
<figureCitation id="578E41B54F2EFF8A22B90F01FC03EDDE" box="[920,1004,1916,1941]" captionStart="FIGURES 58 – 63" captionStartId="11.[151,264,1736,1759]" captionTargetBox="[151,1434,176,1715]" captionTargetId="figure@11.[151,1435,159,1715]" captionTargetPageId="11" captionText="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." httpUri="https://zenodo.org/record/270601/files/figure.png" pageId="6" pageNumber="141">Fig. 63</figureCitation>
).
</paragraph>
<caption id="9BCA0DB84F2FFF8B21B60F26FF23EDA7" httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="7" pageNumber="142" targetBox="[156,1432,199,1855]" targetPageId="7">
<paragraph id="CF0A5D304F2FFF8B21B60F26FF23EDA7" blockId="7.[151,1436,1883,2028]" pageId="7" pageNumber="142">
<emphasis id="FDC181224F2FFF8B21B60F26FEB9ED3B" bold="true" box="[151,342,1883,1906]" pageId="7" pageNumber="142">FIGURES 19–29.</emphasis>
19–23, 
<taxonomicName id="08B526B34F2FFF8B208C0F26FD71ED3A" box="[429,670,1883,1905]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F2FFF8B208C0F26FD71ED3A" box="[429,670,1883,1905]" italics="true" pageId="7" pageNumber="142">Cheleocloeon yolandae</emphasis>
</taxonomicName>
: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24–25, 
<taxonomicName id="08B526B34F2FFF8B22050FE4FC73EDE4" box="[804,924,1945,1967]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F2FFF8B22050FE4FC73EDE4" box="[804,924,1945,1967]" italics="true" pageId="7" pageNumber="142">Ch. soldani</emphasis>
</taxonomicName>
: 24, claw; 25, larval fore leg, anterior view. 26, 
<taxonomicName id="08B526B34F2FFF8B21B60FCAFECDED86" box="[151,290,1975,1997]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F2FFF8B21B60FCAFECDED86" box="[151,290,1975,1997]" italics="true" pageId="7" pageNumber="142">Cheleocloeon</emphasis>
</taxonomicName>
sp. 3, larval fore leg, anterior view. 27–28, 
<taxonomicName id="08B526B34F2FFF8B23FD0FCAFC97ED86" box="[732,888,1975,1997]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F2FFF8B23FD0FCAFC97ED86" box="[732,888,1975,1997]" italics="true" pageId="7" pageNumber="142">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2FFF8B225F0FC5FC59ED86" box="[894,950,1976,1997]" pageId="7" pageNumber="142" rank="species">
<emphasis id="FDC181224F2FFF8B225F0FC5FC59ED86" bold="true" box="[894,950,1976,1997]" pageId="7" pageNumber="142">sp. n.</emphasis>
</taxonomicNameLabel>
, larval claws. 29, 
<taxonomicName id="08B526B34F2FFF8B254E0FCAFAF9ED86" box="[1135,1302,1975,1997]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F2FFF8B254E0FCAFAF9ED86" box="[1135,1302,1975,1997]" italics="true" pageId="7" pageNumber="142">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F2FFF8B243C0FC5FAB9ED86" box="[1309,1366,1976,1997]" pageId="7" pageNumber="142" rank="species">
<emphasis id="FDC181224F2FFF8B243C0FC5FAB9ED86" bold="true" box="[1309,1366,1976,1997]" pageId="7" pageNumber="142">sp. n.</emphasis>
</taxonomicNameLabel>
, larval claw.
</paragraph>
</caption>
<caption id="9BCA0DB84F20FF8421B60E6FFC4BEC0C" httpUri="https://zenodo.org/record/270600/files/figure.png" pageId="8" pageNumber="143" targetBox="[153,1432,201,1529]" targetPageId="8">
<paragraph id="CF0A5D304F20FF8421B60E6FFC4BEC0C" blockId="8.[151,1436,1554,1607]" pageId="8" pageNumber="143">
<emphasis id="FDC181224F20FF8421B60E6FFEB6EC63" bold="true" box="[151,345,1554,1577]" pageId="8" pageNumber="143">FIGURES 30–57.</emphasis>
Tergalii I–VII: 30–36, 
<taxonomicName id="08B526B34F20FF84237B0E6EFC8DEC62" box="[602,866,1555,1577]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F20FF84237B0E6EFC8DEC62" box="[602,866,1555,1577]" italics="true" pageId="8" pageNumber="143">Cheleocloeon clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F20FF84224F0E6EFC43EC63" box="[878,940,1555,1576]" pageId="8" pageNumber="143" rank="species">
<emphasis id="FDC181224F20FF84224F0E6EFC43EC63" bold="true" box="[878,940,1555,1576]" pageId="8" pageNumber="143">sp. n.</emphasis>
</taxonomicNameLabel>
; 37–43, 
<taxonomicName id="08B526B34F20FF8425310E6EFB25EC62" box="[1040,1226,1555,1577]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F20FF8425310E6EFB25EC62" box="[1040,1226,1555,1577]" italics="true" pageId="8" pageNumber="143">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F20FF8425F70E6EFAFBEC63" box="[1238,1300,1555,1576]" pageId="8" pageNumber="143" rank="species">
<emphasis id="FDC181224F20FF8425F70E6EFAFBEC63" bold="true" box="[1238,1300,1555,1576]" pageId="8" pageNumber="143">sp. n.</emphasis>
</taxonomicNameLabel>
; 44–50, 
<taxonomicName id="08B526B34F20FF8424590E6EFEFBEC0C" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F20FF8424590E6EFEFBEC0C" italics="true" pageId="8" pageNumber="143">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F20FF84203B0E4FFEBCEC0C" box="[282,339,1586,1607]" pageId="8" pageNumber="143" rank="species">
<emphasis id="FDC181224F20FF84203B0E4FFEBCEC0C" bold="true" box="[282,339,1586,1607]" pageId="8" pageNumber="143">sp. n.</emphasis>
</taxonomicNameLabel>
(male, holotype); 51–57, 
<taxonomicName id="08B526B34F20FF84237B0E4CFCEEEC0C" box="[602,769,1585,1607]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F20FF84237B0E4CFCEEEC0C" box="[602,769,1585,1607]" italics="true" pageId="8" pageNumber="143">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F20FF8422290E4FFCAFEC0C" box="[776,832,1586,1607]" pageId="8" pageNumber="143" rank="species">
<emphasis id="FDC181224F20FF8422290E4FFCAFEC0C" bold="true" box="[776,832,1586,1607]" pageId="8" pageNumber="143">sp. n.</emphasis>
</taxonomicNameLabel>
(female).
</paragraph>
</caption>
<paragraph id="CF0A5D304F20FF8421E60E13FB66EC84" blockId="8.[151,1436,1646,2030]" pageId="8" pageNumber="143">
<emphasis id="FDC181224F20FF8421E60E13FE1CECCC" bold="true" box="[199,499,1646,1671]" pageId="8" pageNumber="143">(19) Structure of tergalii</emphasis>
(
<figureCitation id="578E41B54F20FF8423270E13FD62ECCC" box="[518,653,1646,1671]" captionStart="FIGURES 30 – 57" captionStartId="8.[151,264,1554,1577]" captionTargetBox="[153,1432,201,1529]" captionTargetId="figure@8.[151,1436,193,1533]" captionTargetPageId="8" captionText="FIGURES 30 – 57. Tergalii I – VII: 30 – 36, Cheleocloeon clavifolium sp. n.; 37 – 43, Ch. lancetofolium sp. n.; 44 – 50, Ch. truncifolium sp. n. (male, holotype); 51 – 57, Ch. truncifolium sp. n. (female)." httpUri="https://zenodo.org/record/270600/files/figure.png" pageId="8" pageNumber="143">Figs 30–57</figureCitation>
). Costal and anal ribs can be rather long, but not reaching apex of tergalius, so that distal margin of tergalius is not bordered by rib. First tergalius is narrower than others, more or less petiolate; its length can be either less, or equal, or greater than length of other tergalii.
</paragraph>
<paragraph id="CF0A5D304F20FF8421E60EA6FBFFEDC8" blockId="8.[151,1436,1646,2030]" pageId="8" pageNumber="143">
<emphasis id="FDC181224F20FF8421E60EA6FE00ECBF" bold="true" box="[199,495,1755,1780]" pageId="8" pageNumber="143">(20) Mobility of tergalii.</emphasis>
Tergalii are able to make rhythmical respiratory movements. 
<bibRefCitation id="AB2420C14F20FF8425E10EA6FF0EED5C" author="Wuillot" pageId="8" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot and Gillies (1993)</bibRefCitation>
wrote about 
<taxonomicName id="08B526B34F20FF8420550E82FDE5ED5C" box="[372,522,1791,1815]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F20FF8420550E82FDE5ED5C" box="[372,522,1791,1815]" italics="true" pageId="8" pageNumber="143">Ch. yolandae</emphasis>
</taxonomicName>
and/or 
<taxonomicName id="08B526B34F20FF84237E0E82FCEFED5C" box="[607,768,1791,1815]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F20FF84237E0E82FCEFED5C" box="[607,768,1791,1815]" italics="true" pageId="8" pageNumber="143">Ch. carinatum</emphasis>
</taxonomicName>
: “In living nymphs, while at rest, the first gill is held above the body like a banner and actively vibrated. In contrast, the remaining gills are held passively, close to the abdomen”. Actually other tergalii are also able to make respiratory vibration. I observed respiratory vibrations of 
<taxonomicName id="08B526B34F20FF8421B60F16FEC3EDC8" box="[151,300,1899,1923]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F20FF8421B60F16FEC3EDC8" box="[151,300,1899,1923]" italics="true" pageId="8" pageNumber="143">Ch. yolandae</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F20FF84201B0F16FE2DEDC8" box="[314,450,1899,1923]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F20FF84201B0F16FE2DEDC8" box="[314,450,1899,1923]" italics="true" pageId="8" pageNumber="143">Ch. excisum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F20FF8420F00F16FD91EDC8" box="[465,638,1899,1923]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F20FF8420F00F16FD91EDC8" box="[465,638,1899,1923]" italics="true" pageId="8" pageNumber="143">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F20FF8423A50F16FD2BEDCF" box="[644,708,1899,1924]" pageId="8" pageNumber="143" rank="species">
<emphasis id="FDC181224F20FF8423A50F16FD2BEDCF" bold="true" box="[644,708,1899,1924]" pageId="8" pageNumber="143">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="08B526B34F20FF8423DB0F16FC2CEDC8" box="[762,963,1899,1923]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F20FF8423DB0F16FC2CEDC8" box="[762,963,1899,1923]" italics="true" pageId="8" pageNumber="143">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F20FF8422EB0F16FBE6EDCF" box="[970,1033,1899,1924]" pageId="8" pageNumber="143" rank="species">
<emphasis id="FDC181224F20FF8422EB0F16FBE6EDCF" bold="true" box="[970,1033,1899,1924]" pageId="8" pageNumber="143">sp. n.</emphasis>
</taxonomicNameLabel>
.
</paragraph>
<paragraph id="CF0A5D304F20FF8521E60FF3FC4EEABC" blockId="8.[151,1436,1646,2030]" lastBlockId="9.[151,1437,151,2024]" lastPageId="9" lastPageNumber="144" pageId="8" pageNumber="143">
<emphasis id="FDC181224F20FF8421E60FF3FE51EDEC" bold="true" box="[199,446,1934,1959]" pageId="8" pageNumber="143">(21) Larval caudalii</emphasis>
(
<figureCitation id="578E41B54F20FF8420F00FF3FDB3EDEC" box="[465,604,1934,1959]" captionStart="FIGURES 58 – 63" captionStartId="11.[151,264,1736,1759]" captionTargetBox="[151,1434,176,1715]" captionTargetId="figure@11.[151,1435,159,1715]" captionTargetPageId="11" captionText="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." httpUri="https://zenodo.org/record/270601/files/figure.png" pageId="8" pageNumber="143">Figs 60, 61</figureCitation>
, 
<figureCitation id="578E41B54F20FF84234F0FF3FD54EDEC" box="[622,699,1934,1959]" captionStart="FIGURES 64 – 70" captionStartId="12.[151,264,1899,1922]" captionTargetBox="[150,1436,193,1878]" captionTargetId="figure@12.[150,1436,193,1878]" captionTargetPageId="12" captionText="FIGURES 64 – 70. Details of exuviae of last instar larva (64 – 69 under the same magnification): 64 – 66, Cheleocloeon yolandae; 67 – 70, Ch. carinatum." httpUri="https://zenodo.org/record/270602/files/figure.png" pageId="8" pageNumber="143">65, 68</figureCitation>
, 
<figureCitation id="578E41B54F20FF8423ED0FF3FD05EDEC" box="[716,746,1934,1959]" captionStart="FIGURES 71 – 78" captionStartId="16.[151,264,1837,1860]" captionTargetBox="[151,1434,193,1816]" captionTargetId="figure@16.[151,1435,193,1816]" captionTargetPageId="16" captionText="FIGURES 71 – 78. Details of exuviae of last instar larvae of Cheleocloeon clavifolium sp. n. under the same magnification (72, middle leg; 77, abdomen of holotype)." httpUri="https://zenodo.org/record/270603/files/figure.png" pageId="8" pageNumber="143">75</figureCitation>
, 
<figureCitation id="578E41B54F20FF8423DA0FF2FCF6EDEC" box="[763,793,1935,1959]" captionStart="FIGURES 79 – 86. 79 – 85" captionStartId="18.[151,264,1620,1643]" captionTargetBox="[151,1435,193,1599]" captionTargetId="figure@18.[151,1435,193,1600]" captionTargetPageId="18" captionText="FIGURES 79 – 86. 79 – 85, details of exuviae of last instar larvae under the same magnification: 79 – 81, Cheleocloeon lancetofolium sp. n.; 82 – 84, Ch. truncifolium sp. n. (83, holotype); 85, Cheleocloeon sp. 2; 86, abdomen of female subimago of the same specimen." httpUri="https://zenodo.org/record/270604/files/figure.png" pageId="8" pageNumber="143">80</figureCitation>
, 83). Paracercus is always as long as cerci; cerci and paracercus up to the apex bear well developed primary swimming setae. Each swimming seta is thick and sometimes darkened in proximal part and contrastingly thinner and always colorless in distal part. Distal part of each cercus bears sparse longitudinal row of delicate secondary swimming setae (
<figureCitation id="578E41B54F21FF85251908EAFB63EAFB" box="[1080,1164,151,176]" captionStart="FIGURES 58 – 63" captionStartId="11.[151,264,1736,1759]" captionTargetBox="[151,1434,176,1715]" captionTargetId="figure@11.[151,1435,159,1715]" captionTargetPageId="11" captionText="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." httpUri="https://zenodo.org/record/270601/files/figure.png" pageId="9" pageNumber="144">Fig. 61</figureCitation>
). These features are not reported in literature, but observed by me for 
<taxonomicName id="08B526B34F21FF8523FF08C6FC93EA98" box="[734,892,187,211]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F21FF8523FF08C6FC93EA98" box="[734,892,187,211]" italics="true" pageId="9" pageNumber="144">Ch. yolandae</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F21FF8522B508C6FBD0EA98" box="[916,1087,187,211]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F21FF8522B508C6FBD0EA98" box="[916,1087,187,211]" italics="true" pageId="9" pageNumber="144">Ch. carinatum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F21FF85257708C6FAE3EA98" box="[1110,1292,187,211]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF85257708C6FAE3EA98" box="[1110,1292,187,211]" italics="true" pageId="9" pageNumber="144">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF85243D08C6FA8AEA9F" box="[1308,1381,187,212]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF85243D08C6FA8AEA9F" bold="true" box="[1308,1381,187,212]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="08B526B34F21FF85245508C6FEDEEABC" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F21FF85245508C6FEDEEABC" italics="true" pageId="9" pageNumber="144">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF85201908A3FE98EABC" box="[312,375,222,247]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF85201908A3FE98EABC" bold="true" box="[312,375,222,247]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F21FF8520A408A2FDE1EABC" box="[389,526,223,247]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F21FF8520A408A2FDE1EABC" box="[389,526,223,247]" italics="true" pageId="9" pageNumber="144">Ch. excisum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F21FF85233D08A2FD3DEABC" authority="Ch." authorityName="Ch." box="[540,722,223,247]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F21FF85233D08A2FD73EABC" box="[540,668,223,247]" italics="true" pageId="9" pageNumber="144">Ch. soldani</emphasis>
, 
<emphasis id="FDC181224F21FF85238A08A2FD23EABC" box="[683,716,223,247]" italics="true" pageId="9" pageNumber="144">Ch</emphasis>
.
</taxonomicName>
sp.2 and 
<emphasis id="FDC181224F21FF85221F08A2FCB0EABC" box="[830,863,223,247]" italics="true" pageId="9" pageNumber="144">Ch</emphasis>
. sp.3.
</paragraph>
<paragraph id="CF0A5D304F21FF8521E6097FFDFAEB87" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
Position of denticles on posterior margins of segments of caudalii have species-specific variability: In most species (
<taxonomicName id="08B526B34F21FF8521D6095BFE4BEB75" box="[247,420,294,318]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8521D6095BFE4BEB75" box="[247,420,294,318]" italics="true" pageId="9" pageNumber="144">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF85208B0958FE06EB75" box="[426,489,293,318]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF85208B0958FE06EB75" bold="true" box="[426,489,293,318]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F21FF8520D7095BFD51EB75" box="[502,702,294,318]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8520D7095BFD51EB75" box="[502,702,294,318]" italics="true" pageId="9" pageNumber="144">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF8523E40958FCEBEB75" box="[709,772,293,318]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF8523E40958FCEBEB75" bold="true" box="[709,772,293,318]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F21FF852230095BFC26EB75" box="[785,969,294,318]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF852230095BFC26EB75" box="[785,969,294,318]" italics="true" pageId="9" pageNumber="144">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF8522F10958FBE1EB75" box="[976,1038,293,318]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF8522F10958FBE1EB75" bold="true" box="[976,1038,293,318]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F21FF85253D095BFB4BEB75" box="[1052,1188,294,318]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F21FF85253D095BFB4BEB75" box="[1052,1188,294,318]" italics="true" pageId="9" pageNumber="144">Ch. excisum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F21FF852593095BFA88EB75" authority="Ch." authorityName="Ch." box="[1202,1383,294,318]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F21FF852593095BFADDEB75" box="[1202,1330,294,318]" italics="true" pageId="9" pageNumber="144">Ch. soldani</emphasis>
, 
<emphasis id="FDC181224F21FF852461095BFA8EEB75" box="[1344,1377,294,318]" italics="true" pageId="9" pageNumber="144">Ch</emphasis>
.
</taxonomicName>
sp.2 and 
<emphasis id="FDC181224F21FF8521E70934FF08EB2A" box="[198,231,329,353]" italics="true" pageId="9" pageNumber="144">Ch</emphasis>
. sp.3) enlarged pointed denticles form short transverse rows on each 4th or each 2nd segment on lateral side of each cercus (
<figureCitation id="578E41B54F21FF85206D0911FE70EBCE" box="[332,415,364,389]" captionStart="FIGURES 58 – 63" captionStartId="11.[151,264,1736,1759]" captionTargetBox="[151,1434,176,1715]" captionTargetId="figure@11.[151,1435,159,1715]" captionTargetPageId="11" captionText="FIGURES 58 – 63. Cheleocloeon clavifolium sp. n., larva: 58 – 59, left and right mandibles of penultimolarva ready to molt to ultimolarva; 60, middle part of cercus; 61, distal part of cercus; 62, posterior margin of abdominal tergum VI; antero-lateral area of sternum IV." httpUri="https://zenodo.org/record/270601/files/figure.png" pageId="9" pageNumber="144">Fig. 60</figureCitation>
) and on dorsal side of paracercus, occasionally also on ventral side of paracercus. In 
<taxonomicName id="08B526B34F21FF8524550910FF11EBE3" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F21FF8524550910FF11EBE3" italics="true" pageId="9" pageNumber="144">Ch. yolandae</emphasis>
</taxonomicName>
such denticles are present only on cerci (one individual examined); in 
<taxonomicName id="08B526B34F21FF85250509EDFB29EBE3" box="[1060,1222,400,424]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F21FF85250509EDFB29EBE3" box="[1060,1222,400,424]" italics="true" pageId="9" pageNumber="144">Ch. carinatum</emphasis>
</taxonomicName>
large denticles are absent (one individual examined).
</paragraph>
<paragraph id="CF0A5D304F21FF8521E609AAFBBBE8EA" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
<emphasis id="FDC181224F21FF8521E609AAFDA5EBBB" bold="true" box="[199,586,471,496]" pageId="9" pageNumber="144">(22) Pose of developing gonostyli</emphasis>
(
<figureCitation id="578E41B54F21FF85237909AAFD35EBBB" box="[600,730,471,496]" captionStart="FIGURES 95 – 98. 95 – 96" captionStartId="21.[151,264,1779,1802]" captionTargetBox="[254,1331,471,1750]" captionTargetId="figure@21.[240,1347,467,1758]" captionTargetPageId="21" captionText="FIGURES 95 – 98. 95 – 96, Cheleocloeon clavifolium sp. n.: 95, larval protogonostyli and subimaginal gonostyli developing under larval cuticle; 96, subimaginal gonostyli of mature larva. 97 – 98, Cheleocloeon sp. 1: 97, subimaginal exuviae of genitals, 98, genitals of male imago (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles and gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/270606/files/figure.png" pageId="9" pageNumber="144">Figs 95, 96</figureCitation>
). Pose of developing subimaginal gonostyli folded under larval cuticle is of the « 
<taxonomicName id="08B526B34F21FF85204A0986FE53E858" box="[363,444,507,531]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF85204A0986FE53E858" box="[363,444,507,531]" italics="true" pageId="9" pageNumber="144">Cloeon</emphasis>
</taxonomicName>
- 
<typeStatus id="100EE3924F21FF8520E50986FE16E858" box="[452,505,507,531]" pageId="9" pageNumber="144">type</typeStatus>
», i.e. with 2nd segments diverging laterally. This is a plesiomorphic character, common with most Protopatellata, Cloeonini, 
<taxonomicName id="08B526B34F21FF8523FF0A63FB34E87D" authority="Baetopus and Afrobaetodes" authorityName="Baetopus and Afrobaetodes" box="[734,1243,541,566]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8523FF0A63FC9DE87D" box="[734,882,542,566]" italics="true" pageId="9" pageNumber="144">Centroptilum</emphasis>
, 
<emphasis id="FDC181224F21FF8522AC0A62FC1AE87D" box="[909,1013,543,566]" italics="true" pageId="9" pageNumber="144">Baetopus</emphasis>
and 
<emphasis id="FDC181224F21FF8525660A63FB34E87D" box="[1095,1243,542,566]" italics="true" pageId="9" pageNumber="144">Afrobaetodes</emphasis>
</taxonomicName>
; but it allows 
<taxonomicName id="08B526B34F21FF8521B60A3CFEDEE812" box="[151,305,577,601]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8521B60A3CFEDEE812" box="[151,305,577,601]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
to be distinguished from 
<taxonomicName id="08B526B34F21FF8523470A3CFD30E812" box="[614,735,577,601]" class="Insecta" family="Baetidae" genus="Bugilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8523470A3CFD30E812" box="[614,735,577,601]" italics="true" pageId="9" pageNumber="144">Bugilliesia</emphasis>
</taxonomicName>
and other Rhithrocloeoninae, whose gonostyli have a quite different pose (
<bibRefCitation id="AB2420C14F21FF8520620A18FE35E835" author="Kluge" box="[323,474,613,638]" pageId="9" pageNumber="167" refString="Kluge, N. J. (2012 b) Systematics of Rhithrocloeninae with new species from Uganda. Russian Entomological Journal, 21 (1), 1 - 13." type="journal article" year="2012" yearSuffix="b">Kluge 2012b</bibRefCitation>
, 
<bibRefCitation id="AB2420C14F21FF8520C60A18FDCBE835" author="Kluge" box="[487,548,613,638]" pageId="9" pageNumber="167" refString="Kluge, N. J. (2015) First description of winged stages of Thraulobaetodes Elouard &amp; Hideux 1991 and reclassification of Rhithrocloeoninae (Ephemeroptera, Baetidae). Zootaxa, 3949 (4), 491 - 514. http: // dx. doi. org / 10.11646 / zootaxa. 3949.4.2" type="journal article" year="2015">2015</bibRefCitation>
). The pose of developing gonostyli is examined for 
<taxonomicName id="08B526B34F21FF8525590A18FAEDE836" box="[1144,1282,613,637]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F21FF8525590A18FAEDE836" box="[1144,1282,613,637]" italics="true" pageId="9" pageNumber="144">Ch. excisum</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F21FF8524330A18FA7AE836" box="[1298,1429,613,637]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F21FF8524330A18FA7AE836" box="[1298,1429,613,637]" italics="true" pageId="9" pageNumber="144">Ch. soldani</emphasis>
</taxonomicName>
, 
<taxonomicName id="08B526B34F21FF8521B60AF5FEACE8EB" box="[151,323,648,672]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8521B60AF5FEACE8EB" box="[151,323,648,672]" italics="true" pageId="9" pageNumber="144">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF85206B0AF5FE65E8EA" box="[330,394,648,673]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF85206B0AF5FE65E8EA" bold="true" box="[330,394,648,673]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F21FF8520B60AF5FD8FE8EB" box="[407,608,648,672]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8520B60AF5FD8FE8EB" box="[407,608,648,672]" italics="true" pageId="9" pageNumber="144">Ch. lancetofolium</emphasis>
</taxonomicName>
<emphasis id="FDC181224F21FF8523460AF5FD48E8EA" bold="true" box="[615,679,648,673]" pageId="9" pageNumber="144">
<taxonomicNameLabel id="E6F23C594F21FF8523460AF5FD48E8EA" box="[615,679,648,673]" pageId="9" pageNumber="144" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="08B526B34F21FF8523950AF5FC82E8EB" box="[692,877,648,672]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8523950AF5FC82E8EB" box="[692,877,648,672]" italics="true" pageId="9" pageNumber="144">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF8522550AF5FC5BE8EA" box="[884,948,648,673]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF8522550AF5FC5BE8EA" bold="true" box="[884,948,648,673]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<emphasis id="FDC181224F21FF8522CB0AF5FBFDE8EB" box="[1002,1042,648,672]" italics="true" pageId="9" pageNumber="144">Ch.</emphasis>
sp. 2.
</paragraph>
<paragraph id="CF0A5D304F21FF8521E60AD1FCCCEE93" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
<emphasis id="FDC181224F21FF8521E60AD1FDFCE88E" bold="true" box="[199,531,684,709]" pageId="9" pageNumber="144">(23) Genitals of male imago</emphasis>
(
<figureCitation id="578E41B54F21FF8523020AD1FD5BE88F" box="[547,692,684,709]" captionStart-0="FIGURES 95 – 98. 95 – 96" captionStart-1="FIGURES 99 – 102" captionStart-2="FIGURES 103 – 106" captionStart-3="FIGURES 107 – 112. 107 – 109" captionStartId-0="21.[151,264,1779,1802]" captionStartId-1="22.[151,264,1439,1462]" captionStartId-2="23.[151,264,1757,1780]" captionStartId-3="24.[151,264,1860,1883]" captionTargetBox-0="[254,1331,471,1750]" captionTargetBox-1="[162,1435,193,1418]" captionTargetBox-2="[227,1341,194,1740]" captionTargetBox-3="[177,1387,202,1819]" captionTargetId-0="figure@21.[240,1347,467,1758]" captionTargetId-1="figure@22.[151,1436,193,1419]" captionTargetId-2="figure@23.[220,1347,193,1741]" captionTargetId-3="figure@24.[177,1400,193,1824]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="FIGURES 95 – 98. 95 – 96, Cheleocloeon clavifolium sp. n.: 95, larval protogonostyli and subimaginal gonostyli developing under larval cuticle; 96, subimaginal gonostyli of mature larva. 97 – 98, Cheleocloeon sp. 1: 97, subimaginal exuviae of genitals, 98, genitals of male imago (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles and gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." captionText-1="FIGURES 99 – 102. Genitals of Cheleocloeon clavifolium sp. n. 99 – 100, cuticle treated with alkali, ventral view: 99, with focus on gonovectes; 100, with focus on unistyligers. 101, cuticle of left half of penis treated by alkali; 102, left half of penis with muscles and gonoduct. Abbreviations: ap, apodeme served for attachment of gonovectal muscle; gd, gonoduct; gv, gonovectis; pb, penial bridge." captionText-2="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." captionText-3="FIGURES 107 – 112. 107 – 109, Cheleocloeon lancetofolium sp. n.; 110 – 112, Ch. truncifolium sp. n. 107, 111, subimaginal exuviae of genitals; 108, 110, imaginal genitals, ventral view (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines; see Fig. 105); 109, 112, hind wing." httpUri-0="https://zenodo.org/record/270606/files/figure.png" httpUri-1="https://zenodo.org/record/270607/files/figure.png" httpUri-2="https://zenodo.org/record/270608/files/figure.png" httpUri-3="https://zenodo.org/record/270609/files/figure.png" pageId="9" pageNumber="144">Figs 98–110</figureCitation>
). Gonovectes are movable, wide, well-sclerotized and pigmented, with distal margin strengthened; apex of each gonovectis (where gonoduct is attached), being directed medially, is pointed and hooked so that its point is directed cranially (i.e. into the body). Protuberance at mid-length of gonovectis, served for attachment of gonovectal muscle, is prominent, located at the same plane as the hooked apex and has the same direction (cranially, i.e., into the body) (
<figureCitation id="578E41B54F21FF8522130B44FC79E919" box="[818,918,825,850]" captionStart="FIGURES 99 – 102" captionStartId="22.[151,264,1439,1462]" captionTargetBox="[162,1435,193,1418]" captionTargetId="figure@22.[151,1436,193,1419]" captionTargetPageId="22" captionText="FIGURES 99 – 102. Genitals of Cheleocloeon clavifolium sp. n. 99 – 100, cuticle treated with alkali, ventral view: 99, with focus on gonovectes; 100, with focus on unistyligers. 101, cuticle of left half of penis treated by alkali; 102, left half of penis with muscles and gonoduct. Abbreviations: ap, apodeme served for attachment of gonovectal muscle; gd, gonoduct; gv, gonovectis; pb, penial bridge." httpUri="https://zenodo.org/record/270607/files/figure.png" pageId="9" pageNumber="144">Fig. 101</figureCitation>
). When gonovectes are turned caudally, their hooked apices are visible between unistyligers (
<figureCitation id="578E41B54F21FF85239C0B20FCF9E93D" box="[701,790,861,886]" captionStart="FIGURES 99 – 102" captionStartId="22.[151,264,1439,1462]" captionTargetBox="[162,1435,193,1418]" captionTargetId="figure@22.[151,1436,193,1419]" captionTargetPageId="22" captionText="FIGURES 99 – 102. Genitals of Cheleocloeon clavifolium sp. n. 99 – 100, cuticle treated with alkali, ventral view: 99, with focus on gonovectes; 100, with focus on unistyligers. 101, cuticle of left half of penis treated by alkali; 102, left half of penis with muscles and gonoduct. Abbreviations: ap, apodeme served for attachment of gonovectal muscle; gd, gonoduct; gv, gonovectis; pb, penial bridge." httpUri="https://zenodo.org/record/270607/files/figure.png" pageId="9" pageNumber="144">Figs 99</figureCitation>
, 
<figureCitation id="578E41B54F21FF8522040B20FCBCE93D" box="[805,851,861,886]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="9" pageNumber="144">106</figureCitation>
); when they are turned cranially, they are invisible from ventral view (
<figureCitation id="578E41B54F21FF8520520BFDFE38E9D2" box="[371,471,896,921]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="9" pageNumber="144">Fig. 105</figureCitation>
). Penial bridge is slightly sclerotized in lateral parts and membranous medially (
<figureCitation id="578E41B54F21FF85244B0BFDFF2CE9F6" captionStart="FIGURES 99 – 102" captionStartId="22.[151,264,1439,1462]" captionTargetBox="[162,1435,193,1418]" captionTargetId="figure@22.[151,1436,193,1419]" captionTargetPageId="22" captionText="FIGURES 99 – 102. Genitals of Cheleocloeon clavifolium sp. n. 99 – 100, cuticle treated with alkali, ventral view: 99, with focus on gonovectes; 100, with focus on unistyligers. 101, cuticle of left half of penis treated by alkali; 102, left half of penis with muscles and gonoduct. Abbreviations: ap, apodeme served for attachment of gonovectal muscle; gd, gonoduct; gv, gonovectis; pb, penial bridge." httpUri="https://zenodo.org/record/270607/files/figure.png" pageId="9" pageNumber="144">Figs 101</figureCitation>
, 
<figureCitation id="578E41B54F21FF8521EF0BD8FF13E9F6" box="[206,252,933,957]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="9" pageNumber="144">104</figureCitation>
). This structure of gonovectes and penial bridge is uniform in all species examined (
<taxonomicName id="08B526B34F21FF8525850BD9FABFE9F7" box="[1188,1360,932,956]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8525850BD9FABFE9F7" box="[1188,1360,932,956]" italics="true" pageId="9" pageNumber="144">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF8524770BD9FA7AE9F6" box="[1366,1429,932,957]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF8524770BD9FA7AE9F6" bold="true" box="[1366,1429,932,957]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F21FF8521B60BB5FE8EE9AB" box="[151,353,968,992]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8521B60BB5FE8EE9AB" box="[151,353,968,992]" italics="true" pageId="9" pageNumber="144">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF8520480BBAFE45E9AB" box="[361,426,967,992]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF8520480BBAFE45E9AB" bold="true" box="[361,426,967,992]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F21FF8520980BB5FD9CE9AB" box="[441,627,968,992]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8520980BB5FD9CE9AB" box="[441,627,968,992]" italics="true" pageId="9" pageNumber="144">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF85235D0BBAFD53E9AB" box="[636,700,967,992]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF85235D0BBAFD53E9AB" bold="true" box="[636,700,967,992]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<emphasis id="FDC181224F21FF8523D40BB5FCF9E9AB" box="[757,790,968,992]" italics="true" pageId="9" pageNumber="144">Ch</emphasis>
. sp. 1). Unpaired styligeral muscle is well developed, so that unistyligers can be brought together (
<figureCitation id="578E41B54F21FF8523490B96FD28EE4F" box="[616,711,1003,1028]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="9" pageNumber="144">Fig. 105</figureCitation>
) and turned apart (
<figureCitation id="578E41B54F21FF8522BB0B96FC13EE4F" box="[922,1020,1003,1028]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="9" pageNumber="144">Fig. 106</figureCitation>
). Median sclerite served for insertion of the styligeral muscle, is either present (
<figureCitation id="578E41B54F21FF85234D0C73FD23EE6C" box="[620,716,1038,1063]" captionStart="FIGURES 107 – 112. 107 – 109" captionStartId="24.[151,264,1860,1883]" captionTargetBox="[177,1387,202,1819]" captionTargetId="figure@24.[177,1400,193,1824]" captionTargetPageId="24" captionText="FIGURES 107 – 112. 107 – 109, Cheleocloeon lancetofolium sp. n.; 110 – 112, Ch. truncifolium sp. n. 107, 111, subimaginal exuviae of genitals; 108, 110, imaginal genitals, ventral view (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines; see Fig. 105); 109, 112, hind wing." httpUri="https://zenodo.org/record/270609/files/figure.png" pageId="9" pageNumber="144">Fig. 110</figureCitation>
), or not (such sclerite occurs also in some Protopatellata). Bases of unistyligers are either contiguous (
<figureCitation id="578E41B54F21FF85231F0C4FFD4BEE00" box="[574,676,1074,1099]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="9" pageNumber="144">Figs 106</figureCitation>
, 
<figureCitation id="578E41B54F21FF8523900C4FFD31EE01" box="[689,734,1074,1098]" captionStart="FIGURES 107 – 112. 107 – 109" captionStartId="24.[151,264,1860,1883]" captionTargetBox="[177,1387,202,1819]" captionTargetId="figure@24.[177,1400,193,1824]" captionTargetPageId="24" captionText="FIGURES 107 – 112. 107 – 109, Cheleocloeon lancetofolium sp. n.; 110 – 112, Ch. truncifolium sp. n. 107, 111, subimaginal exuviae of genitals; 108, 110, imaginal genitals, ventral view (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines; see Fig. 105); 109, 112, hind wing." httpUri="https://zenodo.org/record/270609/files/figure.png" pageId="9" pageNumber="144">108</figureCitation>
), or narrowly separated (
<figureCitation id="578E41B54F21FF8522DA0C4FFBA0EE01" box="[1019,1103,1074,1099]" captionStart="FIGURES 95 – 98. 95 – 96" captionStartId="21.[151,264,1779,1802]" captionTargetBox="[254,1331,471,1750]" captionTargetId="figure@21.[240,1347,467,1758]" captionTargetPageId="21" captionText="FIGURES 95 – 98. 95 – 96, Cheleocloeon clavifolium sp. n.: 95, larval protogonostyli and subimaginal gonostyli developing under larval cuticle; 96, subimaginal gonostyli of mature larva. 97 – 98, Cheleocloeon sp. 1: 97, subimaginal exuviae of genitals, 98, genitals of male imago (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles and gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/270606/files/figure.png" pageId="9" pageNumber="144">Fig. 98</figureCitation>
), but never widely separated. Gonostylus has 2nd segment widened apically and 3rd segment small and clavate (this is similar to Cloeonini, 
<taxonomicName id="08B526B34F21FF8521B60C04FEC4EEDA" box="[151,299,1145,1169]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8521B60C04FEC4EEDA" box="[151,299,1145,1169]" italics="true" pageId="9" pageNumber="144">Centroptilum</emphasis>
</taxonomicName>
and 
<taxonomicName id="08B526B34F21FF8520460C04FE14EEDA" box="[359,507,1145,1169]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8520460C04FE14EEDA" box="[359,507,1145,1169]" italics="true" pageId="9" pageNumber="144">Anafroptilum</emphasis>
</taxonomicName>
). Shape of 1st segment of gonostylus is species-specific; in certain species (
<taxonomicName id="08B526B34F21FF8524550C04FEFAEEFE" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F21FF8524550C04FEFAEEFE" italics="true" pageId="9" pageNumber="144">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F21FF85203C0CE1FEB1EEFE" box="[285,350,1180,1205]" pageId="9" pageNumber="144" rank="species">
<emphasis id="FDC181224F21FF85203C0CE1FEB1EEFE" bold="true" box="[285,350,1180,1205]" pageId="9" pageNumber="144">sp. n.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="08B526B34F21FF8520B70CE0FDF6EEFE" box="[406,537,1181,1205]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F21FF8520B70CE0FDF6EEFE" box="[406,537,1181,1205]" italics="true" pageId="9" pageNumber="144">Ch. soldani</emphasis>
</taxonomicName>
) it has a sclerotized and pigmented concavity on median side close to the base (
<figureCitation id="578E41B54F21FF8521BE0CC2FEC2EE93" box="[159,301,1215,1240]" captionStart="FIGURES 99 – 102" captionStartId="22.[151,264,1439,1462]" captionTargetBox="[162,1435,193,1418]" captionTargetId="figure@22.[151,1436,193,1419]" captionTargetPageId="22" captionText="FIGURES 99 – 102. Genitals of Cheleocloeon clavifolium sp. n. 99 – 100, cuticle treated with alkali, ventral view: 99, with focus on gonovectes; 100, with focus on unistyligers. 101, cuticle of left half of penis treated by alkali; 102, left half of penis with muscles and gonoduct. Abbreviations: ap, apodeme served for attachment of gonovectal muscle; gd, gonoduct; gv, gonovectis; pb, penial bridge." httpUri="https://zenodo.org/record/270607/files/figure.png" pageId="9" pageNumber="144">Figs 99, 100</figureCitation>
, 
<figureCitation id="578E41B54F21FF8520180CC2FE70EE93" box="[313,415,1215,1240]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="9" pageNumber="144">104–106</figureCitation>
; Gattolliat &amp; Sartori 2008: 
<figureCitation id="578E41B54F21FF8523F30CC2FCFAEE93" box="[722,789,1215,1240]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="9" pageNumber="144">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="CF0A5D304F21FF8521E60C9EFC98EF08" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
Genitals of the 
<typeStatus id="100EE3924F21FF8520520C99FE4CEEB7" box="[371,419,1252,1276]" pageId="9" pageNumber="144">type</typeStatus>
species, 
<taxonomicName id="08B526B34F21FF8523280C9EFD70EEB0" box="[521,671,1251,1275]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F21FF8523280C9EFD70EEB0" box="[521,671,1251,1275]" italics="true" pageId="9" pageNumber="144">Ch. yolandae</emphasis>
</taxonomicName>
, have not been examined by me, and structure of their internal parts (gonovectes and penial bridge) is not described; shape of their external parts (gonostyli and unistyligers) well agree with the description given above (
<bibRefCitation id="AB2420C14F21FF8523320D57FC85EF08" author="Wuillot" box="[531,874,1322,1347]" pageId="9" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">
Wuillot &amp; Gillies 1993: 
<figureCitation id="578E41B54F21FF8522040D57FC85EF08" box="[805,874,1322,1347]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="9" pageNumber="144">Fig. 2</figureCitation>
</bibRefCitation>
).
</paragraph>
<paragraph id="CF0A5D304F21FF8521E60D32FCDDEFBF" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
In contrast to this, genitals of the species described as 
<taxonomicName id="08B526B34F21FF8522130D33FB2DEF2D" authority="Gillies 2001" authorityName="Gillies" authorityYear="2001" box="[818,1218,1357,1382]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="sigiense">
<emphasis id="FDC181224F21FF8522130D33FBC0EF2D" box="[818,1071,1358,1382]" italics="true" pageId="9" pageNumber="144">Cheleocloeon sigiense</emphasis>
<bibRefCitation id="AB2420C14F21FF8525160D30FB2DEF2D" author="Gillies" box="[1079,1218,1357,1382]" pageId="9" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">Gillies 2001</bibRefCitation>
</taxonomicName>
and genitals of the species originally described as 
<taxonomicName id="08B526B34F21FF8520D60D0CFC94EFC1" authority="Crass 1947" authorityName="Crass" authorityYear="1947" box="[503,891,1393,1418]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="falcatum">
<emphasis id="FDC181224F21FF8520D60D0CFD1AEFC2" box="[503,757,1393,1417]" italics="true" pageId="9" pageNumber="144">Centroptilum falcatum</emphasis>
<bibRefCitation id="AB2420C14F21FF8523DC0D0FFC94EFC1" author="Crass" box="[765,891,1394,1418]" pageId="9" pageNumber="167" refString="Crass, R. S. (1947) The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum, 11 (1), 37 - 100." type="journal article" year="1947">Crass 1947</bibRefCitation>
</taxonomicName>
and subsequently attributed to 
<taxonomicName id="08B526B34F21FF8525FF0D0CFA97EFC2" box="[1246,1400,1393,1417]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8525FF0D0CFA97EFC2" box="[1246,1400,1393,1417]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
by 
<bibRefCitation id="AB2420C14F21FF8521B60DE9FDCCEFE6" author="Lugo-Ortiz" box="[151,547,1428,1453]" pageId="9" pageNumber="167" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) Cheleocloeon falcatum (Crass), a new combination for a southern African species previously assigned to Afroptilum Gillies (Ephemeroptera: Baetidae). African Entomology, 6 (2), 379 - 380." type="journal article" year="1998">Lugo-Ortiz and McCafferty (1998)</bibRefCitation>
, differ from genitals of 
<taxonomicName id="08B526B34F21FF8522130DE8FC23EFE6" box="[818,972,1429,1453]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8522130DE8FC23EFE6" box="[818,972,1429,1453]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
by widely separated unistyligers (
<bibRefCitation id="AB2420C14F21FF8524710DE9FF3CEF9B" author="Gillies" pageId="9" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">Gillies 2001</bibRefCitation>
: 
<figureCitation id="578E41B54F21FF8521FF0DC5FECDEF9B" box="[222,290,1464,1489]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="9" pageNumber="144">Fig. 3</figureCitation>
; 
<bibRefCitation id="AB2420C14F21FF85200E0DC5FE5FEF9B" author="Crass" box="[303,432,1464,1488]" pageId="9" pageNumber="167" refString="Crass, R. S. (1947) The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum, 11 (1), 37 - 100." type="journal article" year="1947">Crass 1947</bibRefCitation>
: 
<figureCitation id="578E41B54F21FF85209A0DC5FDE1EF9A" box="[443,526,1464,1489]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="9" pageNumber="144">Fig. 20</figureCitation>
b). These two species do not belong to 
<taxonomicName id="08B526B34F21FF8522E20DC5FBB2EF9B" box="[963,1117,1464,1488]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8522E20DC5FBB2EF9B" box="[963,1117,1464,1488]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
: the first of them belongs to Protopatellata and the second one to Cloeonini (see below).
</paragraph>
<paragraph id="CF0A5D304F21FF8521E60D82FCF6ED78" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
Structure of gonovectes is unique for 
<taxonomicName id="08B526B34F21FF8523490D82FCEDEC5C" box="[616,770,1535,1559]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8523490D82FCEDEC5C" box="[616,770,1535,1559]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
, not found in any other taxa. In all Cloeonini, 
<taxonomicName id="08B526B34F21FF8524240E7DFA82EC5C" box="[1285,1389,1536,1559]" class="Insecta" family="Baetidae" genus="Baetopus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8524240E7DFA82EC5C" box="[1285,1389,1536,1559]" italics="true" pageId="9" pageNumber="144">Baetopus</emphasis>
</taxonomicName>
and 
<taxonomicName id="08B526B34F21FF8521B60E5EFEC4EC70" box="[151,299,1571,1595]" class="Insecta" family="Baetidae" genus="Anafroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8521B60E5EFEC4EC70" box="[151,299,1571,1595]" italics="true" pageId="9" pageNumber="144">Anafroptilum</emphasis>
</taxonomicName>
, gonovectes are immovably fused by their apices with penial bridge (e.g., 
<bibRefCitation id="AB2420C14F21FF8525BA0E5FFADCEC70" author="Kluge" box="[1179,1331,1570,1595]" pageId="9" pageNumber="167" refString="Kluge, N. J. (2012 a) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 2011, 20 (4), 361 - 376." type="journal article" year="2012" yearSuffix="a">Kluge 2012a</bibRefCitation>
: 
<figureCitation id="578E41B54F21FF8524600E5FFA78EC70" box="[1345,1431,1570,1595]" captionStart="FIGURES 30 – 57" captionStartId="8.[151,264,1554,1577]" captionTargetBox="[153,1432,201,1529]" captionTargetId="figure@8.[151,1436,193,1533]" captionTargetPageId="8" captionText="FIGURES 30 – 57. Tergalii I – VII: 30 – 36, Cheleocloeon clavifolium sp. n.; 37 – 43, Ch. lancetofolium sp. n.; 44 – 50, Ch. truncifolium sp. n. (male, holotype); 51 – 57, Ch. truncifolium sp. n. (female)." httpUri="https://zenodo.org/record/270600/files/figure.png" pageId="9" pageNumber="144">Fig. 45</figureCitation>
; 
<bibRefCitation id="AB2420C14F21FF8521B60E3BFDF2EC14" author="Kluge" box="[151,541,1606,1631]" pageId="9" pageNumber="167" refString="Kluge, N. J., Tiunova, T. M. &amp; Novikova, E. A. (2014) A new species, Procloeon monilistylus sp. n. (Ephemeroptera, Baetidae), from the Russian Far East. Zootaxa, 3786 (4), 483 - 491. http: // dx. doi. org / 10.11646 / zootaxa. 3786.4.6" type="journal article" year="2014">Kluge, Tiunova &amp; Novikova 2014</bibRefCitation>
: 
<figureCitation id="578E41B54F21FF8523080E3BFD45EC14" box="[553,682,1606,1631]" captionStart="FIGURES 30 – 57" captionStartId="8.[151,264,1554,1577]" captionTargetBox="[153,1432,201,1529]" captionTargetId="figure@8.[151,1436,193,1533]" captionTargetPageId="8" captionText="FIGURES 30 – 57. Tergalii I – VII: 30 – 36, Cheleocloeon clavifolium sp. n.; 37 – 43, Ch. lancetofolium sp. n.; 44 – 50, Ch. truncifolium sp. n. (male, holotype); 51 – 57, Ch. truncifolium sp. n. (female)." httpUri="https://zenodo.org/record/270600/files/figure.png" pageId="9" pageNumber="144">Figs 30, 31</figureCitation>
). In 
<taxonomicName id="08B526B34F21FF8523FF0E3BFC9DEC15" box="[734,882,1606,1630]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8523FF0E3BFC9DEC15" box="[734,882,1606,1630]" italics="true" pageId="9" pageNumber="144">Centroptilum</emphasis>
</taxonomicName>
gonovectes are movable, pointed and hooked; in contrast to 
<taxonomicName id="08B526B34F21FF8520030E14FE53ECCA" box="[290,444,1641,1665]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8520030E14FE53ECCA" box="[290,444,1641,1665]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
, their points are directed not cranially, but caudally (
<bibRefCitation id="AB2420C14F21FF85256C0E14FA85ECC9" author="Grandi" box="[1101,1386,1641,1666]" pageId="9" pageNumber="167" refString="Grandi, M. (1960) Contributi allo studio degli Efemeroidei italiani. XXIII. Gli organi genitali esterni maschili degli Efemeroidei. Bollettino dell'Istituto di Entomologia della Universita di Bologna, 24, 67 - 120." type="journal article" year="1960">
Grandi 1960: Fig. 
<date id="BB0B7BF04F21FF8524150E17FA85ECC9" box="[1332,1386,1642,1666]" pageId="9" pageNumber="144">XI.3</date>
</bibRefCitation>
). In 
<taxonomicName id="08B526B34F21FF8521B60EF0FEC3ECEE" box="[151,300,1677,1701]" class="Insecta" family="Baetidae" genus="Afrobaetodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8521B60EF0FEC3ECEE" box="[151,300,1677,1701]" italics="true" pageId="9" pageNumber="144">Afrobaetodes</emphasis>
</taxonomicName>
gonovectes are also movable and hooked; in contrast to 
<taxonomicName id="08B526B34F21FF8522830EF0FBD3ECEE" box="[930,1084,1677,1701]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8522830EF0FBD3ECEE" box="[930,1084,1677,1701]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
, the protuberance which serves for muscle attachment and the hooked apex are directed to opposite sides (unpublished data). In all Baetovectata gonovectes are also movable, but slender, sharply bent and deeply evaginated into the body. In various taxa of Protopatellata, gonovectes are either fused with penial bridge (e.g. in Rhithrocloeoninae, 
<taxonomicName id="08B526B34F21FF85255D0E8AFB15ED44" box="[1148,1274,1783,1807]" class="Insecta" family="Baetidae" genus="Indocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF85255D0E8AFB15ED44" box="[1148,1274,1783,1807]" italics="true" pageId="9" pageNumber="144">Indocloeon</emphasis>
</taxonomicName>
and others) or movable, but always different from that of 
<taxonomicName id="08B526B34F21FF8523590F66FCFDED78" box="[632,786,1819,1843]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F21FF8523590F66FCFDED78" box="[632,786,1819,1843]" italics="true" pageId="9" pageNumber="144">Cheleocloeon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F21FF8521E60F43FE35ED31" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
<emphasis id="FDC181224F21FF8521E60F43FED3ED1C" bold="true" box="[199,316,1854,1879]" pageId="9" pageNumber="144">(24) Eggs</emphasis>
(
<figureCitation id="578E41B54F21FF8520710F43FE1AED1C" box="[336,501,1854,1879]" captionStart-0="FIGURES 122 – 127" captionStart-1="FIGURES 128 – 133" captionStartId-0="28.[151,264,1786,1809]" captionStartId-1="30.[151,264,1616,1639]" captionTargetBox-0="[150,1435,303,1764]" captionTargetBox-1="[177,1402,195,1589]" captionTargetId-0="figure@28.[150,1436,303,1764]" captionTargetId-1="figure@30.[177,1402,195,1589]" captionTargetPageId-0="28" captionTargetPageId-1="30" captionText-0="FIGURES 122 – 127. Eggs. 122 – 123, Cheleocloeon yolandae; 124 – 125, Cheleocloeon clavifolium sp. n., 126 – 127, Cheleocloeon lancetofolium sp. n." captionText-1="FIGURES 128 – 133. Eggs. 128 – 131, Cheleocloeon truncifolium sp. n.; 132 – 133, Cheleocloeon sp. 2." httpUri-0="https://zenodo.org/record/270613/files/figure.png" httpUri-1="https://zenodo.org/record/270614/files/figure.png" pageId="9" pageNumber="144">Figs 122–131</figureCitation>
). Eggs are oval, with more or less expressed convex net-like relief, somewhat different in different species.
</paragraph>
</subSubSection>
<subSubSection id="87AF0EBB4F21FF8621E60FF4FD49EE5F" lastPageId="10" lastPageNumber="145" pageId="9" pageNumber="144" type="distribution">
<paragraph id="CF0A5D304F21FF8521E60FF4FA66EDA3" blockId="9.[151,1437,151,2024]" pageId="9" pageNumber="144">
<emphasis id="FDC181224F21FF8521E60FF4FE8FEDE9" bold="true" box="[199,352,1929,1954]" pageId="9" pageNumber="144">Distribution.</emphasis>
Whole of Africa, 
<collectingCountry id="B7A21DA04F21FF8523100FF4FD57EDE9" box="[561,696,1929,1954]" name="Madagascar" pageId="9" pageNumber="144">Madagascar</collectingCountry>
and Arabian Peninsula. Such area of distribution is unknown for other mayfly taxa, which are either Afrotropical (often including 
<collectingCountry id="B7A21DA04F21FF85225E0FD1FBE4ED8E" box="[895,1035,1964,1989]" name="Madagascar" pageId="9" pageNumber="144">Madagascar</collectingCountry>
), or Palaearctic (including northern Africa), or widespread, including both Afrotropical and Palaearctic regions, but not limited to Africa and Arabia.
</paragraph>
<paragraph id="CF0A5D304F22FF8621E608E6FCA6EA93" blockId="10.[151,1437,155,1044]" pageId="10" pageNumber="145">
<emphasis id="FDC181224F22FF8621E608E6FE88EAFF" bold="true" box="[199,359,155,180]" pageId="10" pageNumber="145">Composition.</emphasis>
Together with three species described below, 
<taxonomicName id="08B526B34F22FF8622AC08E1FBC8EAFF" box="[909,1063,156,180]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8622AC08E1FBC8EAFF" box="[909,1063,156,180]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
contains 10 formally described species, which can be presumably arranged into three groups:
</paragraph>
<paragraph id="CF0A5D304F22FF8621E60899FB78EB6B" blockId="10.[151,1437,155,1044]" pageId="10" pageNumber="145">
A. Group 
<taxonomicName id="08B526B34F22FF8620600899FE47EAB7" box="[321,424,228,252]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F22FF8620600899FE47EAB7" box="[321,424,228,252]" italics="true" pageId="10" pageNumber="145">yolandae</emphasis>
</taxonomicName>
(hind wings absent; claws without large denticles); includes 2 species— 
<taxonomicName id="08B526B34F22FF8624230899FF03EB6B" authority="Wuillot" authorityName="Wuillot" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="yolandae">
<emphasis id="FDC181224F22FF8624230899FA73EAB7" box="[1282,1436,228,252]" italics="true" pageId="10" pageNumber="145">Ch. yolandae</emphasis>
Wuillot
</taxonomicName>
(in 
<bibRefCitation id="AB2420C14F22FF862039097AFDCBEB6B" author="Wuillot" box="[280,548,263,288]" pageId="10" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot &amp; Gillies) 1993</bibRefCitation>
and 
<taxonomicName id="08B526B34F22FF86237B097AFCB8EB6B" authority="Wuillot" authorityName="Wuillot" box="[602,855,263,288]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="carinatum">
<emphasis id="FDC181224F22FF86237B097AFD13EB54" box="[602,764,263,287]" italics="true" pageId="10" pageNumber="145">Ch. carinatum</emphasis>
Wuillot
</taxonomicName>
(in 
<bibRefCitation id="AB2420C14F22FF8622A5097AFB7DEB6B" author="Wuillot" box="[900,1170,263,288]" pageId="10" pageNumber="167" refString="Wuillot, J. &amp; Gillies, M. T. (1993) Cheleocloeon, a new genus of Baetidae (Ephemeroptera) from West Africa. Revue d'Hydrobiologie Tropicale, 26 (3), 213 - 217." type="journal article" year="1993">Wuillot &amp; Gillies) 1993</bibRefCitation>
.
</paragraph>
<paragraph id="CF0A5D304F22FF8621E60951FB31EBC7" blockId="10.[151,1437,155,1044]" pageId="10" pageNumber="145">
B. Group 
<taxonomicName id="08B526B34F22FF86201D0951FE55EB0F" box="[316,442,300,324]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="clavifolium">
<emphasis id="FDC181224F22FF86201D0951FE55EB0F" box="[316,442,300,324]" italics="true" pageId="10" pageNumber="145">clavifolium</emphasis>
</taxonomicName>
(hind wings present in male; claws without large denticles); includes 5 species:— 
<taxonomicName id="08B526B34F22FF8624550951FEFAEB2C" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="clavifolium" status="sp. nov.">
<emphasis id="FDC181224F22FF8624550951FEFAEB2C" italics="true" pageId="10" pageNumber="145">Ch. clavifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F22FF86203D0932FEB2EB23" box="[284,349,335,360]" pageId="10" pageNumber="145" rank="species">
<emphasis id="FDC181224F22FF86203D0932FEB2EB23" bold="true" box="[284,349,335,360]" pageId="10" pageNumber="145">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F22FF86204A0932FDDBEB2C" box="[363,564,335,359]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="lancetofolium" status="sp. nov.">
<emphasis id="FDC181224F22FF86204A0932FDDBEB2C" box="[363,564,335,359]" italics="true" pageId="10" pageNumber="145">Ch. lancetofolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F22FF86231D0932FD93EB23" box="[572,636,335,360]" pageId="10" pageNumber="145" rank="species">
<emphasis id="FDC181224F22FF86231D0932FD93EB23" bold="true" box="[572,636,335,360]" pageId="10" pageNumber="145">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F22FF8623AB0932FCAAEB2C" box="[650,837,335,359]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="truncifolium" status="sp. nov.">
<emphasis id="FDC181224F22FF8623AB0932FCAAEB2C" box="[650,837,335,359]" italics="true" pageId="10" pageNumber="145">Ch. truncifolium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6F23C594F22FF86226D0932FC63EB23" box="[844,908,335,360]" pageId="10" pageNumber="145" rank="species">
<emphasis id="FDC181224F22FF86226D0932FC63EB23" bold="true" box="[844,908,335,360]" pageId="10" pageNumber="145">sp. n.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="08B526B34F22FF8622BB0932FA73EB23" authority="Gattolliat &amp; Monaghan" authorityName="Gattolliat &amp; Monaghan" box="[922,1436,335,360]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="madagascariense">
<emphasis id="FDC181224F22FF8622BB0932FB61EB2C" box="[922,1166,335,359]" italics="true" pageId="10" pageNumber="145">Ch. madagascariense</emphasis>
Gattolliat &amp; Monaghan
</taxonomicName>
(in 
<bibRefCitation id="AB2420C14F22FF86219C090EFD27EBC7" author="Gattolliat" box="[189,712,371,396]" pageId="10" pageNumber="167" refString="Gattolliat, J. - L., Barber-James, H. M. &amp; Monaghan, M. T. (2009) New species and generic delimitation of the Afrotropical genera Bugilliesia Lugo-Ortiz &amp; McCafferty, 1996; Cheleocloeon Wuillot &amp; Gillies, 1993 and Delouardus Lugo-Ortiz &amp; McCafferty, 1999 (Ephemeroptera: Baetidae). Aquatic Insects, 31 (3), 167 - 186. http: // dx. doi. org / 10.1080 / 01650420902782987" type="journal article" year="2009">Gattolliat &amp; Barber-James &amp; Monaghan) 2009</bibRefCitation>
and 
<taxonomicName id="08B526B34F22FF8623DF0909FB36EBC7" authority="Soldan &amp; Tomas 1985" authorityName="Soldan &amp; Tomas" authorityYear="1985" box="[766,1241,371,396]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="dimorphicum">
<emphasis id="FDC181224F22FF8623DF0909FC2DEBC7" box="[766,962,372,396]" italics="true" pageId="10" pageNumber="145">Ch. dimorphicum</emphasis>
(
<bibRefCitation id="AB2420C14F22FF8622F0090EFB3EEBC7" author="Soldan" box="[977,1233,371,396]" pageId="10" pageNumber="167" refString="Soldan, T. &amp; Thomas, A. G. B. (1985) Centroptilum dimorphicum sp. n., a new species of mayfly (Ephemeroptera, Baetidae) from Algeria. Acta Entomologica Bohemoslovaca, 82, 180 - 186." type="journal article" year="1985">Soldán &amp; Tomas 1985</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F22FF8621E609E5FA92EB9F" blockId="10.[151,1437,155,1044]" pageId="10" pageNumber="145">
C. Group 
<taxonomicName id="08B526B34F22FF86201E09E5FE76EBE4" box="[319,409,408,431]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F22FF86201E09E5FE76EBE4" box="[319,409,408,431]" italics="true" pageId="10" pageNumber="145">excisum</emphasis>
</taxonomicName>
(hind wings present in male; claws with large pointed denticles); includes 3 described species— 
<taxonomicName id="08B526B34F22FF86202509C1FD47EB9F" authority="Gattolliat &amp; Sartori 2008" authorityName="Gattolliat &amp; Sartori" authorityYear="2008" box="[260,680,443,468]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="soldani">
<emphasis id="FDC181224F22FF86202509C1FE6AEB9F" box="[260,389,444,468]" italics="true" pageId="10" pageNumber="145">Ch. soldani</emphasis>
Gattolliat &amp; Sartori 2008
</taxonomicName>
, 
<taxonomicName id="08B526B34F22FF86239209C1FC01EB9F" authority="Barnard 1932" authorityName="Barnard" authorityYear="1932" box="[691,1006,443,468]" class="Insecta" family="Baetidae" genus="Afroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="excisum">
<emphasis id="FDC181224F22FF86239209C1FCD4EB98" box="[691,827,444,468]" italics="true" pageId="10" pageNumber="145">Ch. excisum</emphasis>
(
<bibRefCitation id="AB2420C14F22FF86226B09C6FC09EB9F" author="Barnard" box="[842,998,443,468]" pageId="10" pageNumber="166" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. http: // dx. doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="08B526B34F22FF86250509C1FA96EB9F" authority="McCafferty 2001" authorityName="McCafferty" authorityYear="2001" box="[1060,1401,443,468]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="littorale">
<emphasis id="FDC181224F22FF86250509C1FB42EB9F" box="[1060,1197,444,468]" italics="true" pageId="10" pageNumber="145">Ch. littorale</emphasis>
<bibRefCitation id="AB2420C14F22FF86259509C6FA96EB9F" author="McCafferty" box="[1204,1401,443,468]" pageId="10" pageNumber="167" refString="McCafferty, W. P. (2001) New Baetidae (Insecta: Ephemeroptera) from Lake Malawi. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 2000, 136, 65 - 72." type="journal article" year="2001">McCafferty 2001</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F22FF8621E609A2FA67E857" blockId="10.[151,1437,155,1044]" pageId="10" pageNumber="145">
<emphasis id="FDC181224F22FF8621E609A2FD03EBB3" bold="true" box="[199,748,479,504]" pageId="10" pageNumber="145">
Species erroneously placed to 
<taxonomicName id="08B526B34F22FF86236909A2FD0AEBB3" box="[584,741,479,504]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF86236909A2FD0AEBB3" bold="true" box="[584,741,479,504]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
.
</emphasis>
<bibRefCitation id="AB2420C14F22FF8623DA09A2FB7CEBB3" author="Lugo-Ortiz" box="[763,1171,479,504]" pageId="10" pageNumber="167" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1997 b) New species and first reports of the genera Cheleocloeon, Dabulamanzia, and Mutelocloeon (Insecta: Ephemeroptera: Baetidae) from Madagascar. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 133 (1), 47 - 53." type="journal article" year="1997" yearSuffix="b">Lugo-Ortiz &amp; McCafferty (1997b)</bibRefCitation>
described the species 
<taxonomicName id="08B526B34F22FF8621B60A79FF13E857" box="[151,252,516,540]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="mirandei">
<emphasis id="FDC181224F22FF8621B60A79FF13E857" box="[151,252,516,540]" italics="true" pageId="10" pageNumber="145">mirandei</emphasis>
</taxonomicName>
[
<taxonomicName id="08B526B34F22FF86202D0A79FE49E857" box="[268,422,516,540]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF86202D0A79FE49E857" box="[268,422,516,540]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
] based on larvae from 
<collectingCountry id="B7A21DA04F22FF8623880A7EFCDBE857" box="[681,820,515,540]" name="Madagascar" pageId="10" pageNumber="145">Madagascar</collectingCountry>
; now attributed to 
<taxonomicName id="08B526B34F22FF8625240A79FA6CE857" authority="Gattolliat et al. 2009" authorityName="Gattolliat et al." authorityYear="2009" box="[1029,1411,515,540]" class="Insecta" family="Baetidae" genus="Bugilliesia" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8625240A79FB91E857" box="[1029,1150,516,540]" italics="true" pageId="10" pageNumber="145">Bugilliesia</emphasis>
(
<bibRefCitation id="AB2420C14F22FF8625AC0A7EFA93E857" author="Gattolliat" box="[1165,1404,515,540]" pageId="10" pageNumber="167" refString="Gattolliat, J. - L., Barber-James, H. M. &amp; Monaghan, M. T. (2009) New species and generic delimitation of the Afrotropical genera Bugilliesia Lugo-Ortiz &amp; McCafferty, 1996; Cheleocloeon Wuillot &amp; Gillies, 1993 and Delouardus Lugo-Ortiz &amp; McCafferty, 1999 (Ephemeroptera: Baetidae). Aquatic Insects, 31 (3), 167 - 186. http: // dx. doi. org / 10.1080 / 01650420902782987" type="journal article" year="2009">
Gattolliat 
<emphasis id="FDC181224F22FF8624210A79FAD7E857" box="[1280,1336,516,540]" italics="true" pageId="10" pageNumber="145">et al.</emphasis>
2009
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="CF0A5D304F22FF8621E60A5AFE9FE89B" blockId="10.[151,1437,155,1044]" pageId="10" pageNumber="145">
<bibRefCitation id="AB2420C14F22FF8621E60A5AFDD0E80B" author="Lugo-Ortiz" box="[199,575,551,576]" pageId="10" pageNumber="167" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1998) Cheleocloeon falcatum (Crass), a new combination for a southern African species previously assigned to Afroptilum Gillies (Ephemeroptera: Baetidae). African Entomology, 6 (2), 379 - 380." type="journal article" year="1998">Lugo-Ortiz &amp; McCafferty (1998)</bibRefCitation>
attributed to 
<taxonomicName id="08B526B34F22FF8623F80A5AFC9CE874" box="[729,883,551,575]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8623F80A5AFC9CE874" box="[729,883,551,575]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
the species 
<taxonomicName id="08B526B34F22FF8622DF0A5AFB09E80B" authority="Crass 1947" authorityName="Crass" authorityYear="1947" box="[1022,1254,551,576]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="falcatum">
<emphasis id="FDC181224F22FF8622DF0A5AFB8FE874" box="[1022,1120,551,575]" italics="true" pageId="10" pageNumber="145">falcatum</emphasis>
<bibRefCitation id="AB2420C14F22FF8625490A55FB09E80B" author="Crass" box="[1128,1254,552,576]" pageId="10" pageNumber="167" refString="Crass, R. S. (1947) The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum, 11 (1), 37 - 100." type="journal article" year="1947">Crass 1947</bibRefCitation>
</taxonomicName>
[
<taxonomicName id="08B526B34F22FF8625D60A5AFA64E874" box="[1271,1419,551,575]" class="Insecta" family="Baetidae" genus="Centroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8625D60A5AFA64E874" box="[1271,1419,551,575]" italics="true" pageId="10" pageNumber="145">Centroptilum</emphasis>
</taxonomicName>
], described as male imagoes only; actually, this species cannot belong to 
<taxonomicName id="08B526B34F22FF8622E20A31FBB2E82F" box="[963,1117,588,612]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8622E20A31FBB2E82F" box="[963,1117,588,612]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
, because its unistyligers are widely separated (
<bibRefCitation id="AB2420C14F22FF8620400A0DFDA3E8C3" author="Crass" box="[353,588,623,648]" pageId="10" pageNumber="167" refString="Crass, R. S. (1947) The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum, 11 (1), 37 - 100." type="journal article" year="1947" yearSuffix="b">
Crass 1947: 
<figureCitation id="578E41B54F22FF8620CF0A12FDD1E8C3" box="[494,574,623,648]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="10" pageNumber="145">Fig. 20</figureCitation>
b
</bibRefCitation>
) [see character (23) of 
<taxonomicName id="08B526B34F22FF86226F0A12FC07E8CC" box="[846,1000,623,647]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF86226F0A12FC07E8CC" box="[846,1000,623,647]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
]; most probably, these imagoes belong to Cloeonini and possibly are conspecific with larvae described as 
<taxonomicName id="08B526B34F22FF8622810AE9FA73E8E7" authority="Lugo-Ortiz 1996" authorityName="Lugo-Ortiz" authorityYear="1996" box="[928,1436,659,684]" class="Insecta" family="Baetidae" genus="Securiops" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="macafertiorum">
<emphasis id="FDC181224F22FF8622810AE9FB51E8E7" box="[928,1214,660,684]" italics="true" pageId="10" pageNumber="145">Securiops macafertiorum</emphasis>
(
<bibRefCitation id="AB2420C14F22FF8625F00AEEFA7BE8E7" author="Lugo-Ortiz" box="[1233,1428,659,684]" pageId="10" pageNumber="167" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1996) The Bugilliesia complex of African Baetidae (Ephemeroptera). Transactions of the American Entomological Society, 122 (4), 175 - 197." type="journal article" year="1996">Lugo-Ortiz 1996</bibRefCitation>
)
</taxonomicName>
(unpublished data).
</paragraph>
<paragraph id="CF0A5D304F22FF8621E60AA6FD49EE5F" blockId="10.[151,1437,155,1044]" pageId="10" pageNumber="145">
<bibRefCitation id="AB2420C14F22FF8621E60AA6FE86E8BF" author="Gillies" box="[199,361,731,756]" pageId="10" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">Gillies (2001)</bibRefCitation>
described the species 
<taxonomicName id="08B526B34F22FF8623580AA1FD3CE8B8" box="[633,723,732,755]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="sigiense">
<emphasis id="FDC181224F22FF8623580AA1FD3CE8B8" box="[633,723,732,755]" italics="true" pageId="10" pageNumber="145">sigiense</emphasis>
</taxonomicName>
[
<taxonomicName id="08B526B34F22FF8623C80AA1FC6CE8BF" box="[745,899,732,756]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8623C80AA1FC6CE8BF" box="[745,899,732,756]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
] based on imagoes and larvae of both sexes associated by rearing. This species differs from all 
<taxonomicName id="08B526B34F22FF8623F00A82FC84E95C" box="[721,875,767,791]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8623F00A82FC84E95C" box="[721,875,767,791]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
by the following characters: labrum without setae on outer surface (
<bibRefCitation id="AB2420C14F22FF86207D0B5EFDD3E977" author="Gillies" box="[348,572,803,828]" pageId="10" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">
Gillies 2001: 
<figureCitation id="578E41B54F22FF8620D90B5EFDD3E977" box="[504,572,803,828]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="10" pageNumber="145">Fig. 4</figureCitation>
</bibRefCitation>
), while labrum of 
<taxonomicName id="08B526B34F22FF86222F0B59FC47E977" box="[782,936,804,828]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF86222F0B59FC47E977" box="[782,936,804,828]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
has several setae [see character (3) and 
<figureCitation id="578E41B54F22FF8624510B5EFF4AE92B" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="10" pageNumber="145">Fig. 1</figureCitation>
]; labial palp with 2nd segment moderately projected (
<bibRefCitation id="AB2420C14F22FF8623DE0B3AFC35E92B" author="Gillies" box="[767,986,839,864]" pageId="10" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">
Gillies 2001: 
<figureCitation id="578E41B54F22FF8622B90B3AFC35E92B" box="[920,986,839,864]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="10" pageNumber="145">Fig. 7</figureCitation>
</bibRefCitation>
), while in 
<taxonomicName id="08B526B34F22FF8625710B3AFB05E914" box="[1104,1258,839,863]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8625710B3AFB05E914" box="[1104,1258,839,863]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
its projection is larger [see character (8) and 
<figureCitation id="578E41B54F22FF8620FC0B16FDCCE9CF" box="[477,547,875,900]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="10" pageNumber="145">Fig. 5</figureCitation>
]; patella-tibial suture is present on middle and hind legs only, being absent on fore legs (
<bibRefCitation id="AB2420C14F22FF8620270BF2FDFFE9E3" author="Gillies" box="[262,528,911,936]" pageId="10" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">
Gillies 2001: 
<figureCitation id="578E41B54F22FF8620BF0BF2FDFFE9E3" box="[414,528,911,936]" captionStart="FIGURES 7 – 18. 7 – 15" captionStartId="5.[151,264,1436,1459]" captionTargetBox="[226,1346,181,1410]" captionTargetId="figure@5.[226,1360,169,1415]" captionTargetPageId="5" captionText="FIGURES 7 – 18. 7 – 15, Cheleocloeon clavifolium sp. n.: 7 – 8, incisor, kinetodontium and prostheca of left and right mandibles; 9, head of female larva; 10 – 11, larval hind protopteron and imaginal hind wing of the same male individual (holotype) at the same magnification; 12 – 13, larval protopteron and imaginal vestige of hind wing (with lateral fragment of metathorax) of the same female individual at the same magnification; 14, larval paraproct; 15, scale on larval abdominal tergum. 16, Ch. lancetofolium sp. n.: incisor, kinetodontium and prostheca of right mandible; 17, Ch. truncifolium sp. n., the same; 18, Cheleocloeon sp. 3, the same. Abbreviations: in 1, in 2, in 3, in 4, denticles of incisor; in-V, ventral denticle of incisor; kd 1, kd 2, kd 3, denticles of kinetodontium." httpUri="https://zenodo.org/record/270598/files/figure.png" pageId="10" pageNumber="145">Figs 9, 10</figureCitation>
</bibRefCitation>
), while in 
<taxonomicName id="08B526B34F22FF8623A50BEDFCF1E9E3" box="[644,798,912,936]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8623A50BEDFCF1E9E3" box="[644,798,912,936]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
it is equally developed on all legs [see character (11) and 
<figureCitation id="578E41B54F22FF8621B60BCEFF04E987" box="[151,235,947,972]" captionStart="FIGURES 19 – 29. 19 – 23" captionStartId="7.[151,264,1883,1906]" captionTargetBox="[156,1432,199,1855]" captionTargetId="figure@7.[151,1436,193,1862]" captionTargetPageId="7" captionText="FIGURES 19 – 29. 19 – 23, Cheleocloeon yolandae: 19, fore leg of female imago, anterior view; 20, fore tibia and tarsus of subimago (the same specimen), anterior view; 21, fore leg of larva (the same specimen), anterior view; 22, knee area of the same fore larval leg (posterior view); 23, larval claw. 24 – 25, Ch. soldani: 24, claw; 25, larval fore leg, anterior view. 26, Cheleocloeon sp. 3, larval fore leg, anterior view. 27 – 28, Ch. clavifolium sp. n., larval claws. 29, Ch. truncifolium sp. n., larval claw." httpUri="https://zenodo.org/record/270599/files/figure.png" pageId="10" pageNumber="145">Fig. 21</figureCitation>
]; unistyligers are widely separated (
<bibRefCitation id="AB2420C14F22FF8623A20BCEFC8DE987" author="Gillies" box="[643,866,947,972]" pageId="10" pageNumber="167" refString="Gillies, M. T. (2001) New species of Baetidae (Ephemeroptera) from the Upper River Sigi, Usambara, Tanzania. Journal of Natural History, 35, 23 - 32. http: // dx. doi. org / 10.1080 / 002229301447862" type="journal article" year="2001">
Gillies 2001: 
<figureCitation id="578E41B54F22FF86223E0BCEFC8DE987" box="[799,866,947,972]" captionStart="FIGURES 1 – 6. 1 – 5" captionStartId="4.[151,264,1942,1965]" captionTargetBox="[168,1406,201,1903]" captionTargetId="figure@4.[168,1407,193,1913]" captionTargetPageId="4" captionText="FIGURES 1 – 6. 1 – 5, Cheleocloeon clavifolium sp. n.: 1, labrum (left ventral view, right dorsal view); 2, maxilla, dorsal view; 3, dentisetae and medio-dorsal setae of another specimen (holotype); 4, labium, dorsal view; 5, the same, ventral view; 6, Ch. lancetofolium sp. n. (holotype): glossa, ventral view. Abbreviations: ds 1, ds 2, ds 3, dentisetae." httpUri="https://zenodo.org/record/270597/files/figure.png" pageId="10" pageNumber="145">Fig. 3</figureCitation>
</bibRefCitation>
), while in 
<taxonomicName id="08B526B34F22FF8622FC0BC9FB98E987" box="[989,1143,948,972]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8622FC0BC9FB98E987" box="[989,1143,948,972]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
they are brought together [see character (23) and 
<figureCitation id="578E41B54F22FF8620BD0BAAFE1FE9BB" box="[412,496,983,1008]" captionStart="FIGURES 95 – 98. 95 – 96" captionStartId="21.[151,264,1779,1802]" captionTargetBox="[254,1331,471,1750]" captionTargetId="figure@21.[240,1347,467,1758]" captionTargetPageId="21" captionText="FIGURES 95 – 98. 95 – 96, Cheleocloeon clavifolium sp. n.: 95, larval protogonostyli and subimaginal gonostyli developing under larval cuticle; 96, subimaginal gonostyli of mature larva. 97 – 98, Cheleocloeon sp. 1: 97, subimaginal exuviae of genitals, 98, genitals of male imago (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; at right half gonostylar muscle shown by interrupted lines, gonovectal muscles and gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: gs 1, gs 2, gs 3, segments of gonostylus; mIX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); usg, unistyliger." httpUri="https://zenodo.org/record/270606/files/figure.png" pageId="10" pageNumber="145">Figs 98</figureCitation>
, 
<figureCitation id="578E41B54F22FF8620DD0BAAFD8FE9BB" box="[508,608,983,1008]" captionStart="FIGURES 103 – 106" captionStartId="23.[151,264,1757,1780]" captionTargetBox="[227,1341,194,1740]" captionTargetId="figure@23.[220,1347,193,1741]" captionTargetPageId="23" captionText="FIGURES 103 – 106. Genitals of Cheleocloeon clavifolium sp. n. 103, subimaginal exuviae; 104, imaginal genitals, posterior view; 105, the same, ventral view with unistyligers and gonostyli brought together and gonovectes turned cranially (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines); 106, the same with unistyligers and gonostyli turned apart and gonovectes turned caudally (hidden parts of penis shown by interrupted lines, gonoducts shown by dotted lines). Abbreviations: gd, gonoduct; gs 1, gs 2, gs 3, segments of gonostylus; gv, gonovectis; mIX-X, areas of attachment of paraproctal muscles; pb, penial bridge; usg, unistyliger." httpUri="https://zenodo.org/record/270608/files/figure.png" pageId="10" pageNumber="145">104–106</figureCitation>
, 
<figureCitation id="578E41B54F22FF86234A0BA5FD3EE9BB" box="[619,721,984,1008]" captionStart="FIGURES 107 – 112. 107 – 109" captionStartId="24.[151,264,1860,1883]" captionTargetBox="[177,1387,202,1819]" captionTargetId="figure@24.[177,1400,193,1824]" captionTargetPageId="24" captionText="FIGURES 107 – 112. 107 – 109, Cheleocloeon lancetofolium sp. n.; 110 – 112, Ch. truncifolium sp. n. 107, 111, subimaginal exuviae of genitals; 108, 110, imaginal genitals, ventral view (on the left half, gonovectal muscles shown by interrupted lines, gonovectis shown by interrupted line and stippled, gonoduct shown by stippled line, gonostylar muscle and external pigmentation not shown; on the right half, gonostylar and gonovectal muscles shown by interrupted lines, gonoduct not shown, external pigmentation shown by stippling; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by stippled lines; see Fig. 105); 109, 112, hind wing." httpUri="https://zenodo.org/record/270609/files/figure.png" pageId="10" pageNumber="145">108, 110</figureCitation>
]. The species 
<taxonomicName id="08B526B34F22FF86224D0BA5FC29E9A4" box="[876,966,984,1007]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="species" species="sigiense">
<emphasis id="FDC181224F22FF86224D0BA5FC29E9A4" box="[876,966,984,1007]" italics="true" pageId="10" pageNumber="145">sigiense</emphasis>
</taxonomicName>
[
<taxonomicName id="08B526B34F22FF8622F70BA5FB9FE9BB" box="[982,1136,984,1008]" class="Insecta" family="Baetidae" genus="Cheleocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="FDC181224F22FF8622F70BA5FB9FE9BB" box="[982,1136,984,1008]" italics="true" pageId="10" pageNumber="145">Cheleocloeon</emphasis>
</taxonomicName>
] should be moved to some taxon within the plesiomorphon Protopatellata.
</paragraph>
</subSubSection>
</treatment>
</document>